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<document id="8BDE115D5AEE0369845CBF0EF508653B" ID-CLB-Dataset="276131" ID-DOI="10.1080/00222933.2012.707240" ID-GBIF-Dataset="1ee471b0-aef3-43f2-98fb-a60802490e74" ID-ISSN="1464-5262" ID-Zenodo-Dep="5200545" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628911929894" checkinUser="felipe" docAuthor="Joharchi, Omid, Halliday, Bruce &amp; Saboori, Alireza" docDate="2012" docId="03F787B5652B5F0788DFFE6EFEFBFA7B" docLanguage="en" docName="JNatHist.46.31-32.1999-2018.pdf" docOrigin="Journal of Natural History 46 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2012.707240" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Laelaspis pennatus Joharchi and Halliday 2012, sp. nov." docType="treatment" docVersion="5" lastPageNumber="2009" masterDocId="FFCEFFCD65215F0B8941FFB2FFB6FF96" masterDocTitle="Three new species of Laelaspis Berlese from Iran (Acari: Laelapidae), with a review of the species occurring in the Western Palaearctic Region" masterLastPageNumber="2018" masterPageNumber="1999" pageNumber="2007" updateTime="1699084505657" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9D8AC350A107DC3034BEEC0D4181E390">Three new species of Laelaspis Berlese from Iran (Acari: Laelapidae), with a review of the species occurring in the Western Palaearctic Region</mods:title>
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<mods:namePart id="396DC2151593C75D5E7E7DCD2206CDDA">Joharchi, Omid</mods:namePart>
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<mods:namePart id="21084193339A28BB6BE2D15F8C501E75">Halliday, Bruce</mods:namePart>
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<mods:namePart id="497FA6EC3E0BA4DB04D8A35DEE382A92">Saboori, Alireza</mods:namePart>
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<mods:title id="DE58A35504BA50F8F93BF03BC5169C7E">Journal of Natural History</mods:title>
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<mods:number id="CA62F7178E27E6BDF24CBD0F39EB94AF">31 - 32</mods:number>
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<mods:identifier id="D94DCCC6730FE095D8882E5F00324E0A" type="DOI">10.1080/00222933.2012.707240</mods:identifier>
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<treatment id="03F787B5652B5F0788DFFE6EFEFBFA7B" ID-GBIF-Taxon="217223924" LSID="urn:lsid:plazi:treatment:03F787B5652B5F0788DFFE6EFEFBFA7B" httpUri="http://treatment.plazi.org/id/03F787B5652B5F0788DFFE6EFEFBFA7B" lastPageId="12" lastPageNumber="2009" pageId="10" pageNumber="2007">
<subSubSection id="C3446528652B5F0188DFFE6EFC7FFE65" box="[414,969,475,500]" pageId="10" pageNumber="2007" type="nomenclature">
<paragraph id="8BE136A3652B5F0188DFFE6EFC7FFE65" blockId="10.[414,969,475,531]" box="[414,969,475,500]" pageId="10" pageNumber="2007">
<heading id="D0A981CF652B5F0188DFFE6EFC7FFE65" box="[414,969,475,500]" centered="true" fontSize="36" level="2" pageId="10" pageNumber="2007" reason="7">
<taxonomicName id="4C5E4D20652B5F0188DFFE6EFCD8FE65" authority="Joharchi and Halliday" authorityName="Joharchi and Halliday" authorityYear="2012" box="[414,878,475,500]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="2007" phylum="Arthropoda" rank="species" species="pennatus" status="sp. nov.">
<emphasis id="B92AEAB1652B5F0188DFFE6EFDD8FE62" bold="true" box="[414,622,476,500]" italics="true" pageId="10" pageNumber="2007">Laelaspis pennatus</emphasis>
Joharchi and Halliday
</taxonomicName>
<taxonomicNameLabel id="A21957CA652B5F018A34FE6EFC7FFE65" box="[885,969,476,499]" pageId="10" pageNumber="2007" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3446528652B5F078B39FE49FEFBFA7B" lastPageId="12" lastPageNumber="2009" pageId="10" pageNumber="2007" type="description">
<paragraph id="8BE136A3652B5F018B39FE49FD5EFD85" blockId="10.[414,969,475,531]" box="[632,744,507,531]" pageId="10" pageNumber="2007">
(
<figureCitation id="13652A26652B5F018BC1FE49FD56FD85" box="[640,736,507,531]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="10" pageNumber="2007">Figure 2</figureCitation>
)
</paragraph>
<paragraph id="8BE136A3652B5F018992FDEAFE00FDE6" blockId="10.[211,438,600,624]" box="[211,438,600,624]" pageId="10" pageNumber="2007">
<emphasis id="B92AEAB1652B5F018992FDEAFE00FDE6" box="[211,438,600,624]" italics="true" pageId="10" pageNumber="2007">Specimens examined</emphasis>
</paragraph>
<paragraph id="8BE136A3652B5F018992FD32FE99FD41" blockId="10.[211,1172,640,727]" pageId="10" pageNumber="2007">
<materialsCitation id="3B363CFE652B5F018992FD32FDD4FD2E" ID-GBIF-Occurrence="4434851304" collectingDate="2011-02-25" collectionCode="JAZM" collectorName="O. Joharchi" country="Iran" location="Esfahan" pageId="10" pageNumber="2007" specimenCount="1" stateProvince="Esfahan" typeStatus="holotype">
<typeStatus id="54E58801652B5F018992FD32FEF6FD0E" box="[211,320,640,664]" pageId="10" pageNumber="2007" type="holotype">Holotype</typeStatus>
, female,
<collectingCountry id="F3497633652B5F0188EFFD33FE57FD0E" box="[430,481,641,664]" name="Iran" pageId="10" pageNumber="2007">Iran</collectingCountry>
,
<collectingRegion id="499AF841652B5F0188B3FD32FDE4FD0E" box="[498,594,640,664]" country="Iran" name="Esfahan" pageId="10" pageNumber="2007">Esfahan</collectingRegion>
,
<date id="FFE01063652B5F018B23FD32FC81FD0E" box="[610,823,640,664]" pageId="10" pageNumber="2007" value="2011-02-25">
<collectingDate id="EFA4E98B652B5F018B23FD32FC81FD0E" box="[610,823,640,664]" pageId="10" pageNumber="2007" value="2011-02-25">25 February 2011</collectingDate>
</date>
,
<collectorName id="26AB5375652B5F018A09FD32FC78FD0E" box="[840,974,640,664]" pageId="10" pageNumber="2007">O. Joharchi</collectorName>
coll., in nest of
<taxonomicName id="4C5E4D20652B5F018992FD13FE6CFD2E" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[211,474,673,696]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="2007" phylum="Arthropoda" rank="species" species="caespitum">
<emphasis id="B92AEAB1652B5F018992FD13FE6CFD2E" box="[211,474,673,696]" italics="true" pageId="10" pageNumber="2007">Tetramorium caespitum</emphasis>
</taxonomicName>
(in
<collectionCode id="ED4FAE66652B5F018B4BFD12FDEFFD2E" box="[522,601,672,696]" pageId="10" pageNumber="2007">JAZM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B363CFE652B5F018B2FFD12FE9DFD41" ID-GBIF-Occurrence="4434851303" collectingDate="2011-02-25" collectionCode="JAZM" collectorName="O. Joharchi" country="Iran" location="Esfahan" pageId="10" pageNumber="2007" specimenCount="1" specimenCount-female="1" stateProvince="Esfahan" typeStatus="paratype">
<typeStatus id="54E58801652B5F018B2FFD12FD62FD2E" box="[622,724,672,696]" pageId="10" pageNumber="2007" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D58FD2A652B5F018BA0FD12FCE8FD2E" box="[737,862,672,696]" pageId="10" pageNumber="2007" type="female">one female</specimenCount>
, same data as holotype (in
<collectionCode id="ED4FAE66652B5F018992FD0DFE94FD41" box="[211,290,703,727]" pageId="10" pageNumber="2007">JAZM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE136A3652B5F018992FCAFFEE4FCA2" blockId="10.[211,338,797,820]" box="[211,338,797,820]" pageId="10" pageNumber="2007">
<emphasis id="B92AEAB1652B5F018992FCAFFEE4FCA2" box="[211,338,797,820]" italics="true" pageId="10" pageNumber="2007">Description</emphasis>
</paragraph>
<paragraph id="8BE136A3652B5F018992FCF6FD61FBED" blockId="10.[211,1172,836,1147]" pageId="10" pageNumber="2007">
<emphasis id="B92AEAB1652B5F018992FCF6FE4FFCCA" box="[211,505,836,860]" italics="true" pageId="10" pageNumber="2007">Female. Dorsal idiosoma.</emphasis>
Dorsal shield length 450488 µm, width 324354 µm (
<emphasis id="B92AEAB1652B5F01899DFCD7FF5FFCEA" box="[220,233,869,892]" italics="true" pageId="10" pageNumber="2007">n</emphasis>
= 2) (
<figureCitation id="13652A26652B5F018812FCD6FE60FCEA" box="[339,470,868,892]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="10" pageNumber="2007">Figure 2A</figureCitation>
). Shield oval-shaped, with reticulation, more distinct in opisthonotal region; with 39 pairs of long setae, 22 podonotal, 17 opisthonotal, including two pairs of Zx setae between J and Z setae, almost all setae slightly swollen at base, with pointed tip (
<figureCitation id="13652A26652B5F0188D2FC71FDB2FC4D" box="[403,516,963,987]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="10" pageNumber="2007">Figure 2B</figureCitation>
) and very long, long enough to reach well past base of next posterior setae (j6 5469 µm, J1 5969 µm), lateral setae thicker than central setae, almost all marginal setae including Z5 slightly serrated (
<figureCitation id="13652A26652B5F018A12FBB1FC7CFB8D" box="[851,970,1027,1051]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="10" pageNumber="2007">Figure 2C</figureCitation>
), 6975 µm long, almost double the length of J5, 4045 µm; opisthonotal region with three unpaired supernumerary seta Jx in each specimen. Shield with five pairs of large circular to oval-shaped pores, other pores inconspicuous.
</paragraph>
<paragraph id="8BE136A3652B5F078992FB10FDC5FE08" blockId="10.[211,1172,1186,1720]" lastBlockId="12.[211,1172,167,414]" lastPageId="12" lastPageNumber="2009" pageId="10" pageNumber="2007">
<emphasis id="B92AEAB1652B5F018992FB10FE27FB2C" box="[211,401,1186,1210]" italics="true" pageId="10" pageNumber="2007">Ventral idiosoma</emphasis>
(
<figureCitation id="13652A26652B5F0188E5FB10FDA9FB2C" box="[420,543,1186,1210]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="10" pageNumber="2007">Figure 2D</figureCitation>
). Tritosternum with narrow base 20 µm long, paired pilose laciniae 5472 µm long, pre-sternal shields absent, pre-sternal area with some weak transverse lines. Sternal shield length 109111 µm, narrowest between coxae II (7982 µm) widest between coxae II and III (136146 µm), with straight anterior margin and concave posterior margin, with three pairs of smooth sternal setae,
<emphasis id="B92AEAB1652B5F018D32FA91FB30FAAC" box="[1139,1158,1315,1338]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
1 3035 µm,
<emphasis id="B92AEAB1652B5F01881AFAF0FED8FACF" box="[347,366,1346,1369]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
2 3742 µm,
<emphasis id="B92AEAB1652B5F018B4CFAF0FD96FACF" box="[525,544,1346,1369]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
3 4043 µm, one pair of lyrifissures adjacent to setae
<emphasis id="B92AEAB1652B5F018992FAD0FF50FAEF" box="[211,230,1378,1401]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
1, a pair of larger lyrifissures between
<emphasis id="B92AEAB1652B5F018BDEFAD0FD04FAEF" box="[671,690,1378,1401]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
2 and
<emphasis id="B92AEAB1652B5F018BBDFAD0FCB9FAEF" box="[764,783,1378,1401]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
3; antero-lateral surface of sternal shield with lineate ornamentation, central area smooth. Metasternal platelets absent, metasternal setae
<emphasis id="B92AEAB1652B5F0188D8FA10FE1AFA2F" box="[409,428,1442,1465]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
4 (2735 µm) and metasternal pores located in soft skin; endopodal plates II/III fused to sternal shield, endopodal plates III/IV elongate, narrow, curved. Genital shield broad, length 235248 µm, maximum width 181196 µm, posterior margin rounded, abutting anal shield, surface with characteristic ornamentation including distinct Λ- shaped lines and polygonal ornamentation, bearing the genital setae
<emphasis id="B92AEAB1652B5F01887FF9F3FEE7F9CE" box="[318,337,1601,1624]" italics="true" pageId="10" pageNumber="2007">st</emphasis>
5 (5052 µm) and two pairs of long setae (60 µm) on its lateral edges. Paragenital pores located on soft skin lateral to shield behind coxae IV. Anal shield subtriangular, length 7989 µm, width 97106 µm; its anterior half with lineate ornamentation and a pair of lateral pores; post-anal seta 40 µm, longer and thicker than para-anal setae, 25 µm. Opisthogastric skin with long, narrow and oval metapodal plates very close to genital shield, one metapodal plate on the right side and two on the left side in both specimens, and 15 pairs of slightly serrate setae (Jv1 6779 µm, Jv2 6774 µm, Jv3 5254 µm, Jv5 7779 µm, Zv1 2225 µm, Zv2 3750 µm) and five pairs of pores. Exopodal plates forming subtriangular extensions behind coxae IV, narrow elongate exopodal plates II/III not fused to peritrematal shield. Peritreme extending from coxa IV to anterior of coxa I, peritrematal shield narrow, post-stigmatal section conspicuous, with two pairs of pores.
</paragraph>
<caption id="DF21662B652A5F0089CEF9E2FCEFF936" ID-DOI="http://doi.org/10.5281/zenodo.5200549" ID-Zenodo-Dep="5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="11" pageNumber="2008" startId="11.[143,210,1616,1638]" targetBox="[151,1102,163,1583]" targetPageId="11">
<paragraph id="8BE136A3652A5F0089CEF9E2FCEFF936" blockId="11.[143,1104,1616,1696]" pageId="11" pageNumber="2008">
Figure 2.
<taxonomicName id="4C5E4D20652A5F0089B7F9E2FD21F9F0" authority="Joharchi and Halliday" authorityName="Joharchi and Halliday" authorityYear="2012" box="[246,663,1616,1638]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="2008" phylum="Arthropoda" rank="species" species="pennatus" status="sp. nov.">
<emphasis id="B92AEAB1652A5F0089B7F9E2FE07F9F0" box="[246,433,1616,1638]" italics="true" pageId="11" pageNumber="2008">Laelaspis pennatus</emphasis>
Joharchi and Halliday
</taxonomicName>
<taxonomicNameLabel id="A21957CA652A5F008BDCF9E3FD5AF9F0" box="[669,748,1617,1638]" pageId="11" pageNumber="2008" rank="species">sp. nov.</taxonomicNameLabel>
, female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale).
</paragraph>
</caption>
<paragraph id="8BE136A3652D5F078992FE74FE37FD6B" blockId="12.[211,1172,454,765]" pageId="12" pageNumber="2009">
<emphasis id="B92AEAB1652D5F078992FE74FED0FE48" box="[211,358,454,478]" italics="true" pageId="12" pageNumber="2009">
<taxonomicName id="4C5E4D20652D5F078992FE74FED7FE48" authorityName="Krefft" authorityYear="1871" box="[211,353,454,478]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="2009" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Epistome triangular, smooth (
<figureCitation id="13652A26652D5F078B85FE74FC8AFE48" box="[708,828,454,478]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="12" pageNumber="2009">Figure 2E</figureCitation>
). Hypostomal groove with six rows of denticles each bearing 610 small teeth, and smooth anterior transverse line (
<figureCitation id="13652A26652D5F07899AFDB7FEE6FD8B" box="[219,336,517,541]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="12" pageNumber="2009">Figure 2F</figureCitation>
). Hypostomal seta h1 30 µm, h2 20 µm, h3 50 µm, palp coxal seta 20 µm. Corniculi robust and horn-like, reaching mid-level of palp femur. Palp chaetotaxy: trochanter 2 (v1 very thick), femur 5, genu 6, tibia 12, tarsus 15; all setae smooth and needle-like, palp tarsal claw two-tined (
<figureCitation id="13652A26652D5F078BC0FDD7FD4FFDEB" box="[641,761,613,637]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="12" pageNumber="2009">Figure 2H</figureCitation>
). Fixed digit of chelicera with a small blunt proximal tooth, followed by a slightly larger median tooth, and two small blunt distal teeth (
<figureCitation id="13652A26652D5F078821FD17FE6FFD2B" box="[352,473,677,701]" captionStart="Figure 2" captionStartId="11.[143,210,1616,1638]" captionTargetBox="[151,1102,163,1583]" captionTargetId="figure-5@11.[145,1102,163,1583]" captionTargetPageId="11" captionText="Figure 2. Laelaspis pennatus Joharchi and Halliday sp. nov., female. (A) Dorsal shield; (B) dorsal shield seta J3; (C) dorsal shield seta Z5; (D) ventral idiosoma; (E) epistome; (F) hypostome; (G) chelicera; (H) palp tarsal claw. (Note: B, C and H are not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5200549" httpUri="https://zenodo.org/record/5200549/files/figure.png" pageId="12" pageNumber="2009">Figure 2G</figureCitation>
); pilus dentilis moderately long and robust; dorsal seta short, prostrate; movable digit with two teeth; arthrodial membrane with a rounded flap and short filaments.
</paragraph>
<paragraph id="8BE136A3652D5F078992FC97FC21FBCD" blockId="12.[211,1172,804,1147]" pageId="12" pageNumber="2009">
<emphasis id="B92AEAB1652D5F078992FC97FEB9FCAA" box="[211,271,805,828]" italics="true" pageId="12" pageNumber="2009">Legs.</emphasis>
Legs II and III short (272298 µm, 272282 µm), I and IV longer (372410 µm, 372382 µm). Leg I: coxa 0 0/1 0/1 0, trochanter 1 0/1 1/2 1, femur 2 3/2 2/1 2 (
<emphasis id="B92AEAB1652D5F078D25FCF6FB3AFCCA" box="[1124,1164,836,860]" italics="true" pageId="12" pageNumber="2009">ad2</emphasis>
,
<emphasis id="B92AEAB1652D5F078992FCD6FF4DFCEA" box="[211,251,868,892]" italics="true" pageId="12" pageNumber="2009">ad3</emphasis>
,
<emphasis id="B92AEAB1652D5F078848FCD6FE9DFCEA" box="[265,299,868,892]" italics="true" pageId="12" pageNumber="2009">al1</emphasis>
,
<emphasis id="B92AEAB1652D5F078878FCD6FEEDFCEA" box="[313,347,868,892]" italics="true" pageId="12" pageNumber="2009">pl1</emphasis>
and
<emphasis id="B92AEAB1652D5F0788D5FCD6FE00FCEA" box="[404,438,868,892]" italics="true" pageId="12" pageNumber="2009">pl2</emphasis>
thick), genu 2 3/2 3/1 2, tibia 2 3/2 3/1 2. Leg II: coxa 0 0/1 0/1 0, trochanter 1 0/1 1/2 1, femur 2 3/1 2/2 1, genu 2 3/1 2/1 2, tibia 2 2/1 2/1 2. Leg III: coxa 0 0/1 0/1 0, trochanter 1 0/1 0/1 1 (all setae thick), femur 1 2/1 1/0 1, genu 2 2/1 2/1 1 (ventral setae thick), tibia: 2 1/1 2/1 1(ventral setae thick). Leg IV: coxa 0 0/1 0/0 0, trochanter 1 0/1 0/1 1 (
<emphasis id="B92AEAB1652D5F078B12FC51FDD1FC6D" box="[595,615,995,1019]" italics="true" pageId="12" pageNumber="2009">al</emphasis>
thick), femur 1 2/1 1/0 1 (
<emphasis id="B92AEAB1652D5F078AD4FC51FC1FFC6D" box="[917,937,995,1019]" italics="true" pageId="12" pageNumber="2009">al</emphasis>
,
<emphasis id="B92AEAB1652D5F078AF9FC51FC65FC6D" box="[952,979,995,1019]" italics="true" pageId="12" pageNumber="2009">ad</emphasis>
1 and
<emphasis id="B92AEAB1652D5F078D5BFC51FB83FC6D" box="[1050,1077,995,1019]" italics="true" pageId="12" pageNumber="2009">ad</emphasis>
2 thick), genu 2 2/1 3/0 1, tibia 2 1/1 3/1 2 (
<emphasis id="B92AEAB1652D5F078B38FBB1FD3BFB8D" box="[633,653,1027,1051]" italics="true" pageId="12" pageNumber="2009">al</emphasis>
1 and
<emphasis id="B92AEAB1652D5F078B98FBB1FD5BFB8D" box="[729,749,1027,1051]" italics="true" pageId="12" pageNumber="2009">al</emphasis>
2 long); all setae fine and needle-like unless otherwise noted. Tarsi IIV with 18 setae 3 3/2 3/2 3 +
<emphasis id="B92AEAB1652D5F078AE6FB96FC71FBAD" box="[935,967,1060,1083]" italics="true" pageId="12" pageNumber="2009">mv</emphasis>
,
<emphasis id="B92AEAB1652D5F078A97FB91FC4EFBAD" box="[982,1016,1059,1083]" italics="true" pageId="12" pageNumber="2009">md</emphasis>
. All pre-tarsi with a pair of claws and a long thin membranous ambulacrum.
</paragraph>
<paragraph id="8BE136A3652D5F0789BFFBD6FC3DFBED" blockId="12.[211,1172,804,1147]" box="[254,907,1123,1147]" pageId="12" pageNumber="2009">
<emphasis id="B92AEAB1652D5F0789BFFBD6FE49FBED" box="[254,511,1124,1147]" italics="true" pageId="12" pageNumber="2009">Insemination structures</emphasis>
not seen, apparently unsclerotized.
</paragraph>
<paragraph id="8BE136A3652D5F078992FB07FEFDFB5B" blockId="12.[211,331,1205,1229]" box="[211,331,1205,1229]" pageId="12" pageNumber="2009">
<emphasis id="B92AEAB1652D5F078992FB07FEFDFB5B" box="[211,331,1205,1229]" italics="true" pageId="12" pageNumber="2009">Etymology</emphasis>
</paragraph>
<paragraph id="8BE136A3652D5F078992FB6EFE58FA82" blockId="12.[211,1171,1244,1300]" pageId="12" pageNumber="2009">
The name
<taxonomicName id="4C5E4D20652D5F078809FB6FFE1FFB62" authorityName="Joharchi and Halliday" authorityYear="2012" box="[328,425,1245,1268]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="2009" phylum="Arthropoda" rank="species" species="pennatus">
<emphasis id="B92AEAB1652D5F078809FB6FFE1FFB62" box="[328,425,1245,1268]" italics="true" pageId="12" pageNumber="2009">pennatus</emphasis>
</taxonomicName>
(Latin
<emphasis id="B92AEAB1652D5F0788BBFB6FFD8DFB62" box="[506,571,1245,1268]" italics="true" pageId="12" pageNumber="2009">penna</emphasis>
, feather) refers to the feather-like appearance of many of the dorsal shield setae.
</paragraph>
<paragraph id="8BE136A3652D5F078992FAFDFEA5FAF0" blockId="12.[211,275,1359,1382]" box="[211,275,1359,1382]" pageId="12" pageNumber="2009">
<emphasis id="B92AEAB1652D5F078992FAFDFEA5FAF0" box="[211,275,1359,1382]" italics="true" pageId="12" pageNumber="2009">Notes</emphasis>
</paragraph>
<paragraph id="8BE136A3652D5F078992FAC4FEFBFA7B" blockId="12.[211,1172,1398,1518]" pageId="12" pageNumber="2009">
<taxonomicName id="4C5E4D20652D5F078992FAC4FE1EFA18" authorityName="Joharchi and Halliday" authorityYear="2012" box="[211,424,1398,1422]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="2009" phylum="Arthropoda" rank="species" species="pennatus">
<emphasis id="B92AEAB1652D5F078992FAC4FE1EFA18" box="[211,424,1398,1422]" italics="true" pageId="12" pageNumber="2009">Laelaspis pennatus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C5E4D20652D5F078B0AFAC5FD74FA18" authorityName="Joharchi and Halliday" authorityYear="2012" box="[587,706,1398,1422]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="2009" phylum="Arthropoda" rank="species" species="kamalii">
<emphasis id="B92AEAB1652D5F078B0AFAC5FDEBFA18" box="[587,605,1399,1422]" italics="true" pageId="12" pageNumber="2009">L</emphasis>
.
<emphasis id="B92AEAB1652D5F078B2EFAC4FD74FA18" box="[623,706,1398,1422]" italics="true" pageId="12" pageNumber="2009">kamalii</emphasis>
</taxonomicName>
, but differs from it by its dorsal shield seta Z5 much longer than J5; seta v1 on the palp trochanter very thick, movable digit of chelicera with two teeth, and the genital shield narrower than that of
<taxonomicName id="4C5E4D20652D5F078992FA65FEF0FA78" authorityName="Joharchi and Halliday" authorityYear="2012" box="[211,326,1494,1518]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="2009" phylum="Arthropoda" rank="species" species="kamalii">
<emphasis id="B92AEAB1652D5F078992FA65FF53FA78" box="[211,229,1495,1518]" italics="true" pageId="12" pageNumber="2009">L</emphasis>
.
<emphasis id="B92AEAB1652D5F0789B2FA64FEF0FA78" box="[243,326,1494,1518]" italics="true" pageId="12" pageNumber="2009">kamalii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>