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<document id="3AFEE1E6910DC167DFBCCEA7C311ADDE" ID-CLB-Dataset="57602" ID-DOI="10.5852/ejt.2023.867.2109" ID-GBIF-Dataset="1d2570da-e150-4d75-94d8-bfc5813062bc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7891021" ID-ZooBank="8DEF295C-A8B1-4A6B-B873-B30949F64E07" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1683117081100" checkinUser="felipe" docAuthor="Iniesta, Luiz Felipe Moretti, Bouzan, Rodrigo Salvador &amp; Brescovit, Antonio Domingos" docDate="2023" docId="03F887BA1350B1054D00FAAAFAAC52DE" docLanguage="en" docName="ejt-867_1-312.pdf.imf" docOrigin="European Journal of Taxonomy 867 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.867.2109" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pseudonannolene aurea Iniesta &amp; Bouzan &amp; Brescovit 2023, sp. nov." docType="treatment" docUuid="0458352D-56B7-4A58-A71A-7E502759CA27" docUuidSource="ZooBank" docVersion="7" lastPageNumber="101" masterDocId="FFC1FFC21333B1614F56FFECFF9A553E" masterDocTitle="A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)" masterLastPageNumber="312" masterPageNumber="1" pageNumber="100" updateTime="1699515670513" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="C21911D1E843B63E56BAAE43CE8B75F0">
<mods:title id="C5F445591FA0609BBB9E4B8A59595A3E">A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)</mods:title>
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<mods:namePart id="3EA65AA4F874A598C4E14315F5AC8B2D">Iniesta, Luiz Felipe Moretti</mods:namePart>
<mods:nameIdentifier id="B9A631F28EA7FE7FF7BC57D63BC43CC7" type="LSID">DEEF048E-97FB-4CCD-875F-5FA6184CA8AB&amp;14A15A7F-730F-4D41-BDAC-D53514FAB85D</mods:nameIdentifier>
<mods:affiliation id="6DAD1CE58B7C417AE41760E79BD47CB7">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="5EF17F073B10797B9024DA19CB94BFCD" type="email">rodrigobouzan@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F76A4CEB091F53A8C046B1A5E1DE2515" type="personal">
<mods:role id="09EE84C31CDC138EFFE180FEA4F9BE29">
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<mods:namePart id="AA4FF99A27AB490D6790CB32EF219CE9">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:nameIdentifier id="F82E40AE54D054A2EA7C3FF27DA4FB5B" type="LSID">DEEF048E-97FB-4CCD-875F-5FA6184CA8AB&amp;14A15A7F-730F-4D41-BDAC-D53514FAB85D</mods:nameIdentifier>
<mods:affiliation id="D47BD2DA346846E608DCD506D6EBB4C1">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="EFC8E5EC11156F5BBB7D074FDD577C77" type="email">rodrigobouzan@outlook.com</mods:nameIdentifier>
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<mods:roleTerm id="1A6107EE48F5B14A3047B8D180DF1E63">Author</mods:roleTerm>
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<mods:namePart id="6AA413DF441CEF9DBBB4B7CBF492378C">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="F508EEB67E01D687909A046225C573B7">Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="684572F05F260250493C42BB3AA80B55" type="email">rodrigobouzan@outlook.com&amp;antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:title id="56A4AE2B3BA9B70EDEB64A5552DB55A0">European Journal of Taxonomy</mods:title>
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<mods:date id="699E2B397DA784A8FD5553CDFA8A0234">2023</mods:date>
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<mods:classification id="7BDA487B67626E2B684DDAC576E5D5F5">journal article</mods:classification>
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<treatment id="03F887BA1350B1054D00FAAAFAAC52DE" ID-DOI="http://doi.org/10.5281/zenodo.7907953" ID-GBIF-Taxon="209588426" ID-Zenodo-Dep="7907953" LSID="urn:lsid:zoobank.org:act:0458352D-56B7-4A58-A71A-7E502759CA27" httpUri="http://treatment.plazi.org/id/03F887BA1350B1054D00FAAAFAAC52DE" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100">
<subSubSection id="C34B65271350B1024D00FAAAFB5B50B8" pageId="99" pageNumber="100" type="nomenclature">
<paragraph id="8BEE36AC1350B1024D00FAAAFC44505C" blockId="99.[370,1217,1350,1449]" box="[598,990,1350,1378]" pageId="99" pageNumber="100">
<heading id="D0A681C01350B1024D00FAAAFC44505C" box="[598,990,1350,1378]" centered="true" fontSize="11" level="2" pageId="99" pageNumber="100" reason="2">
<taxonomicName id="4C514D2F1350B1024D00FAAAFCE6505F" ID-CoL="041ae964-e7e3-4813-83c7-fb6b789eccbd" authority="Iniesta &amp; Bouzan &amp; Brescovit, 2023" authorityName="Iniesta &amp; Bouzan &amp; Brescovit" authorityYear="2023" box="[598,892,1350,1377]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="aurea" status="sp. nov.">
<emphasis id="B925EABE1350B1024D00FAAAFCE6505F" bold="true" box="[598,892,1350,1377]" italics="true" pageId="99" pageNumber="100">Pseudonannolene aurea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21657C51350B1024CD5FAA4FC44505C" box="[899,990,1352,1378]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34B65271350B1024D0CFA62FC405097" box="[602,986,1422,1449]" pageId="99" pageNumber="100" type="description">
<paragraph id="8BEE36AC1350B1024D0CFA62FC405097" blockId="99.[370,1217,1350,1449]" box="[602,986,1422,1449]" pageId="99" pageNumber="100">
<figureCitation id="136A2A291350B1024D0CFA62FD675097" box="[602,765,1422,1449]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="251.[189,232,760,786]" captionStartId-1="252.[189,232,1665,1691]" captionTargetBox-0="[189,1390,283,719]" captionTargetBox-1="[189,1398,265,1622]" captionTargetId-0="figure-11@251.[189,1398,267,719]" captionTargetId-1="figure-14@252.[189,1398,265,1622]" captionTargetPageId-0="251" captionTargetPageId-1="252" captionText-0="Fig. 147. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854), in lateral view. A. Anterior region. B. Posterior region. Scale bars = 1 mm." captionText-1="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7891363" figureDoi-1="http://doi.org/10.5281/zenodo.7891365" httpUri-0="https://zenodo.org/record/7891363/files/figure.png" httpUri-1="https://zenodo.org/record/7891365/files/figure.png" pageId="99" pageNumber="100">Figs 147–148</figureCitation>
,
<figureCitation id="136A2A291350B1024C5FFA62FCD75097" box="[777,845,1422,1449]" captionStart="Fig" captionStartId="278.[189,232,1928,1954]" captionTargetBox="[199,1388,265,1886]" captionTargetId="figure-14@278.[199,1388,265,1886]" captionTargetPageId="278" captionText="Fig. 175. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. granulata sp. nov. B. P. alata sp. nov. C. P. curvata sp. nov. D. P. aurea sp. nov. E. P. alegrensis. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891435" httpUri="https://zenodo.org/record/7891435/files/figure.png" pageId="99" pageNumber="100">175D</figureCitation>
,
<figureCitation id="136A2A291350B1024C0FFA63FC3B5097" box="[857,929,1423,1449]" captionStart="Fig" captionStartId="282.[189,232,1857,1883]" captionTargetBox="[225,1363,265,1811]" captionTargetId="figure-14@282.[225,1363,265,1811]" captionTargetPageId="282" captionText="Fig. 179. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. strinatii Mauriès, 1974. B. P. tocaiensis Fontanetti, 1996. C. P. tricolor Brölemann, 1902. D. P. typica Silvestri, 1895. E. P. xavieri Iniesta &amp; Ferreira, 2014. F. P. bucculenta sp. nov. G. P. morettii sp. nov. H. P. insularis sp. nov. I. P. nicolau sp. nov. J. P. granulata sp. nov. K. P. alata sp. nov. L. P. curvata sp. nov. M. P. aurea sp. nov. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891443" httpUri="https://zenodo.org/record/7891443/files/figure.png" pageId="99" pageNumber="100">179M</figureCitation>
,
<figureCitation id="136A2A291350B1024CF8FA63FC405097" box="[942,986,1423,1449]" captionStart="Fig" captionStartId="293.[189,232,1363,1389]" captionTargetBox="[193,1368,289,1302]" captionTargetId="figure-12@293.[189,1398,265,1319]" captionTargetPageId="293" captionText="Fig. 190. Distribution map of the speciesPseudonannolene alata sp. nov.,P. aureasp. nov., P. bucculenta sp. nov., P. curvata sp. nov., and P. granulata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7891465" httpUri="https://zenodo.org/record/7891465/files/figure.png" pageId="99" pageNumber="100">190</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B65271350B1024FEBFA3CFCFC535F" pageId="99" pageNumber="100" type="diagnosis">
<paragraph id="8BEE36AC1350B1024FEBFA3CFEAC50D4" blockId="99.[189,310,1488,1514]" box="[189,310,1488,1514]" pageId="99" pageNumber="100">
<heading id="D0A681C01350B1024FEBFA3CFEAC50D4" bold="true" box="[189,310,1488,1514]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="B925EABE1350B1024FEBFA3CFEAC50D4" bold="true" box="[189,310,1488,1514]" pageId="99" pageNumber="100">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1350B1024FEBFA13FCFC535F" blockId="99.[187,1399,1535,1633]" pageId="99" pageNumber="100">
Males of
<taxonomicName id="4C514D2F1350B1024E7EF9ECFE1C5327" authority="Iniesta &amp; Bouzan &amp; Brescovit, 2023" authorityName="Iniesta &amp; Bouzan &amp; Brescovit" authorityYear="2023" box="[296,390,1536,1561]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="aurea" status="sp. nov.">
<emphasis id="B925EABE1350B1024E7EF9ECFE1C5327" box="[296,390,1536,1561]" italics="true" pageId="99" pageNumber="100">P. aurea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21657C51350B1024ED8F9ECFE7D5324" box="[398,487,1536,1562]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
slightly resemble those of
<taxonomicName id="4C514D2F1350B1024C4AF9ECFC0D5327" authorityName="Brolemann" authorityYear="1902" box="[796,919,1535,1561]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="paulista">
<emphasis id="B925EABE1350B1024C4AF9ECFC0D5327" box="[796,919,1535,1561]" italics="true" pageId="99" pageNumber="100">P. paulista</emphasis>
</taxonomicName>
by having a solenomere with a spiniform ectal process deeply notched separating from apicomesal process (
<figureCitation id="136A2A291350B1024CE0F9CFFBAB5303" box="[950,1073,1571,1597]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="99" pageNumber="100">Fig. 148D</figureCitation>
), but differing by an internal branch subtriangular with distal projection (
<figureCitation id="136A2A291350B1024DE8F9ABFCC0535F" box="[702,858,1607,1633]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="99" pageNumber="100">Fig. 148D–F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271350B1024FEBF965FB6A53C8" pageId="99" pageNumber="100" type="etymology">
<paragraph id="8BEE36AC1350B1024FEBF965FEDE539D" blockId="99.[189,324,1673,1699]" box="[189,324,1673,1699]" pageId="99" pageNumber="100">
<heading id="D0A681C01350B1024FEBF965FEDE539D" bold="true" box="[189,324,1673,1699]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="B925EABE1350B1024FEBF965FEDE539D" bold="true" box="[189,324,1673,1699]" pageId="99" pageNumber="100">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1350B1024FEBF954FB6A53C8" blockId="99.[189,1399,1720,1782]" pageId="99" pageNumber="100">
The species epithet is derived from the Latin adjective ‘
<emphasis id="B925EABE1350B1024C3CF955FC2D53EC" box="[874,951,1721,1746]" italics="true" pageId="99" pageNumber="100">aurum</emphasis>
’ = ‘golden’; in reference to the
<typeStatus id="54EA880E1350B1024A12F955FAED53ED" box="[1348,1399,1721,1747]" pageId="99" pageNumber="100">type</typeStatus>
locality Dianópolis, which is historically known for the gold mining activities in the region.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271350B1054FEBF8F1FC70574B" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100" type="materials_examined">
<paragraph id="8BEE36AC1350B1024FEBF8F1FE305209" blockId="99.[189,426,1821,1847]" box="[189,426,1821,1847]" pageId="99" pageNumber="100">
<heading id="D0A681C01350B1024FEBF8F1FE305209" bold="true" box="[189,426,1821,1847]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="B925EABE1350B1024FEBF8F1FE305209" bold="true" box="[189,426,1821,1847]" pageId="99" pageNumber="100">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1350B1024FBCF8A1FDC15291" blockId="99.[189,1399,1869,1967]" pageId="99" pageNumber="100">
<materialsCitation id="3B393CF11350B1024FBCF8A1FDC15291" ID-GBIF-Occurrence="4101310053" collectingDate="2017-12-04" collectingDateMax="2017-12-09" collectingDateMin="2017-12-04" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" elevation="672" latitude="-11.624226" location="Mojadores cave" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" pageId="99" pageNumber="100" specimenCode="IBSP 5858" specimenCount="1" specimenCount-male="1" stateProvince="Tocantins" typeStatus="holotype">
<emphasis id="B925EABE1350B1024FBCF8A1FEC65259" bold="true" box="[234,348,1869,1895]" pageId="99" pageNumber="100">
<typeStatus id="54EA880E1350B1024FBCF8A1FEC65259" box="[234,348,1869,1895]" pageId="99" pageNumber="100" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F346763C1350B1024FEBF89DFEBC52B5" box="[189,294,1905,1931]" name="Brazil" pageId="99" pageNumber="100">BRAZIL</collectingCountry>
•
<specimenCount id="9D57FD251350B1024E68F89DFECD52B5" box="[318,343,1905,1931]" pageId="99" pageNumber="100" type="male">♂</specimenCount>
;
<collectingRegion id="4995F84E1350B1024E34F89CFE4D52B4" box="[354,471,1904,1930]" country="Brazil" name="Tocantins" pageId="99" pageNumber="100">Tocantins</collectingRegion>
,
<collectingMunicipality id="6B8AACD61350B1024EB4F89CFDFC52B4" box="[482,614,1904,1930]" pageId="99" pageNumber="100">Dianópolis</collectingMunicipality>
,
<location id="8E8E60771350B1024D27F89CFCB452B5" LSID="urn:lsid:plazi:treatment:03F887BA1350B1054D00FAAAFAAC52DE:8E8E60771350B1024D27F89CFCB452B5" box="[625,814,1904,1931]" country="Brazil" latitude="-11.624226" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" name="Mojadores cave" pageId="99" pageNumber="100" stateProvince="Tocantins">Mojadores cave</location>
; [
<geoCoordinate id="EE65506B1350B1024C17F89CFC5652B4" box="[833,972,1904,1930]" degrees="11.624226" direction="south" orientation="latitude" pageId="99" pageNumber="100" precision="1" value="-11.624226">-11.624226</geoCoordinate>
,
<geoCoordinate id="EE65506B1350B1024C8EF89CFBFE52B5" box="[984,1124,1904,1931]" degrees="46.820593" direction="west" orientation="longitude" pageId="99" pageNumber="100" precision="1" value="-46.820595">-46.820593</geoCoordinate>
];
<elevation id="007CD19F1350B1024B21F89CFA9A52B4" box="[1143,1280,1904,1931]" metricMagnitude="2" metricUnit="m" metricValue="6.72" pageId="99" pageNumber="100" unit="m" value="672.0">
<quantity id="4CA99B491350B1024B21F89CFB5B52B5" box="[1143,1217,1904,1931]" metricMagnitude="2" metricUnit="m" metricValue="6.72" pageId="99" pageNumber="100" unit="m" value="672.0">672 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C1350B1024A5BF89DFF615290" pageId="99" pageNumber="100" value="2017-12-04" valueMax="2017-12-09" valueMin="2017-12-04">
<collectingDate id="EFABE9841350B1024A5BF89DFF615290" pageId="99" pageNumber="100" value="2017-12-04" valueMax="2017-12-09" valueMin="2017-12-04">4–9 Dec. 2017</collectingDate>
</date>
;
<collectorName id="26A4537A1350B1024E51F879FE095290" box="[263,403,1940,1967]" pageId="99" pageNumber="100">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71350B1024E83F878FDCD5291" box="[469,599,1940,1967]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="99" pageNumber="100">IBSP 5858</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE36AC1357B1054FBCFEE5FE1454B4" blockId="100.[189,1399,265,394]" pageId="100" pageNumber="101">
<materialsCitation id="3B393CF11357B1054FBCFEE5FD735456" ID-GBIF-Occurrence="4101309577" collectingDate="2008-05-21" collectingDateMax="2008-05-29" collectingDateMin="2008-05-21" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" elevation="670" latitude="-11.624226" location="Areia cave" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" pageId="100" pageNumber="101" specimenCode="IBSP 5854" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Tocantins" typeStatus="paratype">
<emphasis id="B925EABE1357B1054FBCFEE5FEFD541D" bold="true" box="[234,359,265,291]" pageId="100" pageNumber="101">
<typeStatus id="54EA880E1357B1054FBCFEE5FEFD541D" box="[234,359,265,291]" pageId="100" pageNumber="101" type="paratype">Paratypes</typeStatus>
</emphasis>
(total:
<specimenCount id="9D57FD251357B1054EEBFEE5FE71541D" box="[445,491,265,291]" pageId="100" pageNumber="101" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD251357B1054EA1FEE5FDAD541D" box="[503,567,265,291]" pageId="100" pageNumber="101" type="female">3 ♀♀</specimenCount>
)
<collectingCountry id="F346763C1357B1054FEBFEC7FEBC547B" box="[189,294,299,325]" name="Brazil" pageId="100" pageNumber="101">BRAZIL</collectingCountry>
–
<emphasis id="B925EABE1357B1054E1FFEC7FE58547B" bold="true" box="[329,450,299,325]" pageId="100" pageNumber="101">
<collectingRegion id="4995F84E1357B1054E1FFEC7FE58547B" box="[329,450,299,325]" country="Brazil" name="Tocantins" pageId="100" pageNumber="101">Tocantins</collectingRegion>
</emphasis>
•
<specimenCount id="9D57FD251357B1054EB7FEC7FD885478" box="[481,530,299,326]" pageId="100" pageNumber="101" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD251357B1054D77FEC7FDFD5478" box="[545,615,299,326]" pageId="100" pageNumber="101" type="female">2 ♀♀</specimenCount>
;
<collectingMunicipality id="6B8AACD61357B1054D2FFEC7FD67547B" box="[633,765,299,325]" pageId="100" pageNumber="101">Dianópolis</collectingMunicipality>
,
<location id="8E8E60771357B1054C5FFEC7FC145478" LSID="urn:lsid:plazi:treatment:03F887BA1350B1054D00FAAAFAAC52DE:8E8E60771357B1054C5FFEC7FC145478" box="[777,910,299,326]" country="Brazil" latitude="-11.624226" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" name="Areia cave" pageId="100" pageNumber="101" stateProvince="Tocantins">Areia cave</location>
; [
<geoCoordinate id="EE65506B1357B1054CF2FEC7FBB5547B" box="[932,1071,299,325]" degrees="11.624226" direction="south" orientation="latitude" pageId="100" pageNumber="101" precision="1" value="-11.624226">-11.624226</geoCoordinate>
,
<geoCoordinate id="EE65506B1357B1054B68FEC6FB51547B" box="[1086,1227,298,325]" degrees="46.820593" direction="west" orientation="longitude" pageId="100" pageNumber="101" precision="1" value="-46.820595">-46.820593</geoCoordinate>
];
<elevation id="007CD19F1357B1054BB6FEC7FAEA547B" box="[1248,1392,299,326]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="100" pageNumber="101" unit="m" value="670.0">
<quantity id="4CA99B491357B1054BB6FEC7FAB45478" box="[1248,1326,299,326]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="100" pageNumber="101" unit="m" value="670.0">670 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C1357B1054FEBFEA1FE115459" box="[189,395,333,360]" pageId="100" pageNumber="101" value="2008-05-21" valueMax="2008-05-29" valueMin="2008-05-21">
<collectingDate id="EFABE9841357B1054FEBFEA1FE115459" box="[189,395,333,360]" pageId="100" pageNumber="101" value="2008-05-21" valueMax="2008-05-29" valueMin="2008-05-21">21–29 May 2008</collectingDate>
</date>
;
<collectorName id="26A4537A1357B1054ECEFEA2FDBF5459" box="[408,549,333,360]" pageId="100" pageNumber="101">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71357B1054D3EFEA1FD735456" box="[616,745,333,360]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5854</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11357B1054C55FEA1FE1354B4" ID-GBIF-Occurrence="4101309349" collectingDate="2008-03-04" collectingDateMax="2008-03-12" collectingDateMin="2008-03-04" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" elevation="672" latitude="-11.624226" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" pageId="100" pageNumber="101" specimenCode="IBSP 5856" specimenCount="1" specimenCount-female="1" stateProvince="Tocantins" typeStatus="holotype">
<specimenCount id="9D57FD251357B1054C55FEA1FCAB5456" box="[771,817,333,360]" pageId="100" pageNumber="101" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<date id="FFEF106C1357B1054BABFEA1FF6154B7" pageId="100" pageNumber="101" value="2008-03-04" valueMax="2008-03-12" valueMin="2008-03-04">
<collectingDate id="EFABE9841357B1054BABFEA1FF6154B7" pageId="100" pageNumber="101" value="2008-03-04" valueMax="2008-03-12" valueMin="2008-03-04">4–12 Mar. 2008</collectingDate>
</date>
;
<specimenCode id="DBF79ED71357B1054E51FE83FE1354B4" box="[263,393,367,394]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5856</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36AC1357B1054FBCFE43FC7954F5" blockId="100.[189,1399,431,629]" box="[234,995,431,459]" pageId="100" pageNumber="101">
<emphasis id="B925EABE1357B1054FBCFE43FDD854F7" bold="true" box="[234,578,431,458]" pageId="100" pageNumber="101">Referred non-type material</emphasis>
(total:
<specimenCount id="9D57FD251357B1054DC0FE5CFD4054F4" box="[662,730,432,458]" pageId="100" pageNumber="101" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D57FD251357B1054DB1FE5CFCB054F4" box="[743,810,432,458]" pageId="100" pageNumber="101" type="female">7 ♀♀</specimenCount>
;
<specimenCount id="9D57FD251357B1054C6EFE5CFC4154F5" box="[824,987,432,459]" pageId="100" pageNumber="101" type="immature">16 immatures</specimenCount>
)
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFE3EFC70574B" blockId="100.[189,1399,431,629]" pageId="100" pageNumber="101">
<materialsCitation id="3B393CF11357B1054FEBFE3EFCF25731" ID-GBIF-Occurrence="4101310130" collectingDate="2008-05-21" collectingDateMax="2008-05-29" collectingDateMin="2008-05-21" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" elevation="672" latitude="-11.624226" location="Vozinha cave" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" pageId="100" pageNumber="101" specimenCode="IBSP 5859" specimenCount="6" specimenCount-female="2" specimenCount-immature="4" stateProvince="Tocantins">
<collectingCountry id="F346763C1357B1054FEBFE3EFEBC54D2" box="[189,294,466,492]" name="Brazil" pageId="100" pageNumber="101">BRAZIL</collectingCountry>
–
<emphasis id="B925EABE1357B1054E1BFE3EFE5C54D2" bold="true" box="[333,454,466,492]" pageId="100" pageNumber="101">
<collectingRegion id="4995F84E1357B1054E1BFE3EFE5C54D2" box="[333,454,466,492]" country="Brazil" name="Tocantins" pageId="100" pageNumber="101">Tocantins</collectingRegion>
</emphasis>
•
<specimenCount id="9D57FD251357B1054EBCFE3EFDA854D2" box="[490,562,466,492]" pageId="100" pageNumber="101" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9D57FD251357B1054D13FE3EFD7A54D2" box="[581,736,466,492]" pageId="100" pageNumber="101" type="immature">4 immatures</specimenCount>
;
<collectingMunicipality id="6B8AACD61357B1054DA7FE3EFCEF54D2" box="[753,885,466,492]" pageId="100" pageNumber="101">Dianópolis</collectingMunicipality>
,
<location id="8E8E60771357B1054CD3FE3EFBB054D2" LSID="urn:lsid:plazi:treatment:03F887BA1350B1054D00FAAAFAAC52DE:8E8E60771357B1054CD3FE3EFBB054D2" box="[901,1066,466,492]" country="Brazil" latitude="-11.624226" longLatPrecision="1" longitude="-46.820595" municipality="Dianopolis" name="Vozinha cave" pageId="100" pageNumber="101" stateProvince="Tocantins">Vozinha cave</location>
; [
<geoCoordinate id="EE65506B1357B1054B14FE3EFB5754D2" box="[1090,1229,466,492]" degrees="11.624226" direction="south" orientation="latitude" pageId="100" pageNumber="101" precision="1" value="-11.624226">-11.624226</geoCoordinate>
,
<geoCoordinate id="EE65506B1357B1054B88FE3DFAF154D2" box="[1246,1387,465,492]" degrees="46.820593" direction="west" orientation="longitude" pageId="100" pageNumber="101" precision="1" value="-46.820595">-46.820593</geoCoordinate>
];
<elevation id="007CD19F1357B1054FEBFE18FEDE5730" box="[189,324,500,527]" metricMagnitude="2" metricUnit="m" metricValue="6.72" pageId="100" pageNumber="101" unit="m" value="672.0">
<quantity id="4CA99B491357B1054FEBFE18FE9C5731" box="[189,262,500,527]" metricMagnitude="2" metricUnit="m" metricValue="6.72" pageId="100" pageNumber="101" unit="m" value="672.0">672 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C1357B1054E19FE18FD825730" box="[335,536,500,527]" pageId="100" pageNumber="101" value="2008-05-21" valueMax="2008-05-29" valueMin="2008-05-21">
<collectingDate id="EFABE9841357B1054E19FE18FD825730" box="[335,536,500,527]" pageId="100" pageNumber="101" value="2008-05-21" valueMax="2008-05-29" valueMin="2008-05-21">21–29 May 2008</collectingDate>
</date>
;
<collectorName id="26A4537A1357B1054D74FE19FD365730" box="[546,684,500,527]" pageId="100" pageNumber="101">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71357B1054DBCFE18FCF25731" box="[746,872,500,527]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5859</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11357B1054C2BFE18FD01570F" ID-GBIF-Occurrence="4101309975" collectingDate="2008-03-04" collectingDateMax="2008-03-12" collectingDateMin="2008-03-04" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" location="Onca cave" pageId="100" pageNumber="101" specimenCode="IBSP 5840" specimenCount="12" specimenCount-female="2" specimenCount-immature="7" specimenCount-male="3" stateProvince="Tocantins">
<specimenCount id="9D57FD251357B1054C2BFE18FC255731" box="[893,959,500,527]" pageId="100" pageNumber="101" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D57FD251357B1054C9CFE18FB915731" box="[970,1035,500,527]" pageId="100" pageNumber="101" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9D57FD251357B1054B41FE19FB315730" box="[1047,1195,500,527]" pageId="100" pageNumber="101" type="immature">7 immatures</specimenCount>
;
<location id="8E8E60771357B1054BE3FE18FAAB5731" LSID="urn:lsid:plazi:treatment:03F887BA1350B1054D00FAAAFAAC52DE:8E8E60771357B1054BE3FE18FAAB5731" box="[1205,1329,500,527]" country="Brazil" name="Onca cave" pageId="100" pageNumber="101" stateProvince="Tocantins">Onça cave</location>
;
<date id="FFEF106C1357B1054A6AFE18FEA0570E" pageId="100" pageNumber="101" value="2008-03-04" valueMax="2008-03-12" valueMin="2008-03-04">
<collectingDate id="EFABE9841357B1054A6AFE18FEA0570E" pageId="100" pageNumber="101" value="2008-03-04" valueMax="2008-03-12" valueMin="2008-03-04">4–12 Mar. 2008</collectingDate>
</date>
;
<collectorName id="26A4537A1357B1054E1EFDFBFE4F570E" box="[328,469,534,561]" pageId="100" pageNumber="101">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71357B1054D4CFDFAFD01570F" box="[538,667,534,561]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5840</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11357B1054DE0FDFAFDC8576D" ID-GBIF-Occurrence="4101309356" collectingDate="2007-12-04" collectingDateMax="2007-12-09" collectingDateMin="2007-12-04" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" location="Onca cave" pageId="100" pageNumber="101" specimenCode="IBSP 5843" specimenCount="5" specimenCount-immature="5" stateProvince="Tocantins">
<specimenCount id="9D57FD251357B1054DE0FDFAFCD4570E" box="[694,846,534,561]" pageId="100" pageNumber="101" type="immature">5 immatures</specimenCount>
; same locality data as for preceding;
<date id="FFEF106C1357B1054A5DFDFAFF61576C" pageId="100" pageNumber="101" value="2007-12-04" valueMax="2007-12-09" valueMin="2007-12-04">
<collectingDate id="EFABE9841357B1054A5DFDFAFF61576C" pageId="100" pageNumber="101" value="2007-12-04" valueMax="2007-12-09" valueMin="2007-12-04">4–9 Dec. 2007</collectingDate>
</date>
;
<collectorName id="26A4537A1357B1054E51FDD5FE08576C" box="[263,402,568,595]" pageId="100" pageNumber="101">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71357B1054E85FDD4FDC8576D" box="[467,594,568,595]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5843</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11357B1054D3FFDD4FAFC576D" ID-GBIF-Occurrence="4101309600" box="[617,1382,568,595]" collectingDate="2008-05-21" collectingDateMax="2008-05-29" collectingDateMin="2008-05-21" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" location="Coluna cave" pageId="100" pageNumber="101" specimenCode="IBSP 5836" specimenCount="1" specimenCount-female="1" stateProvince="Tocantins">
<specimenCount id="9D57FD251357B1054D3FFDD4FD0C576D" box="[617,662,568,595]" pageId="100" pageNumber="101" type="female">1 ♀</specimenCount>
;
<location id="8E8E60771357B1054DF2FDD4FCA2576D" LSID="urn:lsid:plazi:treatment:03F887BA1350B1054D00FAAAFAAC52DE:8E8E60771357B1054DF2FDD4FCA2576D" box="[676,824,568,595]" country="Brazil" name="Coluna cave" pageId="100" pageNumber="101" stateProvince="Tocantins">Coluna cave</location>
;
<date id="FFEF106C1357B1054C12FDD4FB95576C" box="[836,1039,568,595]" pageId="100" pageNumber="101" value="2008-05-21" valueMax="2008-05-29" valueMin="2008-05-21">
<collectingDate id="EFABE9841357B1054C12FDD4FB95576C" box="[836,1039,568,595]" pageId="100" pageNumber="101" value="2008-05-21" valueMax="2008-05-29" valueMin="2008-05-21">21–29 May 2008</collectingDate>
</date>
;
<collectorName id="26A4537A1357B1054B4DFDD5FB3C576C" box="[1051,1190,568,595]" pageId="100" pageNumber="101">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71357B1054BB1FDD4FAFC576D" box="[1255,1382,568,595]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5836</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11357B1054FEBFDB7FC7F574B" ID-GBIF-Occurrence="4101309365" box="[189,997,602,629]" collectingDate="2009-05-21" collectingDateMax="2009-05-29" collectingDateMin="2009-05-21" collectionCode="IBSP" collectorName="F. Pellegatti" country="Brazil" location="Vertebra cave" pageId="100" pageNumber="101" specimenCode="IBSP 5837" specimenCount="2" specimenCount-female="2" stateProvince="Tocantins">
<specimenCount id="9D57FD251357B1054FEBFDB7FE9A574B" box="[189,256,603,629]" pageId="100" pageNumber="101" type="female">2 ♀♀</specimenCount>
;
<location id="8E8E60771357B1054E58FDB6FE28574B" LSID="urn:lsid:plazi:treatment:03F887BA1350B1054D00FAAAFAAC52DE:8E8E60771357B1054E58FDB6FE28574B" box="[270,434,602,629]" country="Brazil" name="Vertebra cave" pageId="100" pageNumber="101" stateProvince="Tocantins">Vertebra cave</location>
;
<date id="FFEF106C1357B1054EE8FDB7FD10574B" box="[446,650,603,629]" pageId="100" pageNumber="101" value="2009-05-21" valueMax="2009-05-29" valueMin="2009-05-21">
<collectingDate id="EFABE9841357B1054EE8FDB7FD10574B" box="[446,650,603,629]" pageId="100" pageNumber="101" value="2009-05-21" valueMax="2009-05-29" valueMin="2009-05-21">21–29 May 2009</collectingDate>
</date>
;
<collectorName id="26A4537A1357B1054DC0FDB7FCBB574A" box="[662,801,602,629]" pageId="100" pageNumber="101">F. Pellegatti</collectorName>
leg.;
<specimenCode id="DBF79ED71357B1054C35FDB7FC7F574B" box="[867,997,602,629]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="100" pageNumber="101">IBSP 5837</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271357B1054FEBFD76FB59524C" pageId="100" pageNumber="101" type="description">
<paragraph id="8BEE36AC1357B1054FEBFD76FED4578A" blockId="100.[189,334,666,692]" box="[189,334,666,692]" pageId="100" pageNumber="101">
<heading id="D0A681C01357B1054FEBFD76FED4578A" bold="true" box="[189,334,666,692]" fontSize="11" level="3" pageId="100" pageNumber="101" reason="3">
<emphasis id="B925EABE1357B1054FEBFD76FED4578A" bold="true" box="[189,334,666,692]" pageId="100" pageNumber="101">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFD25FEC85619" blockId="100.[189,1399,712,807]" pageId="100" pageNumber="101">
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. 58–63 body rings (1–2 apodous + telson). Males: body length
<quantity id="4CA99B491357B1054BD6FD24FB6A57DD" box="[1152,1264,712,739]" metricMagnitude="-2" metricUnit="m" metricValue="6.31" pageId="100" pageNumber="101" unit="mm" value="63.1">63.1 mm</quantity>
; maximum midbody diameter
<quantity id="4CA99B491357B1054EF5FD06FDAD563B" box="[419,567,746,773]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="100" pageNumber="101" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
. Females: body length
<quantity id="4CA99B491357B1054C01FD06FB93563B" box="[855,1033,746,773]" metricMagnitude="-2" metricUnit="m" metricValue="6.459999999999999" metricValueMax="6.779999999999999" metricValueMin="6.14" pageId="100" pageNumber="101" unit="mm" value="64.6" valueMax="67.8" valueMin="61.4">61.4–67.8 mm</quantity>
; maximum midbody diameter
<quantity id="4CA99B491357B1054FEBFCE1FED75619" box="[189,333,781,807]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="100" pageNumber="101" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 mm</quantity>
.
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFCA1FC6256B7" blockId="100.[189,1398,845,906]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBFCA1FE93565B" baselines="864,864" box="[189,265,845,871]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="100" pageNumber="101">COLOR</smallCapsWord>
. Body color brownish red; head, antennae, collum, and legs brownish; prozonites anteriorly greyish; metazonites with a medial band brown and a posterior lighter.
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFC5CFD605130" blockId="100.[188,1399,943,1038]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBFC5CFF6656F6" baselines="963,963" box="[189,252,944,970]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="100" pageNumber="101">HEAD</smallCapsWord>
. Antennae short, just reaching back to end of ring 5 when extended dorsally; relative antennomere lengths 1&lt;2&lt;3&gt;4≈5≈6&gt;7. Mandibular cardo with ventral margin narrow. Ommatidial cluster well-developed, elliptical; ca 35 ommatidia in 5 rows.
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFBD9FCAB518B" blockId="100.[189,1399,1076,1205]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBFBD9FF645173" baselines="1096,1096" box="[189,254,1077,1103]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="100" pageNumber="101">BODY</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054E5EFBD6FEDC5173" baselines="1096" box="[264,326,1082,1101]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="100" pageNumber="101">RINGS</smallCapsWord>
. Collum with lateral lobes rounded, with ca 10 striae, slightly curved ectad (
<figureCitation id="136A2A291357B1054BBDFBD8FAF05170" box="[1259,1386,1076,1102]" captionStart="Fig" captionStartId="251.[189,232,760,786]" captionTargetBox="[189,1390,283,719]" captionTargetId="figure-11@251.[189,1398,267,719]" captionTargetPageId="251" captionText="Fig. 147. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854), in lateral view. A. Anterior region. B. Posterior region. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891363" httpUri="https://zenodo.org/record/7891363/files/figure.png" pageId="100" pageNumber="101">Fig. 147A</figureCitation>
). Very faintly constricted between prozonite and metazonite; prozonites smooth; metazonites laterally with transverse striae slightly above ozopore in anterior body rings. Anterior sterna in midbody rings subrectangular, without transverse striae (
<figureCitation id="136A2A291357B1054DFEFB76FCBE518B" box="[680,804,1178,1205]" captionStart="Fig" captionStartId="278.[189,232,1928,1954]" captionTargetBox="[199,1388,265,1886]" captionTargetId="figure-14@278.[199,1388,265,1886]" captionTargetPageId="278" captionText="Fig. 175. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. granulata sp. nov. B. P. alata sp. nov. C. P. curvata sp. nov. D. P. aurea sp. nov. E. P. alegrensis. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891435" httpUri="https://zenodo.org/record/7891435/files/figure.png" pageId="100" pageNumber="101">Fig. 175D</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFB37FDD45065" blockId="100.[187,1398,1243,1372]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBFB37FF6051CD" baselines="1262,1262" box="[189,250,1243,1269]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="First" pageId="100" pageNumber="101">FIRST</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054E57FB0CFEFB51CD" baselines="1262,1262,1262" box="[257,353,1243,1269]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="100" pageNumber="101">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054E3EFB0CFE1851CD" baselines="1262" box="[360,386,1248,1267]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="100" pageNumber="101">OF</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054EDFFB0CFE5551CD" baselines="1262" box="[393,463,1248,1267]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="100" pageNumber="101">MALES</smallCapsWord>
. Coxae (
<emphasis id="B925EABE1357B1054D6FFB37FDCF51CB" bold="true" box="[569,597,1243,1269]" italics="true" pageId="100" pageNumber="101">cx</emphasis>
) short (less than half of remaining podomere lengths), subtriangular, with the base arched and strongly expanded, densely setose (
<figureCitation id="136A2A291357B1054C23FB11FC6A5029" box="[885,1008,1277,1303]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="100" pageNumber="101">Fig. 148A</figureCitation>
); prefemoral process (
<emphasis id="B925EABE1357B1054BA7FB10FA8F5028" bold="true" box="[1265,1301,1276,1302]" italics="true" pageId="100" pageNumber="101">prf</emphasis>
) as long as prefemur, subcylindrical, densely setose up to its median region (
<figureCitation id="136A2A291357B1054CB5FAF3FBC45007" box="[995,1118,1311,1337]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="100" pageNumber="101">Fig. 148B</figureCitation>
); remaining podomeres with setae along the mesal region.
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFA6EFAFA5080" blockId="100.[188,1398,1409,1470]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBFA6EFE8E50A4" baselines="1429,1429" box="[189,276,1410,1436]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Second" pageId="100" pageNumber="101">SECOND</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054E4AFA6BFEE150A4" baselines="1429,1429,1429" box="[284,379,1410,1436]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="100" pageNumber="101">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054ED4FA6BFE0650A4" baselines="1429" box="[386,412,1415,1434]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="100" pageNumber="101">OF</smallCapsWord>
<smallCapsWord id="8D08A0701357B1054EF5FA6BFE7350A4" baselines="1429" box="[419,489,1415,1434]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="100" pageNumber="101">MALES</smallCapsWord>
. Coxa (
<emphasis id="B925EABE1357B1054D10FA6EFDF850A2" bold="true" box="[582,610,1410,1436]" italics="true" pageId="100" pageNumber="101">cx</emphasis>
) large and rounded; penis (
<emphasis id="B925EABE1357B1054CF6FA6EFC2550A2" bold="true" box="[928,959,1410,1436]" italics="true" pageId="100" pageNumber="101">pn</emphasis>
) located at proximal region, rounded, not extended basally (
<figureCitation id="136A2A291357B1054EEBFA48FDA25080" box="[445,568,1444,1470]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="100" pageNumber="101">Fig. 148C</figureCitation>
); prefemur compressed dorsoventrally; remaining podomeres setose.
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBFA08FCDA5231" blockId="100.[188,1399,1508,1808]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBFA08FEAE50C3" baselines="1527,1528" box="[189,308,1508,1534]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="100" pageNumber="101">GONOPODS</smallCapsWord>
. Gonocoxa (
<emphasis id="B925EABE1357B1054E8DFA08FD9F50C0" bold="true" box="[475,517,1508,1534]" italics="true" pageId="100" pageNumber="101">gcx</emphasis>
) elongated, almost twice as long as telopodite, with the base arched; antero-posteriorly flattened (
<figureCitation id="136A2A291357B1054D5AF9EAFD33531E" box="[524,681,1542,1568]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="100" pageNumber="101">Fig. 148D–F</figureCitation>
); with rows of papillae mesally. Seminal groove (
<emphasis id="B925EABE1357B1054BA5F9EBFA97531F" bold="true" box="[1267,1293,1543,1569]" italics="true" pageId="100" pageNumber="101">sg</emphasis>
) curved; arising medially on mesal cavity and terminating apically on the seminal apophysis (
<emphasis id="B925EABE1357B1054B83F9C5FB75537D" bold="true" box="[1237,1263,1577,1603]" italics="true" pageId="100" pageNumber="101">sa</emphasis>
). Shoulder (
<emphasis id="B925EABE1357B1054F91F9A6FF79535B" bold="true" box="[199,227,1610,1637]" italics="true" pageId="100" pageNumber="101">sh</emphasis>
) subtriangular. Telopodite (
<emphasis id="B925EABE1357B1054D60F9A7FDD7535B" bold="true" box="[566,589,1611,1637]" italics="true" pageId="100" pageNumber="101">tp</emphasis>
) almost as wide as
<emphasis id="B925EABE1357B1054C1FF9A7FCE9535B" bold="true" box="[841,883,1611,1637]" italics="true" pageId="100" pageNumber="101">gcx</emphasis>
, with deep depression separating from
<emphasis id="B925EABE1357B1054A0DF9A6FAED535B" bold="true" box="[1371,1399,1610,1637]" italics="true" pageId="100" pageNumber="101">sh</emphasis>
(
<figureCitation id="136A2A291357B1054F90F980FEDB53B9" box="[198,321,1644,1671]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="100" pageNumber="101">Fig. 148D</figureCitation>
); solenomere (
<emphasis id="B925EABE1357B1054EBBF980FD9B53B9" bold="true" box="[493,513,1644,1671]" italics="true" pageId="100" pageNumber="101">sl</emphasis>
) with apicomesal process (
<emphasis id="B925EABE1357B1054C6BF981FCEB53B9" bold="true" box="[829,881,1645,1671]" italics="true" pageId="100" pageNumber="101">amp</emphasis>
) subtriangular; ectal process (
<emphasis id="B925EABE1357B1054B87F981FB7753B9" bold="true" box="[1233,1261,1645,1671]" italics="true" pageId="100" pageNumber="101">ep</emphasis>
) spiniform, elongated, separating from
<emphasis id="B925EABE1357B1054D51F963FDA15397" bold="true" box="[519,571,1679,1705]" italics="true" pageId="100" pageNumber="101">amp</emphasis>
by deep notch;
<emphasis id="B925EABE1357B1054C54F963FC865397" bold="true" box="[770,796,1679,1705]" italics="true" pageId="100" pageNumber="101">sa</emphasis>
located at mesal portion, slightly visible apically. Internal branch (
<emphasis id="B925EABE1357B1054ED3F95CFE0653F5" bold="true" box="[389,412,1712,1739]" italics="true" pageId="100" pageNumber="101">ib</emphasis>
) subtriangular, narrow, surrounding basally
<emphasis id="B925EABE1357B1054CE2F95DFC5153F5" bold="true" box="[948,971,1713,1739]" italics="true" pageId="100" pageNumber="101">tp</emphasis>
as a shield; with torsion of 180° in the distal portion and projection directed diagonally upwards;
<emphasis id="B925EABE1357B1054CEFF93EFC4A53D3" bold="true" box="[953,976,1746,1773]" italics="true" pageId="100" pageNumber="101">ib</emphasis>
with setae along its entire margin exceeding apically seminal region of
<emphasis id="B925EABE1357B1054D22F919FD12522E" bold="true" box="[628,648,1781,1808]" italics="true" pageId="100" pageNumber="101">sl</emphasis>
(
<figureCitation id="136A2A291357B1054DCEF919FCAE5231" box="[664,820,1781,1807]" captionStart="Fig" captionStartId="252.[189,232,1665,1691]" captionTargetBox="[189,1398,265,1622]" captionTargetId="figure-14@252.[189,1398,265,1622]" captionTargetPageId="252" captionText="Fig. 148. Pseudonannolene aurea sp. nov., paratype, ♂ (IBSP 5854). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891365" httpUri="https://zenodo.org/record/7891365/files/figure.png" pageId="100" pageNumber="101">Fig. 148D–F</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBF8DAFB59524C" blockId="100.[187,1398,1845,1906]" pageId="100" pageNumber="101">
<smallCapsWord id="8D08A0701357B1054FEBF8DAFE8E5270" baselines="1865,1865" box="[189,276,1846,1872]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="100" pageNumber="101">VULVAE</smallCapsWord>
. As typical for the genus. Bursa subtriangular, glabrous (
<figureCitation id="136A2A291357B1054CC9F8D9FB85526E" box="[927,1055,1845,1872]" captionStart="Fig" captionStartId="282.[189,232,1857,1883]" captionTargetBox="[225,1363,265,1811]" captionTargetId="figure-14@282.[225,1363,265,1811]" captionTargetPageId="282" captionText="Fig. 179. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. strinatii Mauriès, 1974. B. P. tocaiensis Fontanetti, 1996. C. P. tricolor Brölemann, 1902. D. P. typica Silvestri, 1895. E. P. xavieri Iniesta &amp; Ferreira, 2014. F. P. bucculenta sp. nov. G. P. morettii sp. nov. H. P. insularis sp. nov. I. P. nicolau sp. nov. J. P. granulata sp. nov. K. P. alata sp. nov. L. P. curvata sp. nov. M. P. aurea sp. nov. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891443" httpUri="https://zenodo.org/record/7891443/files/figure.png" pageId="100" pageNumber="101">Fig. 179M</figureCitation>
); internal valve subtriangular, slightly rounded mesally; operculum narrow, curved ectad; external valve subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271357B1054FEBF874FAAC52DE" box="[189,1334,1944,2016]" pageId="100" pageNumber="101" type="distribution">
<paragraph id="8BEE36AC1357B1054FEBF874FECC528C" blockId="100.[189,342,1944,1970]" box="[189,342,1944,1970]" pageId="100" pageNumber="101">
<heading id="D0A681C01357B1054FEBF874FECC528C" bold="true" box="[189,342,1944,1970]" fontSize="11" level="3" pageId="100" pageNumber="101" reason="3">
<emphasis id="B925EABE1357B1054FEBF874FECC528C" bold="true" box="[189,342,1944,1970]" pageId="100" pageNumber="101">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1357B1054FEBF82AFAAC52DE" blockId="100.[189,1334,1989,2016]" box="[189,1334,1989,2016]" pageId="100" pageNumber="101">
Known only from caves and surrounding forests in Dianópolis,
<collectingRegion id="4995F84E1357B1054CFEF829FBC552DE" box="[936,1119,1989,2016]" country="Brazil" name="Tocantins" pageId="100" pageNumber="101">Tocantins State</collectingRegion>
,
<collectingCountry id="F346763C1357B1054B3DF829FB2952E1" box="[1131,1203,1989,2015]" name="Brazil" pageId="100" pageNumber="101">Brazil</collectingCountry>
(
<figureCitation id="136A2A291357B1054B94F829FAB352DE" box="[1218,1321,1989,2016]" captionStart="Fig" captionStartId="293.[189,232,1363,1389]" captionTargetBox="[193,1368,289,1302]" captionTargetId="figure-12@293.[189,1398,265,1319]" captionTargetPageId="293" captionText="Fig. 190. Distribution map of the speciesPseudonannolene alata sp. nov.,P. aureasp. nov., P. bucculenta sp. nov., P. curvata sp. nov., and P. granulata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7891465" httpUri="https://zenodo.org/record/7891465/files/figure.png" pageId="100" pageNumber="101">Fig. 190</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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