196 lines
32 KiB
XML
196 lines
32 KiB
XML
<document id="1A1E238A61B6BBEAC9405DB08DA8DA55" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFACFFC87EFCF932F7EFDC89" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis Molineri, 2010, gen. nov." docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="22" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="1B8E055182287EA38CEED4C30EA1BDD7">A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:namePart id="ADC9AA314ADAF25B366E2311ACB92139">Molineri, Carlos</mods:namePart>
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<mods:typeOfResource id="5B0972BB095B0517F8319370344417DD">text</mods:typeOfResource>
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<mods:title id="64246D3846EA0C1D9D2C39650C6F5DBE">Zootaxa</mods:title>
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<mods:part id="0E5B10675CB0177555000F4BAAB29560">
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<mods:date id="CCC86448B79778C648E856D771A6D18A">2010</mods:date>
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<mods:number id="EDEB18B7DE023C87E1C240248DF2DBA2">2481</mods:number>
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<mods:identifier id="4F1656DECC4CB762A1E243F5C9606733" type="DOI">10.5281/zenodo.195431</mods:identifier>
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<treatment id="03F39207FFACFFC87EFCF932F7EFDC89" ID-DOI="http://doi.org/10.5281/zenodo.5690577" ID-GBIF-Taxon="119386659" ID-Zenodo-Dep="5690577" LSID="urn:lsid:plazi:treatment:03F39207FFACFFC87EFCF932F7EFDC89" httpUri="http://treatment.plazi.org/id/03F39207FFACFFC87EFCF932F7EFDC89" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection id="C340709AFFACFFCB7EFCF932F472DFCB" box="[151,377,1712,1738]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8BE52311FFACFFCB7EFCF932F472DFCB" blockId="21.[151,377,1712,1738]" box="[151,377,1712,1738]" pageId="21" pageNumber="22">
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<heading id="D0AD947DFFACFFCB7EFCF932F472DFCB" bold="true" box="[151,377,1712,1738]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<emphasis id="B92EFF03FFACFFCB7EFCF932F472DFCB" bold="true" box="[151,377,1712,1738]" pageId="21" pageNumber="22">
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<taxonomicName id="4C5A5892FFACFFCB7EFCF932F409DFCB" ID-CoL="7XX2" box="[151,258,1712,1738]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B92EFF03FFACFFCB7EFCF932F409DFCB" bold="true" box="[151,258,1712,1738]" italics="true" pageId="21" pageNumber="22">Tortopsis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFACFFCB7F67F932F472DFCB" box="[268,377,1712,1738]" pageId="21" pageNumber="22" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFACFFCB7EFCF971F667DE4B" pageId="21" pageNumber="22" type="reference_group">
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<paragraph id="8BE52311FFACFFCB7EFCF971F79FDE2B" blockId="21.[151,1436,1779,1930]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="AB4A043FFFACFFCB7EFCF971F79FDE2B" pageId="21" pageNumber="22">
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<taxonomicName id="4C5A5892FFACFFCB7EFCF971F4D5DE0B" ID-CoL="7XX3" authority="Needham & Murphy" authorityName="Needham & Murphy" box="[151,478,1779,1802]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B92EFF03FFACFFCB7EFCF971F5FEDE0B" box="[151,245,1779,1802]" italics="true" pageId="21" pageNumber="22">Tortopus</emphasis>
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Needham & Murphy
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</taxonomicName>
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(in part), 1924: 23; Ulmer (in part), 1933: 197; Traver (in part), 1950: 596;
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<treatmentCitation id="0AFB0500FFACFFCB7B70F971F43EDE2B" author="McCafferty" page="489" pageId="21" pageNumber="22" year="1975">
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<bibRefCitation id="EFCB5EE0FFACFFCB7B70F971F40BDE2B" author="McCafferty" pageId="21" pageNumber="34" refString="McCafferty, W. P. (1975) The burrowing mayflies (Ephemeroptera: Ephemeroidea) of the United States. Transactions of the American Entomological Society, 101, 447 - 504." type="journal article" year="1975">McCafferty, 1975</bibRefCitation>
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: 489
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</treatmentCitation>
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;
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<bibRefCitation id="EFCB5EE0FFACFFCB7F2BF891F4FBDE2B" author="Scott" box="[320,496,1811,1834]" pageId="21" pageNumber="35" refString="Scott, D. C., Berner, L., Hirsch, A. (1959) The nymph of the mayfly genus Tortopus (Ephemeroptera: Polymitarcidae). Annals of the Entomological Society of America, 52, 205 - 213." type="journal article" year="1959">Scott et al. 1959</bibRefCitation>
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;
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<bibRefCitation id="EFCB5EE0FFACFFCB7F90F891F79FDE2B" author="Molineri" box="[507,660,1811,1834]" pageId="21" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE52311FFACFFCB7EFCF8B1F667DE4B" blockId="21.[151,1436,1779,1930]" box="[151,876,1843,1866]" pageId="21" pageNumber="22">
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Type-species:
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<taxonomicName id="4C5A5892FFACFFCB7F5BF8B1F78DDE4B" authority="Ulmer" authorityName="Ulmer" box="[304,646,1843,1866]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="unguiculatus">
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<emphasis id="B92EFF03FFACFFCB7F5BF8B1F731DE4B" box="[304,570,1843,1866]" italics="true" pageId="21" pageNumber="22">Campsurus unguiculatus</emphasis>
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Ulmer
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</taxonomicName>
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, original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFACFFC87EFCF8D1F7EFDC89" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BE52311FFACFFCB7EFCF8D1F464DE8B" blockId="21.[151,1436,1779,1930]" pageId="21" pageNumber="22">
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Species composition:
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<taxonomicName id="4C5A5892FFACFFCB7FEBF8D1F718DE6B" box="[384,531,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bruchianus">
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<emphasis id="B92EFF03FFACFFCB7FEBF8D1F718DE6B" box="[384,531,1875,1898]" italics="true" pageId="21" pageNumber="22">T. bruchianus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5A5892FFACFFCB7C74F8D1F7BEDE6B" box="[543,693,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="limoncocha" status="sp. nov.">
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<emphasis id="B92EFF03FFACFFCB7C74F8D1F7BEDE6B" box="[543,693,1875,1898]" italics="true" pageId="21" pageNumber="22">T. limoncocha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFACFFCB7CD7F8D1F61ADE6B" box="[700,785,1875,1898]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C5A5892FFACFFCB7D77F8D1F6C1DE6B" box="[796,970,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="obscuripennis">
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<emphasis id="B92EFF03FFACFFCB7D77F8D1F6C1DE6B" box="[796,970,1875,1898]" italics="true" pageId="21" pageNumber="22">T. obscuripennis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5A5892FFACFFCB7DB3F8D1F137DE6B" box="[984,1084,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="parishi">
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<emphasis id="B92EFF03FFACFFCB7DB3F8D1F137DE6B" box="[984,1084,1875,1898]" italics="true" pageId="21" pageNumber="22">T. parishi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5A5892FFACFFCB7A20F8D1F1A6DE6B" box="[1099,1197,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="B92EFF03FFACFFCB7A20F8D1F1A6DE6B" box="[1099,1197,1875,1898]" italics="true" pageId="21" pageNumber="22">T. primus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5A5892FFACFFCB7AD7F8D1F01CDE6B" box="[1212,1303,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="puella">
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<emphasis id="B92EFF03FFACFFCB7AD7F8D1F01CDE6B" box="[1212,1303,1875,1898]" italics="true" pageId="21" pageNumber="22">T. puella</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5A5892FFACFFCB7B4DF8D1F070DE6B" box="[1318,1403,1875,1898]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sarae">
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<emphasis id="B92EFF03FFACFFCB7B4DF8D1F070DE6B" box="[1318,1403,1875,1898]" italics="true" pageId="21" pageNumber="22">T. sarae</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5A5892FFACFFCB7BE3F8D1F41DDE8B" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
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<emphasis id="B92EFF03FFACFFCB7BE3F8D1F41DDE8B" italics="true" pageId="21" pageNumber="22">T. spatula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFACFFCB7F77F8F1F464DE8B" box="[284,367,1907,1930]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BE52311FFACFFC87EFCF830F4C6DB75" blockId="21.[151,1436,1970,2034]" lastBlockId="22.[151,1437,152,1416]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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Adult: fore legs of male normal, remaining legs of male and all legs of female distorted and non-functional. Mesosternum: furcasternal plates approximating each other on basal half (
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<figureCitation id="13613F94FFACFFCB7D93F85AF146DEF3" box="[1016,1101,2008,2034]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="21" pageNumber="22">Fig. 54</figureCitation>
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) so furcasternal longitudinal impression narrow almost entirely, becoming wider only near posterior margin. Female pronotum with anterior ring well developed, at least 1/3 of total length of pronotum. Wings sexually dimorphic, all veins of female thickened; posterior margin of hind wings of female with a reticulated pattern of veinlets. Female fore wing without R3 and short intercalary vein before it (similar to
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<figureCitation id="13613F94FFAFFFC87DF9FE8AF6E7D823" box="[914,1004,264,290]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="22" pageNumber="23">Fig. 44</figureCitation>
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), in some large females one or both forewings may show 1 or 2 weak veins in this area, but their aspect is different (thinner, weaker) from the other longitudinal veins, as they are formed by the coalescence of few crossveins. Abdomen. Female abdominal sternum VIII with relatively large sublateral parastyli receptors, sockets opening towards the median line of the sternum (
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<figureCitation id="13613F94FFAFFFC87F92FE22F798D8BB" box="[505,659,416,442]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="22" pageNumber="23">Figs. 46, 51</figureCitation>
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). Male genitalia: abdominal sternum IX entire (
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<figureCitation id="13613F94FFAFFFC87A82FE22F04CD8BB" box="[1257,1351,416,442]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Fig. 57</figureCitation>
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); large parastyli present, curved dorsally, more than 5 times the length of their bases (pedestals); penes cylindrical, long and slender, completely divided (
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<figureCitation id="13613F94FFAFFFC87C3DFE68F7A5DB05" box="[598,686,490,516]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Fig. 57</figureCitation>
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); apex of penes with a hooked (or rarely blunt) sclerotized spine (
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<figureCitation id="13613F94FFAFFFC87EF4FD92F5FCDB2B" box="[159,247,528,554]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Fig. 59</figureCitation>
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). Penial arms formed by a single transverse bar which is articulated to the posterior corners of tergum IX. Forceps two-segmented, basal segment short and small, with a ventral knob (
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<figureCitation id="13613F94FFAFFFC87A1BFDB7F1DCDB4E" box="[1136,1239,565,591]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Figs. 57</figureCitation>
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); distal segment long, slender and clublike.
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</paragraph>
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<paragraph id="8BE52311FFAFFFC87EADFD02F420DA7B" blockId="22.[151,1437,152,1416]" pageId="22" pageNumber="23">
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Nymph (for a complete characterization see
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<bibRefCitation id="EFCB5EE0FFAFFFC87CB8FD02F689DB9B" author="Molineri" box="[723,898,640,666]" pageId="22" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
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): head with prominent frontal ridge (
|
||
<figureCitation id="13613F94FFAFFFC87B5EFD02F085DB9B" box="[1333,1422,640,666]" captionStart="FIGURES 93 – 96" captionStartId="30.[151,272,1872,1896]" captionTargetBox="[188,1398,197,1845]" captionTargetId="figure@30.[188,1398,197,1846]" captionTargetPageId="30" captionText="FIGURES 93 – 96. Polymitarcyidae, nymphal head: 93, Asthenopus sp.; 94, Campsurus near violaceus; 95, Tortopus harrisi; 96, Tortopsis obscuripennis." httpUri="https://zenodo.org/record/195446/files/figure.png" pageId="22" pageNumber="23">Fig. 96</figureCitation>
|
||
); fronto-clypeal region expanded and concave, extending ventrally beyond mandibular tusks; mandibular tusks long with a large subapical denticle on inner margin (
|
||
<figureCitation id="13613F94FFAFFFC87D4EFD48F68ADBE5" box="[805,897,714,740]" captionStart="FIGURES 93 – 96" captionStartId="30.[151,272,1872,1896]" captionTargetBox="[188,1398,197,1845]" captionTargetId="figure@30.[188,1398,197,1846]" captionTargetPageId="30" captionText="FIGURES 93 – 96. Polymitarcyidae, nymphal head: 93, Asthenopus sp.; 94, Campsurus near violaceus; 95, Tortopus harrisi; 96, Tortopsis obscuripennis." httpUri="https://zenodo.org/record/195446/files/figure.png" pageId="22" pageNumber="23">Fig. 96</figureCitation>
|
||
); maxilla with a basal finger like gill. Large filtering setae present on mouthparts and fore legs. Legs modified for burrowing with robust segments and flat tibiae. Fore tibia-tarsus with a prominent distal projection (dorsal projection 2/3 the length of the claw). Abdomen with vestigial unilamellate gill on segment I, gills on segment II–VII normal (large and bilamellate).
|
||
</paragraph>
|
||
<paragraph id="8BE52311FFAFFFC87EADFC07F7C0DD0E" blockId="22.[151,1437,152,1416]" pageId="22" pageNumber="23">
|
||
Eggs. Hemispherical, bowl-shaped (
|
||
<figureCitation id="13613F94FFAFFFC87C13FC07F600DA9E" box="[632,779,901,927]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="22" pageNumber="23">Figs. 79–80</figureCitation>
|
||
). Length, 355–450 µm; width, 290–375 µm. No polar caps or attachment structures; only one species (
|
||
<taxonomicName id="4C5A5892FFAFFFC87CA0FC29F630DAC5" box="[715,827,939,964]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
|
||
<emphasis id="B92EFF03FFAFFFC87CA0FC29F630DAC5" box="[715,827,939,964]" italics="true" pageId="22" pageNumber="23">T. spatula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D4278FFAFFFC87D2FFC28F694DAC5" box="[836,927,938,964]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) with a long filament coiled along the main axis of the egg (
|
||
<figureCitation id="13613F94FFAFFFC87F31FC52F4E4DAEB" box="[346,495,976,1002]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="22" pageNumber="23">Figs. 79–80</figureCitation>
|
||
). Chorion with microsculptures consisting of slightly elevated pentagonal and hexagonal cells, or shallow subcircular grooves.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FFAFFFC87EADFB98F7EFDC89" blockId="22.[151,1437,152,1416]" pageId="22" pageNumber="23">
|
||
Discussion and diagnosis. The adults of
|
||
<taxonomicName id="4C5A5892FFAFFFC87CF7FB99F60DDD35" box="[668,774,1051,1076]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92EFF03FFAFFFC87CF7FB99F60DDD35" box="[668,774,1051,1076]" italics="true" pageId="22" pageNumber="23">Tortopsis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D4278FFAFFFC87D65FB98F672DD35" box="[782,889,1050,1076]" pageId="22" pageNumber="23" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
can be separated from its sister taxon
|
||
<taxonomicName id="4C5A5892FFAFFFC87B5EFB99F097DD35" box="[1333,1436,1051,1076]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EFF03FFAFFFC87B5EFB99F097DD35" box="[1333,1436,1051,1076]" italics="true" pageId="22" pageNumber="23">Tortopus</emphasis>
|
||
</taxonomicName>
|
||
by: 1) mesofurcasternal plates approximating each other basally and medially, with inner margins diverging only on distal half (
|
||
<figureCitation id="13613F94FFAFFFC87F12FBE7F4D0DD7E" box="[377,475,1125,1151]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="22" pageNumber="23">Figs. 45</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FFAFFFC87F8DFBE7F70CDD7E" box="[486,519,1125,1151]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">54</figureCitation>
|
||
); 2) male abdominal sternum entire (
|
||
<figureCitation id="13613F94FFAFFFC87DDEFBE7F11CDD7E" box="[949,1047,1125,1151]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Figs. 57</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FFAFFFC87A49FBE7F148DD7E" box="[1058,1091,1125,1151]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="22" pageNumber="23">73</figureCitation>
|
||
); 3) parastyli very long, more than 5 times the length of pedestals (
|
||
<figureCitation id="13613F94FFAFFFC87C28FB08F7D8DDA5" box="[579,723,1162,1188]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Figs. 57–58</figureCitation>
|
||
); 4) penes cylindrical at least basally and completely divided (
|
||
<figureCitation id="13613F94FFAFFFC87ECBFB32F5F2DDCB" box="[160,249,1200,1226]" captionStart="FIGURES 53 – 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 – 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="22" pageNumber="23">Fig. 57</figureCitation>
|
||
); 5) female fore wing without R3 and short intercalary before it (
|
||
<figureCitation id="13613F94FFAFFFC87D97FB32F15EDDCB" box="[1020,1109,1200,1226]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="22" pageNumber="23">Fig. 44</figureCitation>
|
||
), rarely expressed as 1 or 2 weak short veins; 6) parastyli receptors on abdominal sternum VIII relatively large and sublateral in position (
|
||
<figureCitation id="13613F94FFAFFFC87ECBFB7CF439DC19" box="[160,306,1278,1304]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="22" pageNumber="23">Figs. 46–52</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FFAFFFC87F54FB7CF486DC19" box="[319,397,1278,1304]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="22" pageNumber="23">75, 78</figureCitation>
|
||
), openings towards medial line. As mentioned previously, the characters distiguishing
|
||
<taxonomicName id="4C5A5892FFAFFFC87EFCFAA1F5F5DC3D" box="[151,254,1315,1340]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EFF03FFAFFFC87EFCFAA1F5F5DC3D" box="[151,254,1315,1340]" italics="true" pageId="22" pageNumber="23">Tortopus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A5892FFAFFFC87F5DFAA1F4A9DC3D" box="[310,418,1315,1340]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EFF03FFAFFFC87F5DFAA1F4A9DC3D" box="[310,418,1315,1340]" italics="true" pageId="22" pageNumber="23">Tortopsis</emphasis>
|
||
</taxonomicName>
|
||
from the other genera of the family are listed in the phylogenetic section (apomorphies of node c). The coupling system in
|
||
<emphasis id="B92EFF03FFAFFFC87C5CFACBF7A1DC63" box="[567,682,1353,1378]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="4C5A5892FFAFFFC87C5CFACBF7ADDC63" box="[567,678,1353,1378]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Tortopsis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
as described in
|
||
<bibRefCitation id="EFCB5EE0FFAFFFC87D03FACAF1FADC63" author="McCafferty" box="[872,1265,1352,1378]" pageId="22" pageNumber="34" refString="McCafferty, W. P. & Bloodgood, D. W. (1989) The female and male coupling apparatus in Tortopus mayflies. Aquatic Insects, 11, 141 - 146." type="journal article" year="1989">McCafferty & Bloodgood (1989)</bibRefCitation>
|
||
, involves very well developed male parastyli and female sockets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |