197 lines
20 KiB
XML
197 lines
20 KiB
XML
<document id="5B9CEA96064CEF25FB5B4A2426A3EB2C" ID-CLB-Dataset="50775" ID-DOI="10.5281/zenodo.172420" ID-GBIF-Dataset="c7b02e9d-c3cb-4c3c-8510-1e3c9b65a53c" ID-ISSN="11755326" ID-Zenodo-Dep="172420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459756714995" checkinUser="plazi" docAuthor="Richards, Stephen J., Oliver, Paul, Dahl, Chris & Tjaturadi, Burhan" docDate="2006" docId="03F087FAFFFEFFBDFEA3FBDCF40F0B45" docLanguage="en" docName="zt01208p068.pdf" docOrigin="Zootaxa 1208" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Litoria infrafrenata" docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFC9FF82FFFFFFBFFFABFE30F0010823" masterDocTitle="A new species of large green treefrog (Anura: Hylidae: Litoria) from northern New Guinea" masterLastPageNumber="68" masterPageNumber="57" pageNumber="58" updateTime="1698218096950" updateUser="ExternalLinkService">
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<mods:titleInfo id="DAA768C2758757C17F9597676556BBA4">
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<mods:title id="B965397E770CBD8BE03B042261BEA30C">A new species of large green treefrog (Anura: Hylidae: Litoria) from northern New Guinea</mods:title>
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<mods:namePart id="ECB4CA2A6EB4402604CF9AB71E35C076">Richards, Stephen J.</mods:namePart>
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<mods:namePart id="EB17A92C847F135D301CC754A62EEE68">Oliver, Paul</mods:namePart>
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<mods:namePart id="E6F5F1650C9242EE3B568A5A1DC617F6">Dahl, Chris</mods:namePart>
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<mods:namePart id="7D7399C0FAFD18E2519CD5FBDF1BFE81">Tjaturadi, Burhan</mods:namePart>
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<mods:date id="E01269A0ABF6B07A5D11777741C6B315">2006</mods:date>
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<treatment id="03F087FAFFFEFFBDFEA3FBDCF40F0B45" ID-DOI="http://doi.org/10.5281/zenodo.5690287" ID-GBIF-Taxon="119343840" ID-Zenodo-Dep="5690287" LSID="urn:lsid:plazi:treatment:03F087FAFFFEFFBDFEA3FBDCF40F0B45" httpUri="http://treatment.plazi.org/id/03F087FAFFFEFFBDFEA3FBDCF40F0B45" lastPageId="2" lastPageNumber="58" pageId="1" pageNumber="58">
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<subSubSection id="C3436567FFFEFFBEFEA3FBDCF2AE0E24" box="[264,687,1516,1543]" pageId="1" pageNumber="58" type="nomenclature">
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<paragraph id="8BE636ECFFFEFFBEFEA3FBDCF2AE0E24" blockId="1.[264,687,1516,1543]" box="[264,687,1516,1543]" pageId="1" pageNumber="58">
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<heading id="D0AE8180FFFEFFBEFEA3FBDCF2AE0E24" bold="true" box="[264,687,1516,1543]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
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<emphasis id="B92DEAFEFFFEFFBEFEA3FBDCF2AE0E24" bold="true" box="[264,687,1516,1543]" pageId="1" pageNumber="58">
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Systematics of
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<taxonomicName id="4C594D6FFFFEFFBEFE69FBDDF2AE0E24" box="[450,687,1517,1543]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="infrafrenata">
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<emphasis id="B92DEAFEFFFEFFBEFE69FBDDF2AE0E24" bold="true" box="[450,687,1517,1543]" italics="true" pageId="1" pageNumber="58">Litoria infrafrenata</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3436567FFFEFFBDFEA3F808F40F0B45" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58" type="description">
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<paragraph id="8BE636ECFFFEFFBEFEA3F808F3B20EEA" blockId="1.[264,1324,1591,1897]" pageId="1" pageNumber="58">
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<taxonomicName id="4C594D6FFFFEFFBEFEA3F808F1EC0E72" box="[264,493,1592,1617]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="infrafrenata">
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<emphasis id="B92DEAFEFFFEFFBEFEA3F808F1EC0E72" box="[264,493,1592,1617]" italics="true" pageId="1" pageNumber="58">Litoria infrafrenata</emphasis>
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</taxonomicName>
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is a large green frog that is superficially similar to our new species and has a complicated taxonomic history. The new species is substantially smaller than
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<taxonomicName id="4C594D6FFFFEFFBEFAB8F850F1950E82" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="infrafrenata">
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<emphasis id="B92DEAFEFFFEFFBEFAB8F850F1950E82" italics="true" pageId="1" pageNumber="58">L. infrafrenata</emphasis>
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</taxonomicName>
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but exhibits a rather poorly defined off white labial stripe, suggesting there is some potential for confusion with juvenile
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<taxonomicName id="4C594D6FFFFEFFBEFCAAF880F3AD0EEA" box="[769,940,1712,1737]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="infrafrenata">
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<emphasis id="B92DEAFEFFFEFFBEFCAAF880F3AD0EEA" box="[769,940,1712,1737]" italics="true" pageId="1" pageNumber="58">L. infrafrenata</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BE636ECFFFEFFBDFE93F8E7F40F0B45" blockId="1.[264,1324,1591,1897]" lastBlockId="2.[264,1324,284,870]" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58">
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Based on published descriptions or examination of specimens, all names synonymised with
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<taxonomicName id="4C594D6FFFFEFFBEFEE5F930F2040F3A" box="[334,517,1792,1817]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="infrafrenata">
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<emphasis id="B92DEAFEFFFEFFBEFEE5F930F2040F3A" box="[334,517,1792,1817]" italics="true" pageId="1" pageNumber="58">L. infrafrenata</emphasis>
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</taxonomicName>
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represent animals clearly assignable to that taxon and easily distinguished from the new species by a number of morphological characters. The
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<typeStatus id="54E2884EFFFEFFBEFB66F917F50A0F62" box="[1229,1291,1831,1857]" pageId="1" pageNumber="58">types</typeStatus>
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of
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<taxonomicName id="4C594D6FFFFEFFBEFEA3F960F28D0F4A" authority="Cope, 1867" authorityName="Cope" authorityYear="1867" box="[264,652,1871,1897]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="dolichopsis">
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<emphasis id="B92DEAFEFFFEFFBEFEA3F960F1E60F4A" box="[264,487,1872,1897]" italics="true" pageId="1" pageNumber="58">Litoria dolichopsis</emphasis>
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(Cope, 1867)
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</taxonomicName>
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,
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<taxonomicName id="4C594D6FFFFEFFBEFD36F960F3430F4A" box="[669,834,1872,1897]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="1" pageNumber="58" phylum="Chordata" rank="species" species="dolichopsis">
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<emphasis id="B92DEAFEFFFEFFBEFD36F960F3430F4A" box="[669,834,1872,1897]" italics="true" pageId="1" pageNumber="58">L. dolichopsis</emphasis>
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</taxonomicName>
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<taxonomicName id="4C594D6FFFFEFFBEFCE0F97FF4F60F4A" authority="Boettger, 1895" authorityName="Boettger" authorityYear="1895" box="[843,1271,1871,1897]" pageId="1" pageNumber="58" rank="variety" variety="tenuigranulata">
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var.
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<emphasis id="B92DEAFEFFFEFFBEFCD4F960F42C0F4A" box="[895,1069,1872,1897]" italics="true" pageId="1" pageNumber="58">tenuigranulata</emphasis>
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(
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<bibRefCitation id="EFC84B1DFFFEFFBEFB96F97FF4EE0F4A" author="Boettger" box="[1085,1263,1871,1897]" pageId="1" pageNumber="66" refString="Boettger, O. (1895) Liste der Reptilien und Batrachier der Insel Halmaheira nach den Sammlungen Prof. Dr. W. Kukenthals's. Zoologischer Anzeiger, 472, 129 - 138." type="journal article" year="1895">Boettger, 1895</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C594D6FFFFDFFBDFEA3FF2CF2640915" authority="Boulenger, 1912" authorityName="Boulenger" authorityYear="1912" box="[264,613,284,310]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="spengeli">
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<emphasis id="B92DEAFEFFFDFFBDFEA3FF2CF1880916" box="[264,393,284,309]" italics="true" pageId="2" pageNumber="59">L. spengeli</emphasis>
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(Boulenger, 1912)
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</taxonomicName>
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are a minimum of
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<quantity id="4CA19B09FFFDFFBDFCEFFF2CF3970915" box="[836,918,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="59" unit="mm" value="90.0">90 mm</quantity>
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SV and often longer, substantially larger than the species described herein; furthermore the
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<typeStatus id="54E2884EFFFDFFBDFC10FF74F3EC097D" box="[955,1005,324,350]" pageId="2" pageNumber="59">type</typeStatus>
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of
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<taxonomicName id="4C594D6FFFFDFFBDFBB1FF74F4C0097E" box="[1050,1217,324,349]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="dolichopsis">
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<emphasis id="B92DEAFEFFFDFFBDFBB1FF74F4C0097E" box="[1050,1217,324,349]" italics="true" pageId="2" pageNumber="59">L. dolichopsis</emphasis>
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</taxonomicName>
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and one other synonym of
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<taxonomicName id="4C594D6FFFFDFFBDFE75FF5CF2C209A6" box="[478,707,364,389]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="infrafrenata">
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<emphasis id="B92DEAFEFFFDFFBDFE75FF5CF2C209A6" box="[478,707,364,389]" italics="true" pageId="2" pageNumber="59">Litoria infrafrenata</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="4C594D6FFFFDFFBDFD7FFF5CF41B09A5" authority="Macleay, 1878" authorityName="Macleay" authorityYear="1878" box="[724,1050,364,390]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="guttata">
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<emphasis id="B92DEAFEFFFDFFBDFD7FFF5CF34709A6" box="[724,838,364,389]" italics="true" pageId="2" pageNumber="59">L. guttata</emphasis>
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<bibRefCitation id="EFC84B1DFFFDFFBDFCE6FF5CF41B09A5" author="Macleay" box="[845,1050,364,390]" pageId="2" pageNumber="66" refString="Macleay W (1878) The Batrachians of the " Chevert " expedition. The Proceedings of the Linnean Society of New South Wales, 2, 135 - 138." type="journal article" year="1878">(Macleay, 1878))</bibRefCitation>
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</taxonomicName>
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, have partiallywebbed (vs fully webbed) hands and a prominent off white labial stripe (
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<bibRefCitation id="EFC84B1DFFFDFFBDFBA2FFA4F4B7098D" author="Macleay" box="[1033,1206,404,430]" pageId="2" pageNumber="66" refString="Macleay W (1878) The Batrachians of the " Chevert " expedition. The Proceedings of the Linnean Society of New South Wales, 2, 135 - 138." type="journal article" year="1878">Macleay 1878</bibRefCitation>
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,
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<bibRefCitation id="EFC84B1DFFFDFFBDFB6FFFA4F14409F5" author="Boettger" pageId="2" pageNumber="66" refString="Boettger, O. (1895) Liste der Reptilien und Batrachier der Insel Halmaheira nach den Sammlungen Prof. Dr. W. Kukenthals's. Zoologischer Anzeiger, 472, 129 - 138." type="journal article" year="1895">Boettger 1895</bibRefCitation>
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,
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<bibRefCitation id="EFC84B1DFFFDFFBDFEF8FF8CF1C309F5" author="Fry" box="[339,450,444,470]" pageId="2" pageNumber="66" refString="Fry, D. B. (1913) A re-examination of Macleay's New Guinea and Queensland frog types. Memoirs of the Queensland Museum, 2, 46 - 50." type="journal article" year="1913">Fry 1913</bibRefCitation>
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,
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<bibRefCitation id="EFC84B1DFFFDFFBDFE7BFF8CF28E09F5" author="Loveridge" box="[464,655,444,470]" pageId="2" pageNumber="66" refString="Loveridge, A. (1948) New Guinea Reptiles and Amphibians in the Museum of Comparitive Zoology and the United States National Museum. Bulletin of the Museum of Comparative Zoology, 101, 303 - 430." type="journal article" year="1948">Loveridge 1948</bibRefCitation>
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,
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<bibRefCitation id="EFC84B1DFFFDFFBDFD36FF8CF34909F5" author="Copland" box="[669,840,444,470]" pageId="2" pageNumber="66" refString="Copland, S. J. (1957) Australian treefrogs of the genus Hyla. Proceedings of the Linnean Society of New South Wales, 82, 99 - 108." type="journal article" year="1957">Copland 1957</bibRefCitation>
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). The males in the
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<typeStatus id="54E2884EFFFDFFBDFB84FF8CF46009F5" box="[1071,1121,444,470]" pageId="2" pageNumber="59">type</typeStatus>
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series of
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<taxonomicName id="4C594D6FFFFDFFBDFB72FF8CF18F09DE" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="dolichopsis">
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<emphasis id="B92DEAFEFFFDFFBDFB72FF8CF18F09DE" italics="true" pageId="2" pageNumber="59">Litoria dolichopsis</emphasis>
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</taxonomicName>
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<taxonomicName id="4C594D6FFFFDFFBDFE34FFD4F31609DD" authority="Werner, 1898" authorityName="Werner" authorityYear="1898" box="[415,791,484,510]" pageId="2" pageNumber="59" rank="variety" variety="pollicaris">
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var.
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<emphasis id="B92DEAFEFFFDFFBDFE70FFD4F24A09DE" box="[475,587,484,509]" italics="true" pageId="2" pageNumber="59">pollicaris</emphasis>
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(
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<bibRefCitation id="EFC84B1DFFFDFFBDFDCFFFD4F30E09DD" author="Werner" box="[612,783,484,510]" pageId="2" pageNumber="66" refString="Werner, F. (1898) Vorlaufige Mittheilung uber die von Herrn Prof. F. Dahl im Bismarck-archipel gesammelten Reptilien und Batrachier. Zoologischer Anzeiger, 21, 552 - 556." type="journal article" year="1898">Werner, 1898</bibRefCitation>
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)
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</taxonomicName>
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have a very hard and protruding pollex rudiment, absent in the new species (
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<bibRefCitation id="EFC84B1DFFFDFFBDFD60FC3CF36D0A05" author="Werner" box="[715,876,524,550]" pageId="2" pageNumber="66" refString="Werner, F. (1898) Vorlaufige Mittheilung uber die von Herrn Prof. F. Dahl im Bismarck-archipel gesammelten Reptilien und Batrachier. Zoologischer Anzeiger, 21, 552 - 556." type="journal article" year="1898">Werner 1898</bibRefCitation>
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). In addition this taxon is from New
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<collectingCountry id="F34E767CFFFDFFBDFEA3FC04F1580A6D" box="[264,345,564,590]" name="United Kingdom" pageId="2" pageNumber="59">Britain</collectingCountry>
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where survey work has found no evidence of any
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<taxonomicName id="4C594D6FFFFDFFBDFC0EFC04F3F60A6E" box="[933,1015,564,589]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="genus">
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<emphasis id="B92DEAFEFFFDFFBDFC0EFC04F3F60A6E" box="[933,1015,564,589]" italics="true" pageId="2" pageNumber="59">Litoria</emphasis>
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</taxonomicName>
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similar to the new species (Richards pers. obs.). The name
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<taxonomicName id="4C594D6FFFFDFFBDFD0BFC6CF3820A56" box="[672,899,604,629]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="dolichopsis">
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<emphasis id="B92DEAFEFFFDFFBDFD0BFC6CF3820A56" box="[672,899,604,629]" italics="true" pageId="2" pageNumber="59">Litoria dolichopsis</emphasis>
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</taxonomicName>
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<taxonomicName id="4C594D6FFFFDFFBDFC3BFC6CF4F10A55" authority="Vogt, 1912" authorityName="Vogt" authorityYear="1912" box="[912,1264,604,630]" pageId="2" pageNumber="59" rank="variety" variety="calcarifera">
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var.
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<emphasis id="B92DEAFEFFFDFFBDFC63FC6CF44B0A56" box="[968,1098,604,629]" italics="true" pageId="2" pageNumber="59">calcarifera</emphasis>
|
||
(Vogt, 1912)
|
||
</taxonomicName>
|
||
was apparently applied erroneously to specimens from the
|
||
<typeStatus id="54E2884EFFFDFFBDFC1AFCB4F3E20ABD" box="[945,995,644,670]" pageId="2" pageNumber="59">type</typeStatus>
|
||
series of
|
||
<emphasis id="B92DEAFEFFFDFFBDFBC1FCB4F52B0ABE" box="[1130,1322,644,669]" italics="true" pageId="2" pageNumber="59">L. d. pollicaris,</emphasis>
|
||
therefore it is regarded as a lapsus (
|
||
<bibRefCitation id="EFC84B1DFFFDFFBDFD10FC9CF4570AE5" author="Zweifel" box="[699,1110,684,710]" pageId="2" pageNumber="66" refString="Zweifel, R. G. (1960) Results of the 1958 - 1959 Archbold Expedition 3. Notes on the frogs of New Britain. American Museum Novitates, 2023, 1 - 27." type="journal article" year="1960">Zweifel 1960, Günther pers. com.</bibRefCitation>
|
||
); in addition, as mentioned above, the specimens involved clearly represent
|
||
<taxonomicName id="4C594D6FFFFDFFBDFC79FCE4F4810ACE" box="[978,1152,724,749]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="infrafrenata">
|
||
<emphasis id="B92DEAFEFFFDFFBDFC79FCE4F4810ACE" box="[978,1152,724,749]" italics="true" pageId="2" pageNumber="59">L. infrafrenata</emphasis>
|
||
</taxonomicName>
|
||
. Finally
|
||
<bibRefCitation id="EFC84B1DFFFDFFBDFB46FCE4F1570B35" author="Tyler" pageId="2" pageNumber="66" refString="Tyler, M. J. (1968) Papuan Hylid Frogs of the genus Hyla. Zoologische Verhandelingen, 96, 1 - 203." type="journal article" year="1968">Tyler (1968)</bibRefCitation>
|
||
and Günther (pers. comm.) have both examined the
|
||
<typeStatus id="54E2884EFFFDFFBDFC43FCCCF4270B35" box="[1000,1062,764,790]" pageId="2" pageNumber="59">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C594D6FFFFDFFBDFBF3FCCCF1900B1D" authority="Ahl, 1929" authorityName="Ahl" authorityYear="1929" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="trinilensis">
|
||
<emphasis id="B92DEAFEFFFDFFBDFBF3FCCCF52B0B36" box="[1112,1322,764,789]" italics="true" pageId="2" pageNumber="59">Litoria trinilensis</emphasis>
|
||
(Ahl, 1929)
|
||
</taxonomicName>
|
||
and found them to possess reduced webbing between the fingers, like
|
||
<taxonomicName id="4C594D6FFFFDFFBDFB72FD14F1950B46" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="infrafrenata">
|
||
<emphasis id="B92DEAFEFFFDFFBDFB72FD14F1950B46" italics="true" pageId="2" pageNumber="59">Litoria infrafrenata</emphasis>
|
||
</taxonomicName>
|
||
and unlike
|
||
<taxonomicName id="4C594D6FFFFDFFBDFDB5FD7CF2AC0B46" box="[542,685,844,869]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="B92DEAFEFFFDFFBDFDB5FD7CF2AC0B46" box="[542,685,844,869]" italics="true" pageId="2" pageNumber="59">L. graminea</emphasis>
|
||
</taxonomicName>
|
||
or the new species from Utai.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |