218 lines
27 KiB
XML
218 lines
27 KiB
XML
<document id="6F9B7DC80B8AAE17AC7A329691B23CDC" ID-CLB-Dataset="45658" ID-DOI="10.11646/zootaxa.3802.4.1" ID-GBIF-Dataset="43524c34-a5b7-4f7e-908a-3e3b61ffc6db" ID-ISSN="1175-5326" ID-Zenodo-Dep="252490" ID-ZooBank="CF479CC3-C014-460D-9C71-3A6C2AB2778B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427181093" checkinUser="plazi" docAuthor="Jørgensen, Aslak, Boesgaard, Tom M., Møbjerg, Nadja & Kristensen, Reinhardt M." docDate="2014" docId="03F487B7FFAEFFF668CE190DD74DA7E9" docLanguage="en" docName="zt03802p443.pdf" docOrigin="Zootaxa 3802 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dipodarctus australiensis" docType="treatment" docVersion="9" lastPageNumber="412" masterDocId="FFCDFFCFFFA7FFFD68591B0DD61FA000" masterDocTitle="The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada" masterLastPageNumber="443" masterPageNumber="401" pageNumber="410" updateTime="1698330854638" updateUser="ExternalLinkService">
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<mods:title id="30FC089A0176DDE76E736D9EA1B20DF6">The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada</mods:title>
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<mods:name id="7639A7488E9F0B3F2DBAD9D2421D63A9" type="personal">
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<mods:namePart id="40235141BEFAC195FE17405C13BCC537">Jørgensen, Aslak</mods:namePart>
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<mods:namePart id="C05AADD251ACA4284F475BCA690AB772">Boesgaard, Tom M.</mods:namePart>
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<mods:namePart id="234A716B7A7C8E8E19E9343C88E3808D">Møbjerg, Nadja</mods:namePart>
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<mods:namePart id="3C9640925DFC520C7F37AB597BC8D5A2">Kristensen, Reinhardt M.</mods:namePart>
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<mods:title id="9232F348AFDC07C061E406CDAE9A9FEF">Zootaxa</mods:title>
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<mods:date id="AB4080F1BDD90EE7425F3B35CB4A1B78">2014</mods:date>
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<mods:number id="421DA129F69A49C65C1A0B39861D9739">3802</mods:number>
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<treatment id="03F487B7FFAEFFF668CE190DD74DA7E9" ID-DOI="http://doi.org/10.5281/zenodo.5691079" ID-Zenodo-Dep="5691079" LSID="urn:lsid:plazi:treatment:03F487B7FFAEFFF668CE190DD74DA7E9" httpUri="http://treatment.plazi.org/id/03F487B7FFAEFFF668CE190DD74DA7E9" lastPageId="11" lastPageNumber="412" pageId="9" pageNumber="410">
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<subSubSection id="C347652AFFAEFFF468CE190DD4F6A21A" box="[151,745,512,538]" pageId="9" pageNumber="410" type="description">
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<paragraph id="8BE236A1FFAEFFF468CE190DD4F6A21A" blockId="9.[151,745,512,538]" box="[151,745,512,538]" pageId="9" pageNumber="410">
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<heading id="D0AA81CDFFAEFFF468CE190DD4F6A21A" bold="true" box="[151,745,512,538]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
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<emphasis id="B929EAB3FFAEFFF468CE190DD4F6A21A" bold="true" box="[151,745,512,538]" pageId="9" pageNumber="410">
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Description of
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<taxonomicName id="4C5D4D22FFAEFFF46916190DD49CA21A" ID-CoL="f990701f-da67-4a99-aba1-7189a43835be" box="[335,643,512,538]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="410" phylum="Tardigrada" rank="species" species="australiensis">
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<emphasis id="B929EAB3FFAEFFF46916190DD49CA21A" bold="true" box="[335,643,512,538]" italics="true" pageId="9" pageNumber="410">Dipodarctus australiensis</emphasis>
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</taxonomicName>
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nov. sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C347652AFFAEFFF468CE194BD762A37F" pageId="9" pageNumber="410" type="diagnosis">
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<paragraph id="8BE236A1FFAEFFF468CE194BD762A37F" blockId="9.[151,1437,582,2011]" pageId="9" pageNumber="410">
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<emphasis id="B929EAB3FFAEFFF468CE194BD70EA25F" bold="true" box="[151,273,582,607]" pageId="9" pageNumber="410">Diagnosis.</emphasis>
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A small
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<taxonomicName id="4C5D4D22FFAEFFF4692C194BD41FA25F" box="[373,512,582,607]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="410" phylum="Tardigrada" rank="genus">
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<emphasis id="B929EAB3FFAEFFF4692C194BD41FA25F" box="[373,512,582,607]" italics="true" pageId="9" pageNumber="410">Dipodarctus</emphasis>
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</taxonomicName>
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-species. The primary clavae are much shorter than half the length of the body. The sausage-shaped secondary clavae are located ventrally behind the internal and external cirri. The head has two triangular projections (cephalic projections) between the common cirrophorus for lateral cirri and primary clavae. Between the two internal cirri a cuticular sheath (frontal cuticular sheath) is present. Two medium-size triangular projections, each with two small posterior indentations, are present between the leg pairs III and IV. The first three leg pairs with a long, dorsal digit IV with three spiraled folds. The leg pair IV with two very long internal digits possessing three spiraled folds, and the two shorter external digits with two spiraled folds. All the four pairs of feet possess two parallel peduncles in the tarsus. The sense organ of leg pair IV has a typical cirrophorus, ovoid scapus and spiny flagellum.
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</paragraph>
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</subSubSection>
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<subSubSection id="C347652AFFAEFFF4689E1887D4ACA433" pageId="9" pageNumber="410" type="materials_examined">
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<paragraph id="8BE236A1FFAEFFF4689E1887D4ACA433" blockId="9.[151,1437,582,2011]" pageId="9" pageNumber="410">
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<emphasis id="B929EAB3FFAEFFF4689E1887D768A3A3" bold="true" box="[199,375,906,931]" pageId="9" pageNumber="410">
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<typeStatus id="54E68803FFAEFFF4689E1887D71DA3A3" box="[199,258,906,931]" pageId="9" pageNumber="410">Type</typeStatus>
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material.
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</emphasis>
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The
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<typeStatus id="54E68803FFAEFFF469ED1887D409A3A3" box="[436,534,906,931]" pageId="9" pageNumber="410" type="holotype">holotype</typeStatus>
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(Fig. 2) is an adult male [
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<collectionCode id="ED4CAE64FFAEFFF46B1F1886D585A3A3" box="[838,922,907,931]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="410">ZMUC</collectionCode>
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<collectionCode id="ED4CAE64FFAEFFF46BFA1886D5C6A3A3" box="[931,985,907,931]" pageId="9" pageNumber="410">TAR</collectionCode>
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1285] with testis, seminal vesicles and fully developed male gonopore; it was collected on
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<date id="FFE31061FFAEFFF46ABD18A2D581A3C7" box="[740,926,943,967]" pageId="9" pageNumber="410" value="1999-01-11">11 January 1999</date>
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from carbonate sand inside Fish Rock Cave. Only three specimens of
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<taxonomicName id="4C5D4D22FFAEFFF4699C18DFD4F4A3EB" box="[453,747,978,1003]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="410" phylum="Tardigrada" rank="species" species="australiensis">
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<emphasis id="B929EAB3FFAEFFF4699C18DFD4F4A3EB" box="[453,747,978,1003]" italics="true" pageId="9" pageNumber="410">Dipodarctus australiensis</emphasis>
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</taxonomicName>
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<emphasis id="B929EAB3FFAEFFF46AA118DFD549A3EB" bold="true" box="[760,854,978,1003]" pageId="9" pageNumber="410">nov. sp.</emphasis>
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were retrieved from the samples (
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<tableCitation id="C6DF031AFFAEFFF46CAF18DFD350A3EB" box="[1270,1359,978,1003]" captionStart="TABLE 1" captionStartId="3.[151,238,152,175]" captionTargetBox="[159,1355,272,620]" captionTargetPageId="3" captionText="TABLE 1. Overview of the specimens collected at Fish Rock Cave (FRC) and Jim’s Cave (JC). If known the sex is stated as female (F), male (M) or larva (L). Undetermined species are also listed to give a better indication of the abundance of tardigrades in the samples." httpUri="http://table.plazi.org/id/DF226629FFA4FFFE68CE1B95D7B1A0EF" pageId="9" pageNumber="410" tableUuid="DF226629FFA4FFFE68CE1B95D7B1A0EF">Table 1</tableCitation>
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). One
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<typeStatus id="54E68803FFAEFFF468CE18FAD6E8A40F" box="[151,247,1015,1039]" pageId="9" pageNumber="410" type="paratype">paratype</typeStatus>
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could be seen in frontal view (Fig. 3). The
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<typeStatus id="54E68803FFAEFFF46AB118FAD507A40F" box="[744,792,1015,1039]" pageId="9" pageNumber="410">type</typeStatus>
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material is deposited at The Natural History Museum of
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<collectingCountry id="F34A7631FFAEFFF468CE1F17D71DA433" box="[151,258,1050,1075]" name="Denmark" pageId="9" pageNumber="410">Denmark</collectingCountry>
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, University of Copenhagen,
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<collectingCountry id="F34A7631FFAEFFF46A1A1F17D4B0A433" box="[579,687,1050,1075]" name="Denmark" pageId="9" pageNumber="410">Denmark</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C347652AFFAEFFF4689E1F33D59AA457" box="[199,901,1086,1111]" pageId="9" pageNumber="410" type="etymology">
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<paragraph id="8BE236A1FFAEFFF4689E1F33D59AA457" blockId="9.[151,1437,582,2011]" box="[199,901,1086,1111]" pageId="9" pageNumber="410">
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<emphasis id="B929EAB3FFAEFFF4689E1F33D751A457" bold="true" box="[199,334,1086,1111]" pageId="9" pageNumber="410">Etymology.</emphasis>
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The name derives from the continent of
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<collectingCountry id="F34A7631FFAEFFF46B4F1F33D59FA457" box="[790,896,1086,1111]" name="Australia" pageId="9" pageNumber="410">Australia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C347652AFFAEFFF6689E1F6FD2CBA6CA" lastPageId="11" lastPageNumber="412" pageId="9" pageNumber="410" type="description">
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<paragraph id="8BE236A1FFAEFFF4689E1F6FD52AA62A" blockId="9.[151,1437,582,2011]" pageId="9" pageNumber="410">
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<emphasis id="B929EAB3FFAEFFF4689E1F6FD7EFA47B" bold="true" box="[199,496,1122,1147]" pageId="9" pageNumber="410">
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Description of
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<typeStatus id="54E68803FFAEFFF469D91F6FD7F4A47B" box="[384,491,1122,1147]" pageId="9" pageNumber="410" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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The holotypic male (Fig. 2) is 73 µm long, with a maximum width of 33 µm between second and third leg pair. The head is relatively large compared to the slender body. The facial plate may be similar to the frontal ala (“ala anterior”, van der
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<bibRefCitation id="EFCC4B50FFAEFFF46A8A1FA7D578A4C3" author="Land" box="[723,871,1194,1219]" pageId="9" pageNumber="410" refString="van der Land, J. (1968) Florarctus antillensis, a new tardigrade from the coral sands of Cura ҫao. Studies on the Fauna of Curacao and other Caribbean Islands, 25, 140 - 146." type="journal article" year="1968">Land (1968))</bibRefCitation>
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in
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<taxonomicName id="4C5D4D22FFAEFFF46BD21FA7D24EA4C3" box="[907,1105,1194,1219]" class="Heterotardigrada" family="Halechiniscidae" genus="Wingstrandarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="410" phylum="Tardigrada" rank="genus">
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<emphasis id="B929EAB3FFAEFFF46BD21FA7D24EA4C3" box="[907,1105,1194,1219]" italics="true" pageId="9" pageNumber="410">Wingstrandarctus</emphasis>
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</taxonomicName>
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. In a frontal view (Fig. 3) the head is somewhat triangular because of the two lateral cephalic projections and the facial plate and it appears distinct from the body. A complete set of cephalic appendages is present and several amoebocytes are embedded in the brain. The amoebocytes are of two sizes: three to four large (1.5–2.0 µm) and 12 to 14 small (0.1–0.3 µm). The 15 µm long median cirrus has a large scapus and a slender composite flagellum divided into proximal and distal sections. The 15 µm internal cirrus consists of a scapus and a thin composite flagellum divided into proximal and distal sections. The 15 µm external cirrus has a long scapus, with a composite flagellum with short proximal and distal sections. The median, internal and external cirri are without distinctive cirrophori. The 13 µm lateral cirrus consists of a flagellum projected from the cirrophorus shared with the 37 µm primary clava. In the common cirrophorus a “van der Land’s organ” is visible (Fig. 2). Large sausage-shaped secondary clavae (13.5 µm long) are lying ventrally on both sides of a small slender mouth cone.
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</paragraph>
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<paragraph id="8BE236A1FFAEFFF4689E1D3BD5E2A74B" blockId="9.[151,1437,582,2011]" pageId="9" pageNumber="410">The buccal tube (15 µm long) ends in a small bulbous tip with the mouth opening in the center of the cone. The needle-shaped stylets are 13 µm long. The thin stylet supports are 3 µm long and attached to the buccal tube and the stylet furcae. Three slightly horn-shaped or arc-shaped placoids are visible in the three-lobed pharyngeal bulb. Through a short esophagus the pharyngeal bulb is connected to the large intestine, which ends at a posterior anus. The crescent-shaped male gonopore is placed on a small ovoid papilla 1.5 µm anterior of the anus. The anus is covered with two overlapping folds with a wave-like mid-ventral suture. Crescent-shaped to roundish spermatids and rod-shaped spermatozoa are visible in the seminal vesicles on each side of the gonopore. The large testis is filled with spermatocytes anteriorly and round-headed spermatids posteriorly.</paragraph>
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<paragraph id="8BE236A1FFAEFFF4689E1C5BD404A7DB" blockId="9.[151,1437,582,2011]" pageId="9" pageNumber="410">Each leg consists of a coxa, femur, tibia and tarsus. Coxa on leg pair II and III possess simple leg sense organs consisting of spines of equal length; 10 µm (II) 10 µm (III). The leg sense organ on leg pair I (8 µm long) consists of a scapus and a flagellum. The sense organ on leg IV consists of a cirrophore-like base, an ovoid scapus and a short flagellum with a distal spine.</paragraph>
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<paragraph id="8BE236A1FFADFFF768CE1C21D2D5A7C2" blockId="10.[151,1437,1836,1989]" pageId="10" pageNumber="411">
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<emphasis id="B929EAB3FFADFFF768CE1C21D703A741" bold="true" box="[151,284,1836,1858]" pageId="10" pageNumber="411">FIGURE 2.</emphasis>
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Drawing of the holotypic male of
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<taxonomicName id="4C5D4D22FFADFFF76A9B1C21D5CCA742" box="[706,979,1836,1858]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="411" phylum="Tardigrada" rank="species" species="australiensis">
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<emphasis id="B929EAB3FFADFFF76A9B1C21D5CCA742" box="[706,979,1836,1858]" italics="true" pageId="10" pageNumber="411">Dipodarctus australiensis</emphasis>
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</taxonomicName>
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<emphasis id="B929EAB3FFADFFF76BBF1C21D25CA741" bold="true" box="[998,1091,1836,1857]" pageId="10" pageNumber="411">nov. sp.</emphasis>
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(ventral view). Abbreviations: am—amoebocyte; an—anus; bt—buccal tube; cE—cirrus E; cg—claw gland; ec—external cirrus; ed1—2—external digit 1—2; fc—facial plate; go—gonopore; ic—internal cirrus; id—internal digit; lc—lateral cirrus; mc—median cirrus; mg—midgut; mo—mouth cone; pc—primary clava; pd—peduncle; pl—placoid; pr4—lateral process 4; sc—secondary clava; se1—4—leg sense organs 1–4; ss—stylet support; st—stylet; te—testis; vo1—“van der Land’s organ” of primary clava.
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</paragraph>
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<paragraph id="8BE236A1FFACFFF668CE1F6BD44AA4DA" blockId="11.[151,1436,1126,1242]" pageId="11" pageNumber="412">
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<emphasis id="B929EAB3FFACFFF668CE1F6BD709A47B" bold="true" box="[151,278,1126,1148]" pageId="11" pageNumber="412">FIGURE 3.</emphasis>
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Drawing of the head of
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<taxonomicName id="4C5D4D22FFACFFF66A721F6BD52BA47C" box="[555,820,1126,1148]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="australiensis">
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<emphasis id="B929EAB3FFACFFF66A721F6BD52BA47C" box="[555,820,1126,1148]" italics="true" pageId="11" pageNumber="412">Dipodarctus australiensis</emphasis>
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</taxonomicName>
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<emphasis id="B929EAB3FFACFFF66B191F6BD587A47B" bold="true" box="[832,920,1126,1147]" pageId="11" pageNumber="412">nov. sp.</emphasis>
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(frontal view). Abbreviations: am—amoebocyte; ce—lateral cephalic projection; cg—claw gland; ec—external cirrus; fc—facial plate; ic—internal cirrus; lc—lateral cirrus; mc—median cirrus; mo—mouth cone; pc—primary clava; sc—secondary clava; se1—leg sense organ 1; vo1—“van der Land’s organ” of primary clava; vr—spiraled folds.
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</paragraph>
|
||
<paragraph id="8BE236A1FFACFFF6689E1E0CD403A5F2" blockId="11.[151,1437,1281,2026]" pageId="11" pageNumber="412">Each tarsus has four digits. On the first three pairs of legs (I–III), three digits (ventral) are of equal length and have claws with a calcar in addition one long digit (dorsal) bears a simple claw. Claw glands are visible in the coxal part of each leg. Claws may be covered by a membranous sheath. At the base of the digits in all four pairs of legs, two horizontal peduncles are present. In the fourth leg pair the two external digits are 6 µm long and shorter than the two internal which are 9 µm long. The dorsal digit on leg pairs I–III have three spiraled-like folds (digits with parallel folds). On leg pair IV all four digits have the spiraled-like folds. The external digits have two folds, while the internal digits have three folds.</paragraph>
|
||
<paragraph id="8BE236A1FFACFFF6689E1EF3D437A6A6" blockId="11.[151,1437,1281,2026]" pageId="11" pageNumber="412">A long cirrus E is attached laterally on the trunk, anteriorly to the insertion of the coxa of leg pair IV. Cirrus E consists of a large cirrophore which is 6 µm long and a 29 µm long flagellum. The scapus is indistinct. A pair of triangular lateral processes, measuring 10 µm in length, is present between leg pair III and IV. The posterior edge of the process has two small indentations. The lateral processes consist only of a thin cuticular membrane and may be folded and therefore overlooked.</paragraph>
|
||
<paragraph id="8BE236A1FFACFFF6689E1DBCD2CBA6CA" blockId="11.[151,1437,1281,2026]" box="[199,1236,1713,1738]" pageId="11" pageNumber="412">
|
||
Morphometric data of the holotypic male of
|
||
<taxonomicName id="4C5D4D22FFACFFF66AE01DBED574A6CA" box="[697,875,1713,1738]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="australiensis">
|
||
<emphasis id="B929EAB3FFACFFF66AE01DBED574A6CA" box="[697,875,1713,1738]" italics="true" pageId="11" pageNumber="412">D. australiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929EAB3FFACFFF66B2B1DBCD5D4A6CA" bold="true" box="[882,971,1713,1738]" pageId="11" pageNumber="412">nov. sp.</emphasis>
|
||
is presented in Table 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347652AFFACFFF6689E1DD8D74DA7E9" pageId="11" pageNumber="412" type="discussion">
|
||
<paragraph id="8BE236A1FFACFFF6689E1DD8D74DA7E9" blockId="11.[151,1437,1281,2026]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAB3FFACFFF6689E1DD8D724A6EE" bold="true" box="[199,315,1749,1774]" pageId="11" pageNumber="412">Remarks.</emphasis>
|
||
Only three specimens were found in Fish Rock Cave.
|
||
<taxonomicName id="4C5D4D22FFACFFF66BFA1DDAD24BA6EE" box="[931,1108,1749,1774]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="australiensis">
|
||
<emphasis id="B929EAB3FFACFFF66BFA1DDAD24BA6EE" box="[931,1108,1749,1774]" italics="true" pageId="11" pageNumber="412">D. australiensis</emphasis>
|
||
</taxonomicName>
|
||
is related to
|
||
<taxonomicName id="4C5D4D22FFACFFF66CBE1DDAD383A6EE" box="[1255,1436,1749,1774]" class="Heterotardigrada" family="Halechiniscidae" genus="Hemitanarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="subterraneus">
|
||
<emphasis id="B929EAB3FFACFFF66CBE1DDAD383A6EE" box="[1255,1436,1749,1774]" italics="true" pageId="11" pageNumber="412">D. subterraneus</emphasis>
|
||
</taxonomicName>
|
||
as indicated by the many similar sensory structures, however, the presence of the triangular cephalic processes, the facial plate and the large lateral processes between leg III and leg IV are diagnostic of
|
||
<taxonomicName id="4C5D4D22FFACFFF66CD51C12D35FA736" box="[1164,1344,1821,1846]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="australiensis">
|
||
<emphasis id="B929EAB3FFACFFF66CD51C12D35FA736" box="[1164,1344,1821,1846]" italics="true" pageId="11" pageNumber="412">D. australiensis</emphasis>
|
||
</taxonomicName>
|
||
. de
|
||
<bibRefCitation id="EFCC4B50FFACFFF66D2C1C10D79FA75A" author="Zio" pageId="11" pageNumber="412" refString="de Zio Grimaldi, S., D'Addabbo Gallo, M., Morone De Lucia, M. R. & Troccoli, A. (1995 / 96) Hemitanarctus chimera n. g., n. sp., new Halechiniscidae from the Ionian Sea (Tardigrada, Heterotardigrada). Zoologischer Anzeiger, 234, 167 - 174." type="journal article" year="1995">
|
||
Zio Grimaldi
|
||
<emphasis id="B929EAB3FFACFFF6695B1C4ED725A75A" box="[258,314,1857,1882]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
|
||
(1995
|
||
</bibRefCitation>
|
||
/96) described two small lateral projections in
|
||
<taxonomicName id="4C5D4D22FFACFFF66BDC1C4ED226A75A" box="[901,1081,1857,1882]" class="Heterotardigrada" family="Halechiniscidae" genus="Hemitanarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="subterraneus">
|
||
<emphasis id="B929EAB3FFACFFF66BDC1C4ED226A75A" box="[901,1081,1857,1882]" italics="true" pageId="11" pageNumber="412">D. subterraneus</emphasis>
|
||
</taxonomicName>
|
||
from the Mediterranean Sea. In the original description by
|
||
<bibRefCitation id="EFCC4B50FFACFFF669911C68D4C9A77E" author="Renaud-Debyser" box="[456,726,1893,1918]" pageId="11" pageNumber="412" refString="Renaud-Debyser, J. (1959) Etudes sur la faune interstitielle des iles Bahamas. III. Tardigrades. Vie et Millieu, 10, 297 - 302." type="journal article" year="1959">Renaud-Debyser (1959)</bibRefCitation>
|
||
these structures were not described in the Bahamas-form even though two small lateral indentations are shown in the drawing. The long dorsal toe of legs I–III in
|
||
<taxonomicName id="4C5D4D22FFACFFF66CB21C86D383A7A2" box="[1259,1436,1929,1954]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="australiensis">
|
||
<emphasis id="B929EAB3FFACFFF66CB21C86D383A7A2" box="[1259,1436,1929,1954]" italics="true" pageId="11" pageNumber="412">D. australiensis</emphasis>
|
||
</taxonomicName>
|
||
is also observed in
|
||
<taxonomicName id="4C5D4D22FFACFFF669351CA2D7C0A7C6" box="[364,479,1965,1990]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="borrori">
|
||
<emphasis id="B929EAB3FFACFFF669351CA2D7C0A7C6" box="[364,479,1965,1990]" italics="true" pageId="11" pageNumber="412">D. borrori</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5D4D22FFACFFF66A4F1CA2D4BFA7C6" box="[534,672,1967,1990]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="susannae">
|
||
<emphasis id="B929EAB3FFACFFF66A4F1CA2D4BFA7C6" box="[534,672,1967,1990]" italics="true" pageId="11" pageNumber="412">D. susannae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929EAB3FFACFFF66AFE1CA0D51FA7C6" bold="true" box="[679,768,1965,1990]" pageId="11" pageNumber="412">nov. sp.</emphasis>
|
||
, but not in
|
||
<taxonomicName id="4C5D4D22FFACFFF66BD91CA2D237A7C6" box="[896,1064,1965,1990]" class="Heterotardigrada" family="Halechiniscidae" genus="Dipodarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="anaholensis">
|
||
<emphasis id="B929EAB3FFACFFF66BD91CA2D237A7C6" box="[896,1064,1965,1990]" italics="true" pageId="11" pageNumber="412">D. anaholensis</emphasis>
|
||
</taxonomicName>
|
||
and in the original description of
|
||
<taxonomicName id="4C5D4D22FFACFFF668CE1CDED754A7EA" box="[151,331,2001,2026]" class="Heterotardigrada" family="Halechiniscidae" genus="Hemitanarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="412" phylum="Tardigrada" rank="species" species="subterraneus">
|
||
<emphasis id="B929EAB3FFACFFF668CE1CDED754A7EA" box="[151,331,2001,2026]" italics="true" pageId="11" pageNumber="412">D. subterraneus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |