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<document id="D6C585E61D5CEC6CCF4CCB3CBCA75D68" ID-CLB-Dataset="55344" ID-DOI="10.5852/ejt.2023.880.2173" ID-GBIF-Dataset="339bce42-a7e7-418b-b9ac-4588e4219b15" ID-ISSN="2118-9773" ID-Zenodo-Dep="8155472" ID-ZooBank="3F806FD6-2EB3-456A-AFD7-780A0FBEB2DA" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1689603386695" checkinUser="felipe" docAuthor="Huber, Bernhard A., Meng, Guanliang, Valdez-Mondragón, Alejandro, Král, Jiří, Ávila Herrera, Ivalú M. &amp; Carvalho, Leonardo S." docDate="2023" docId="03F087F1AC6AFFC8095D6130E23AFDE0" docLanguage="en" docName="ejt-880_1-89.pdf" docOrigin="European Journal of Taxonomy 880 (1)" docSource="http://zoobank.org/3f806fd6-2eb3-456a-afd7-780a0fbeb2da" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pholcophora texana Gertsch 1935" docType="treatment" docVersion="6" lastPageNumber="31" masterDocId="FFC9FF89AC76FFD60B146426E305FFD0" masterDocTitle="Short-legged daddy-long-leg spiders in North America: the genera Pholcophora and Tolteca (Araneae, Pholcidae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="29" updateTime="1699540719498" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8DEFD889C936230210479F5780B0EEEE">Short-legged daddy-long-leg spiders in North America: the genera Pholcophora and Tolteca (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="2815619FE6431EF68CE2643FC2548F5B">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="56D6A767710393CC01A0669871C11173">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany. Laboratory of Arachnology (LATLAX), Institute of Biology, Universidad Nacional Autónoma de México-Tlaxcala, San Miguel Contla, Santa Cruz Tlaxcala, Tlaxcala, México. Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic. Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil.</mods:affiliation>
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<mods:namePart id="9A27B30817E3C6F93C5909C9E3E80559">Meng, Guanliang</mods:namePart>
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<mods:affiliation id="37450E559D6EE39404E5ABC3C138BDDC">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany. Laboratory of Arachnology (LATLAX), Institute of Biology, Universidad Nacional Autónoma de México-Tlaxcala, San Miguel Contla, Santa Cruz Tlaxcala, Tlaxcala, México. Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic. Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil.</mods:affiliation>
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<mods:namePart id="BEE65B7050306F56CED8055822B657D7">Valdez-Mondragón, Alejandro</mods:namePart>
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<mods:namePart id="C4F5E00E1A48A048573CE48B81BD7084">Král, Jiří</mods:namePart>
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<treatment id="03F087F1AC6AFFC8095D6130E23AFDE0" ID-DOI="http://doi.org/10.5281/zenodo.8160850" ID-GBIF-Taxon="211663644" ID-Zenodo-Dep="8160850" LSID="urn:lsid:plazi:treatment:03F087F1AC6AFFC8095D6130E23AFDE0" httpUri="http://treatment.plazi.org/id/03F087F1AC6AFFC8095D6130E23AFDE0" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C343656CAC6AFFCA095D6130E0EEFAE1" box="[585,1003,1302,1329]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BE636E7AC6AFFCA095D6130E0EEFAE1" blockId="28.[585,1003,1302,1365]" box="[585,1003,1302,1329]" pageId="28" pageNumber="29">
<heading id="D0AE818BAC6AFFCA095D6130E0EEFAE1" box="[585,1003,1302,1329]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="4C594D64AC6AFFCA095D6130E0EEFAE1" ID-CoL="4GKZR" authority="Gertsch, 1935" authorityName="Gertsch" authorityYear="1935" box="[585,1003,1302,1329]" class="Arachnida" family="Pholcidae" genus="Pholcophora" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="B92DEAF5AC6AFFCA095D6130E03BFAE0" bold="true" box="[585,830,1302,1329]" italics="true" pageId="28" pageNumber="29">Pholcophora texana</emphasis>
Gertsch, 1935
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<subSubSection id="C343656CAC6AFFCA099F611CE0ADFA85" box="[651,936,1338,1365]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BE636E7AC6AFFCA099F611CE0ADFA85" blockId="28.[585,1003,1302,1365]" box="[651,936,1338,1365]" pageId="28" pageNumber="29">
<figureCitation id="13622A62AC6AFFCA099F611CE006FA85" box="[651,771,1338,1365]" captionStart="Fig" captionStartId="12.[189,232,1822,1848]" captionTargetBox="[189,1398,566,1776]" captionTargetId="figure-121@12.[189,1398,566,1776]" captionTargetPageId="12" captionText="Fig. 3. Pholcophora Banks, 1896, live specimens. A. P. americana Banks, 1896, male from USA, Colorado, near Golden.B–E. P. mazatlan Huber sp. nov., males and females with egg-sacs from Mexico, Guerrero, N of Mazatlán. F–G. P. papanoa Huber sp. nov., male and female from Mexico, Guerrero, S of Papanoa. H–I. P. texana Gertsch, 1935, male and female with egg-sac from Mexico, Hidalgo, SW of Jacala. J–K. P. tehuacan Huber sp. nov., male and female with egg-sac from Mexico, Puebla, SE of Tehuacan. L. P. “Mex354”, female with egg-sac from Mexico, Puebla, SE of Tehuacan." figureDoi="http://doi.org/10.5281/zenodo.8155480" httpUri="https://zenodo.org/record/8155480/files/figure.png" pageId="28" pageNumber="29">Figs 3H–I</figureCitation>
,
<figureCitation id="13622A62AC6AFFCA0804611DE05EFA85" box="[784,859,1339,1365]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="29.[189,232,1928,1954]" captionStartId-1="30.[189,232,1892,1918]" captionStartId-2="31.[189,232,909,935]" captionStartId-3="31.[189,232,1891,1917]" captionStartId-4="32.[189,232,1754,1780]" captionStartId-5="33.[189,232,1766,1792]" captionTargetBox-0="[189,1398,1145,1879]" captionTargetBox-1="[191,1398,739,1848]" captionTargetBox-2="[189,1398,265,869]" captionTargetBox-3="[189,1398,1079,1846]" captionTargetBox-4="[212,1376,265,1711]" captionTargetBox-5="[189,1398,266,1709]" captionTargetId-0="figure-228@29.[189,1398,1145,1879]" captionTargetId-1="figure-78@30.[189,1398,739,1848]" captionTargetId-2="figure-11@31.[189,1398,265,869]" captionTargetId-3="figure-12@31.[189,1398,1079,1846]" captionTargetId-4="figure-12@32.[212,1376,265,1711]" captionTargetId-5="figure-11@33.[189,1398,266,1709]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="31" captionTargetPageId-4="32" captionTargetPageId-5="33" captionText-0="Fig. 19. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). Left palp, prolateral, dorsal, and retrolateral views; arrows point at unique brush of modified hairs. Scale bar = 0.3 mm." captionText-1="Fig. 20. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Chelicerae, frontal and lateral views. C–E. Left palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. F–H. Left genital bulb, prolateral, dorsal, and retrolateral views. Scale bars = 0.2 mm." captionText-2="Fig. 21. Pholcophora texana Gertsch, 1935, female from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952), epigynum. A. Ventral view. B. Lateral view. Scale bar = 0.3 mm (both at same scale)." captionText-3="Fig. 22. Pholcophora texana Gertsch, 1935, females from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Cleared female genitalia, ventral and dorsal views. C–D. Detail of median internal structures in two different specimens (slightly different angles of view). Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." captionText-4="Fig. 23. Pholcophora texana Gertsch, 1935, female from Mexico, Hidalgo, 2.5 km SW of Jacala (ZFMK Mex341). A. Ocular area, dorsal view. B. Lateral face of chelicera, showing absence of stridulatory file. C. Epigynum, ventral (slightly posterior) view. D. ALS. E. PMS and anal hairs (arrows). F. Tip of left palpal tarsus, dorsal view. G. Detail of right palpal tarsus, dorsal view. H. Palpal tarsal organ. Abbreviations: ep = epigynum (main epigynal plate); pep = posterior epigynal plate; to = tarsal organ. Scale bars: A–B = 20 µm; C = 100 µm; D–G = 10 µm; H = 2 µm." captionText-5="Fig. 24. Pholcophora texana Gertsch, 1935, female from Mexico, Hidalgo, 2.5 km SW of Jacala (ZFMK Mex341). A. Detail of femur 2, retrolateral view. B. Detail of tibia 4, retrolateral view. C. Patella-tibia 1 joint, retrolateral view. D. Metatarsus-tarsus 2 joint, retrolateral view. E. Retrolateral trichobothrium of tibia 1. F. Detail of tarsus 1, showing pseudosegmentation. G. Tip of tarsus 2, retrolateral view. H. Tip of tarsus 4, prolateral view, showing comb-hair (arrow). Scale bars: A–B, D–H = 10 µm; C = 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.8155532" figureDoi-1="http://doi.org/10.5281/zenodo.8155534" figureDoi-2="http://doi.org/10.5281/zenodo.8155536" figureDoi-3="http://doi.org/10.5281/zenodo.8155538" figureDoi-4="http://doi.org/10.5281/zenodo.8155540" figureDoi-5="http://doi.org/10.5281/zenodo.8155542" httpUri-0="https://zenodo.org/record/8155532/files/figure.png" httpUri-1="https://zenodo.org/record/8155534/files/figure.png" httpUri-2="https://zenodo.org/record/8155536/files/figure.png" httpUri-3="https://zenodo.org/record/8155538/files/figure.png" httpUri-4="https://zenodo.org/record/8155540/files/figure.png" httpUri-5="https://zenodo.org/record/8155542/files/figure.png" pageId="28" pageNumber="29">19–24</figureCitation>
,
<figureCitation id="13622A62AC6AFFCA087C611DE0ADFA85" box="[872,936,1339,1365]" captionStart="Fig" captionStartId="43.[189,232,1681,1707]" captionTargetBox="[246,1335,267,1633]" captionTargetId="figure-12@43.[246,1342,265,1633]" captionTargetPageId="43" captionText="Fig. 33. Pholcophora internal female genitalia, median sections of main transversal internal sclerite. A–B. P. americana Banks, 1896, females from USA, California, Inyo Nat. Forest (ZFMK G089) and Colorado, near Golden (ZFMK USA16). C–D. P. mazatlan Huber sp. nov., females from Mexico, Guerrero, N of Mazatlán (ZFMK Ar 23943). E–F. P. papanoa Huber sp. nov., females from Mexico, Guerrero, S of Papanoa (ZFMK Ar 23945). G–H. P. “Mex354”, females from Mexico, Puebla, SE of Tehuacan (ZFMK Mex354). I–J. P. texana Gertsch, 1935, females from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). K–L. P. tehuacan Huber sp. nov., females from Mexico, Puebla, SE of Tehuacan (ZFMK Ar 23949). M. Pholcophora? “Car544”, female from Puerto Rico, Isla Mono (USNM ENT 783464). N. Pholcophora? “Cu12-325”, female from Cuba, Santiago de Cuba, Siboney (ZFMK Cu12- 325). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155572" httpUri="https://zenodo.org/record/8155572/files/figure.png" pageId="28" pageNumber="29">33I–J</figureCitation>
</paragraph>
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<paragraph id="8BE636E7AC6AFFCA0BA9615BE221FA47" blockId="28.[189,292,1405,1431]" box="[189,292,1405,1431]" pageId="28" pageNumber="29">
<heading id="D0AE818BAC6AFFCA0BA9615BE221FA47" bold="true" box="[189,292,1405,1431]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B92DEAF5AC6AFFCA0BA9615BE221FA47" bold="true" box="[189,292,1405,1431]" pageId="28" pageNumber="29">Remark</emphasis>
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<paragraph id="8BE636E7AC6AFFCA0BA9618BE0CCFA17" blockId="28.[189,969,1452,1479]" box="[189,969,1452,1479]" pageId="28" pageNumber="29">
For synonymy,
<typeStatus id="54E28845AC6AFFCA0A60618BE2A2FA17" box="[372,423,1453,1479]" pageId="28" pageNumber="29">type</typeStatus>
material, and redescription, see
<bibRefCitation id="EFC84B16AC6AFFCA0837618AE0C3FA17" author="Huber B. A." box="[803,966,1452,1479]" pageId="28" pageNumber="29" pagination="1 - 348" refId="ref34125" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / brh 26 h" type="journal article" year="2000">Huber (2000)</bibRefCitation>
.
</paragraph>
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<subSubSection id="C343656CAC6AFFCA0BA961C9E781F976" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BE636E7AC6AFFCA0BA961C9E233F9D9" blockId="28.[189,310,1519,1545]" box="[189,310,1519,1545]" pageId="28" pageNumber="29">
<heading id="D0AE818BAC6AFFCA0BA961C9E233F9D9" bold="true" box="[189,310,1519,1545]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B92DEAF5AC6AFFCA0BA961C9E233F9D9" bold="true" box="[189,310,1519,1545]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636E7AC6AFFCA0BA96239E781F976" blockId="28.[189,1398,1567,1702]" pageId="28" pageNumber="29">
Easily distinguished from known congeners by unique brush of modified hairs on male palpal femur (
<figureCitation id="13622A62AC6AFFCA0BD26265E25CF98D" box="[198,345,1603,1629]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,1145,1879]" captionTargetId="figure-228@29.[189,1398,1145,1879]" captionTargetPageId="29" captionText="Fig. 19. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). Left palp, prolateral, dorsal, and retrolateral views; arrows point at unique brush of modified hairs. Scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8155532" httpUri="https://zenodo.org/record/8155532/files/figure.png" pageId="28" pageNumber="29">Fig. 19A, C</figureCitation>
) and by pair of round sacs in female internal genitalia (
<figureCitation id="13622A62AC6AFFCA0F106265E79DF98D" box="[1028,1176,1603,1629]" captionStart="Fig" captionStartId="31.[189,232,1891,1917]" captionTargetBox="[189,1398,1079,1846]" captionTargetId="figure-12@31.[189,1398,1079,1846]" captionTargetPageId="31" captionText="Fig. 22. Pholcophora texana Gertsch, 1935, females from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Cleared female genitalia, ventral and dorsal views. C–D. Detail of median internal structures in two different specimens (slightly different angles of view). Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155538" httpUri="https://zenodo.org/record/8155538/files/figure.png" pageId="28" pageNumber="29">Fig. 22B–D</figureCitation>
); also by shape of procursus (
<figureCitation id="13622A62AC6AFFCA0A2B6241E2CEF951" box="[319,459,1639,1665]" captionStart="Fig" captionStartId="30.[189,232,1892,1918]" captionTargetBox="[191,1398,739,1848]" captionTargetId="figure-78@30.[189,1398,739,1848]" captionTargetPageId="30" captionText="Fig. 20. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Chelicerae, frontal and lateral views. C–E. Left palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. F–H. Left genital bulb, prolateral, dorsal, and retrolateral views. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8155534" httpUri="https://zenodo.org/record/8155534/files/figure.png" pageId="28" pageNumber="29">Fig. 20C–E</figureCitation>
; tip in lateral view with wide transparent flap), by shape of distal bulbal process (
<figureCitation id="13622A62AC6AFFCA0BD262ADE257F976" box="[198,338,1675,1702]" captionStart="Fig" captionStartId="30.[189,232,1892,1918]" captionTargetBox="[191,1398,739,1848]" captionTargetId="figure-78@30.[189,1398,739,1848]" captionTargetPageId="30" captionText="Fig. 20. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Chelicerae, frontal and lateral views. C–E. Left palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. F–H. Left genital bulb, prolateral, dorsal, and retrolateral views. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8155534" httpUri="https://zenodo.org/record/8155534/files/figure.png" pageId="28" pageNumber="29">Fig. 20F–H</figureCitation>
), and by indistinct plate in front of main epigynal plate (
<figureCitation id="13622A62AC6AFFCA08F362ADE773F976" box="[999,1142,1675,1702]" captionStart="Fig" captionStartId="31.[189,232,909,935]" captionTargetBox="[189,1398,265,869]" captionTargetId="figure-11@31.[189,1398,265,869]" captionTargetPageId="31" captionText="Fig. 21. Pholcophora texana Gertsch, 1935, female from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952), epigynum. A. Ventral view. B. Lateral view. Scale bar = 0.3 mm (both at same scale)." figureDoi="http://doi.org/10.5281/zenodo.8155536" httpUri="https://zenodo.org/record/8155536/files/figure.png" pageId="28" pageNumber="29">Fig. 21A, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C343656CAC6AFFCA0BA962E8E1B2F879" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BE636E7AC6AFFCA0BA962E8E14DF938" blockId="28.[189,584,1742,1768]" box="[189,584,1742,1768]" pageId="28" pageNumber="29">
<emphasis id="B92DEAF5AC6AFFCA0BA962E8E2AFF938" bold="true" box="[189,426,1742,1768]" pageId="28" pageNumber="29">Material examined</emphasis>
(new record)
</paragraph>
<paragraph id="8BE636E7AC6AFFCA0BA962D8E1B2F879" blockId="28.[189,1400,1789,1961]" pageId="28" pageNumber="29">
<materialsCitation id="3B313CBAAC6AFFCA0BA962D8E7C0F8ED" ID-GBIF-Occurrence="4161533336" collectingDate="2019-10-21" collectionCode="ZFMK" collectorName="B. A. Huber &amp; A. Valdez-Mondragon" country="Mexico" elevation="1430" latitude="20.9948" location="Jacala" longLatPrecision="7" longitude="-99.2138" pageId="28" pageNumber="29" specimenCount="5" specimenCount-female="1" specimenCount-male="1" stateProvince="Hidalgo">
<collectingCountry id="F34E7677AC6AFFCA0BA962D8E234F8C8" box="[189,305,1790,1816]" name="Mexico" pageId="28" pageNumber="29">MEXICO</collectingCountry>
–
<emphasis id="B92DEAF5AC6AFFCA0A5B62DBE2B7F8C7" bold="true" box="[335,434,1789,1815]" pageId="28" pageNumber="29">
<collectingRegion id="499DF805AC6AFFCA0A5B62DBE2B7F8C7" box="[335,434,1789,1815]" country="Mexico" name="Hidalgo" pageId="28" pageNumber="29">Hidalgo</collectingRegion>
</emphasis>
•
<specimenCount id="9D5FFD6EAC6AFFCA0AD862D8E2FFF8C8" box="[460,506,1790,1816]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5FFD6EAC6AFFCA091362D8E136F8C8" box="[519,563,1790,1816]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D5FFD6EAC6AFFCA095662D8E1ACF8C7" box="[578,681,1789,1816]" pageId="28" pageNumber="29" type="female">3 female</specimenCount>
abdomens; ~
<quantity id="4CA19B02AC6AFFCA085262DBE09DF8C7" box="[838,920,1789,1816]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="29" unit="km" value="2.5">2.5 km</quantity>
SW of
<location id="8E86603CAC6AFFCA08E762DBE73BF8C7" LSID="urn:lsid:plazi:treatment:03F087F1AC6AFFC8095D6130E23AFDE0:8E86603CAC6AFFCA08E762DBE73BF8C7" box="[1011,1086,1789,1815]" country="Mexico" latitude="20.9948" longLatPrecision="7" longitude="-99.2138" name="Jacala" pageId="28" pageNumber="29" stateProvince="Hidalgo">Jacala</location>
;
<geoCoordinate id="EE6D5020AC6AFFCA0F5F62D8E7D0F8C8" box="[1099,1237,1790,1816]" degrees="20.9948" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="5" value="20.9948">20.9948° N</geoCoordinate>
,
<geoCoordinate id="EE6D5020AC6AFFCA0FF562D8E677F8C8" box="[1249,1394,1790,1816]" degrees="99.2138" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-99.2138">99.2138° W</geoCoordinate>
;
<elevation id="0074D1D4AC6AFFCA0BA96304E256F8EC" box="[189,339,1826,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="28" pageNumber="29" unit="m" value="1430.0">
<quantity id="4CA19B02AC6AFFCA0BA96304E210F8EC" box="[189,277,1826,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="28" pageNumber="29" unit="m" value="1430.0">1430 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE71027AC6AFFCA0A4B6304E2FDF8EC" box="[351,504,1826,1852]" pageId="28" pageNumber="29" value="2019-10-21">
<collectingDate id="EFA3E9CFAC6AFFCA0A4B6304E2FDF8EC" box="[351,504,1826,1852]" pageId="28" pageNumber="29" value="2019-10-21">21 Oct. 2019</collectingDate>
</date>
;
<collectorName id="26AC5331AC6AFFCA09176304E18CF8EC" box="[515,649,1826,1852]" pageId="28" pageNumber="29">B.A. Huber</collectorName>
and
<collectorName id="26AC5331AC6AFFCA09AA6304E0C7F8EC" box="[702,962,1826,1852]" pageId="28" pageNumber="29">A. Valdez-Mondragón</collectorName>
leg.;
<collectionCode id="ED48AE22AC6AFFCA0F166304E756F8EC" box="[1026,1107,1826,1852]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="28" pageNumber="29" type="Museum">ZFMK</collectionCode>
<collectorName id="26AC5331AC6AFFCA0F436304E773F8EC" box="[1111,1142,1826,1852]" pageId="28" pageNumber="29">Ar</collectorName>
23952
</materialsCitation>
•
<materialsCitation id="3B313CBAAC6AFFCA0FC86304E0B1F854" ID-GBIF-Occurrence="4161533350" collectingDate="2019-10-21" collectionCode="ZFMK" collectorName="B. A. Huber &amp; A. Valdez-Mondragon" country="Mexico" elevation="1430" latitude="20.9948" location="Jacala" longLatPrecision="7" longitude="-99.2138" pageId="28" pageNumber="29" specimenCount="9" specimenCount-female="7" specimenCount-juvenile="2" stateProvince="Hidalgo">
<specimenCount id="9D5FFD6EAC6AFFCA0FC86304E618F8EC" box="[1244,1309,1826,1852]" pageId="28" pageNumber="29" type="female">7 ♀♀</specimenCount>
, 2 juvs, in pure ethanol (
<specimenCount id="9D5FFD6EAC6AFFCA0A956360E113F8B0" box="[385,534,1862,1888]" pageId="28" pageNumber="29" type="female">three female</specimenCount>
prosomata used for molecular work, abdomens transferred to
<collectionCode id="ED48AE22AC6AFFCA0FEB6361E655F8B1" box="[1279,1360,1863,1889]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="28" pageNumber="29" type="Museum">ZFMK</collectionCode>
Ar 23952); same collection data as for preceding;
<collectionCode id="ED48AE22AC6AFFCA09E3634DE04DF855" box="[759,840,1899,1925]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="28" pageNumber="29" type="Museum">ZFMK</collectionCode>
Mex341
</materialsCitation>
•
<materialsCitation id="3B313CBAAC6AFFCA08C7634CE1B6F879" ID-GBIF-Occurrence="4161533347" collectingDate="2019-10-21" collectionCode="LATLAX" collectorName="B. A. Huber &amp; A. Valdez-Mondragon" country="Mexico" elevation="1430" latitude="20.9948" location="Jacala" longLatPrecision="7" longitude="-99.2138" pageId="28" pageNumber="29" specimenCount="8" specimenCount-female="6" specimenCount-juvenile="2" stateProvince="Hidalgo">
<specimenCount id="9D5FFD6EAC6AFFCA08C7634CE71CF855" box="[979,1049,1898,1925]" pageId="28" pageNumber="29" type="female">6 ♀♀</specimenCount>
, 2 juvs (subadult
<specimenCount id="9D5FFD6EAC6AFFCA0FE0634CE621F854" box="[1268,1316,1898,1924]" pageId="28" pageNumber="29" type="male">♂♂</specimenCount>
); same collection data as for preceding;
<collectionCode id="ED48AE22AC6AFFCA092963A9E1B6F879" box="[573,691,1935,1961]" pageId="28" pageNumber="29">LATLAX</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C343656CAC6BFFCB0BA9652FE1A1FC5C" pageId="29" pageNumber="30" type="description">
<paragraph id="8BE636E7AC6BFFCB0BA9652FE1C0FEF3" blockId="29.[189,709,265,291]" box="[189,709,265,291]" pageId="29" pageNumber="30">
<emphasis id="B92DEAF5AC6BFFCB0BA9652FE24BFEF3" bold="true" box="[189,334,265,291]" pageId="29" pageNumber="30">Description</emphasis>
(amendments; see
<bibRefCitation id="EFC84B16AC6BFFCB0924652FE1B9FEF3" author="Huber B. A." box="[560,700,265,291]" pageId="29" pageNumber="30" pagination="1 - 348" refId="ref34125" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / brh 26 h" type="journal article" year="2000">Huber 2000</bibRefCitation>
)
</paragraph>
<paragraph id="8BE636E7AC6BFFCB0BFE651EE22FFE82" blockId="29.[188,1401,312,698]" box="[234,298,312,338]" pageId="29" pageNumber="30">
<emphasis id="B92DEAF5AC6BFFCB0BFE651EE22FFE82" bold="true" box="[234,298,312,338]" pageId="29" pageNumber="30">Male</emphasis>
</paragraph>
<paragraph id="8BE636E7AC6BFFCB0BA9657BE7A7FD6A" blockId="29.[188,1401,312,698]" pageId="29" pageNumber="30">
Measurements of male from SW of Jacala,
<collectingRegion id="499DF805AC6BFFCB09C6657AE034FEA6" box="[722,817,348,374]" country="Mexico" name="Hidalgo" pageId="29" pageNumber="30">Hidalgo</collectingRegion>
(ZFMK Ar 23952): body length 1.75, carapace width 0.70; tibia 1 length 1.25, metatarsus 1 length 1.30, tibia 2 length 0.95, tibia 4 length 1.25. Distance PME-PME 70 µm; diameter PME 50 µm; distance PME-ALE 25 µm; distance AME-AME 15 µm; diameter AME 30 µm. Diameters of leg femora 0.14–0.16, of leg tibiae 0.08–0.09. Clypeus unmodified, but slightly more protruding than in female; clypeus rim to ALE 0.21. Distances between cheliceral stridulatory ridges proximally 2.0 µm, distally 3.2 µm. Sternum humps very distinct, ~0.08 long. Procursus as in
<figureCitation id="13622A62AC6BFFCB0A696612E10FFD9E" box="[381,522,564,590]" captionStart="Fig" captionStartId="30.[189,232,1892,1918]" captionTargetBox="[191,1398,739,1848]" captionTargetId="figure-78@30.[189,1398,739,1848]" captionTargetPageId="30" captionText="Fig. 20. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Chelicerae, frontal and lateral views. C–E. Left palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. F–H. Left genital bulb, prolateral, dorsal, and retrolateral views. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8155534" httpUri="https://zenodo.org/record/8155534/files/figure.png" pageId="29" pageNumber="30">Fig. 20C–E</figureCitation>
, with distinctive transparent element distally (barely visible in dissecting microscope); genital bulb as in
<figureCitation id="13622A62AC6BFFCB0954667EE1CBFDA2" box="[576,718,600,626]" captionStart="Fig" captionStartId="30.[189,232,1892,1918]" captionTargetBox="[191,1398,739,1848]" captionTargetId="figure-78@30.[189,1398,739,1848]" captionTargetPageId="30" captionText="Fig. 20. Pholcophora texana Gertsch, 1935, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Chelicerae, frontal and lateral views. C–E. Left palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. F–H. Left genital bulb, prolateral, dorsal, and retrolateral views. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8155534" httpUri="https://zenodo.org/record/8155534/files/figure.png" pageId="29" pageNumber="30">Fig. 20F–H</figureCitation>
. Leg tibiae with slightly higher number of vertical hairs than in female (proximally on tibiae 1 and 2); prolateral trichobothrium absent on tibia 1. Tibia
<quantity id="4CA19B02AC6BFFCB0E53665AE672FD46" box="[1351,1399,636,662]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="30" unit="in" value="1.0">1 in</quantity>
previously measured specimens: 0.72 (
<bibRefCitation id="EFC84B16AC6BFFCB09906686E02DFD6A" author="Gertsch W. J." box="[644,808,672,698]" pageId="29" pageNumber="30" pagination="95 - 144" refId="ref33706" refString="Gertsch W. J. 1982. The spider genera Pholcophora and Anopsicus (Araneae, Pholcidae) in North America, Central America and the West Indies. Association of Mexican Cave Studies, Bulletin 8: 95 - 144 / Texas Memorial Museum, Bulletin 28: 95 - 144." type="journal article" year="1982">Gertsch 1982</bibRefCitation>
); 0.96, 0.97, 1.39 (
<bibRefCitation id="EFC84B16AC6BFFCB0F176686E793FD6A" author="Huber B. A." box="[1027,1174,672,698]" pageId="29" pageNumber="30" pagination="1 - 348" refId="ref34125" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / brh 26 h" type="journal article" year="2000">Huber 2000</bibRefCitation>
).
</paragraph>
<paragraph id="8BE636E7AC6BFFCB0BFE66C5E240FD2D" blockId="29.[187,1400,739,909]" box="[234,325,739,765]" pageId="29" pageNumber="30">
<emphasis id="B92DEAF5AC6BFFCB0BFE66C5E240FD2D" bold="true" box="[234,325,739,765]" pageId="29" pageNumber="30">Female</emphasis>
</paragraph>
<paragraph id="8BE636E7AC6BFFCB0BA96721E1A1FC5C" blockId="29.[187,1400,739,909]" pageId="29" pageNumber="30">
In general, similar to male but sternum without pair of anterior humps, and chelicerae without stridulatory files. Tibia
<quantity id="4CA19B02AC6BFFCB0A57670DE274FC94" box="[323,369,810,837]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="30" unit="in" value="1.0">1 in</quantity>
<specimenCount id="9D5FFD6EAC6BFFCB0A6E670DE2FCFC94" box="[378,505,810,837]" pageId="29" pageNumber="30" type="female">11 females</specimenCount>
from SW of Jacala: 0.90–1.18 (mean 1.03). Epigynum and internal female genitalia as in
<figureCitation id="13622A62AC6BFFCB0A666768E2CAFCB9" box="[370,463,846,873]" captionStart="Fig" captionStartId="31.[189,232,909,935]" captionTargetBox="[189,1398,265,869]" captionTargetId="figure-11@31.[189,1398,265,869]" captionTargetPageId="31" captionText="Fig. 21. Pholcophora texana Gertsch, 1935, female from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952), epigynum. A. Ventral view. B. Lateral view. Scale bar = 0.3 mm (both at same scale)." figureDoi="http://doi.org/10.5281/zenodo.8155536" httpUri="https://zenodo.org/record/8155536/files/figure.png" pageId="29" pageNumber="30">Figs 21</figureCitation>
,
<figureCitation id="13622A62AC6BFFCB0ACB6769E2FAFCB9" box="[479,511,847,873]" captionStart="Fig" captionStartId="31.[189,232,1891,1917]" captionTargetBox="[189,1398,1079,1846]" captionTargetId="figure-12@31.[189,1398,1079,1846]" captionTargetPageId="31" captionText="Fig. 22. Pholcophora texana Gertsch, 1935, females from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). A–B. Cleared female genitalia, ventral and dorsal views. C–D. Detail of median internal structures in two different specimens (slightly different angles of view). Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155538" httpUri="https://zenodo.org/record/8155538/files/figure.png" pageId="29" pageNumber="30">22</figureCitation>
,
<figureCitation id="13622A62AC6BFFCB091A6769E147FCB9" box="[526,578,847,873]" captionStart="Fig" captionStartId="32.[189,232,1754,1780]" captionTargetBox="[212,1376,265,1711]" captionTargetId="figure-12@32.[212,1376,265,1711]" captionTargetPageId="32" captionText="Fig. 23. Pholcophora texana Gertsch, 1935, female from Mexico, Hidalgo, 2.5 km SW of Jacala (ZFMK Mex341). A. Ocular area, dorsal view. B. Lateral face of chelicera, showing absence of stridulatory file. C. Epigynum, ventral (slightly posterior) view. D. ALS. E. PMS and anal hairs (arrows). F. Tip of left palpal tarsus, dorsal view. G. Detail of right palpal tarsus, dorsal view. H. Palpal tarsal organ. Abbreviations: ep = epigynum (main epigynal plate); pep = posterior epigynal plate; to = tarsal organ. Scale bars: A–B = 20 µm; C = 100 µm; D–G = 10 µm; H = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.8155540" httpUri="https://zenodo.org/record/8155540/files/figure.png" pageId="29" pageNumber="30">23C</figureCitation>
, with pair of receptacles (shape varies slightly with angle of view), without (or with very small?) pore plates.
</paragraph>
</subSubSection>
<caption id="DF26666FAC6BFFCB0BA963AEE2F7F839" ID-DOI="http://doi.org/10.5281/zenodo.8155532" ID-Zenodo-Dep="8155532" httpUri="https://zenodo.org/record/8155532/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1928,1954]" targetBox="[189,1398,1145,1879]" targetPageId="29">
<paragraph id="8BE636E7AC6BFFCB0BA963AEE2F7F839" blockId="29.[189,1398,1928,2025]" pageId="29" pageNumber="30">
<emphasis id="B92DEAF5AC6BFFCB0BA963AEE224F872" bold="true" box="[189,289,1928,1954]" pageId="29" pageNumber="30">Fig. 19.</emphasis>
<taxonomicName id="4C594D64AC6BFFCB0A3B63AEE1E4F873" authority="Gertsch, 1935" authorityName="Gertsch" authorityYear="1935" box="[303,737,1928,1955]" class="Arachnida" family="Pholcidae" genus="Pholcophora" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="B92DEAF5AC6BFFCB0A3B63AEE120F872" box="[303,549,1928,1954]" italics="true" pageId="29" pageNumber="30">Pholcophora texana</emphasis>
Gertsch, 1935
</taxonomicName>
, male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). Left palp, prolateral, dorsal, and retrolateral views; arrows point at unique brush of modified hairs. Scale bar = 0.3 mm.
</paragraph>
</caption>
<subSubSection id="C343656CAC68FFC80BA9652EE610FE4A" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BE636E7AC68FFC80BA9652EE253FEF2" blockId="30.[189,342,264,290]" box="[189,342,264,290]" pageId="30" pageNumber="31">
<heading id="D0AE818BAC68FFC80BA9652EE253FEF2" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B92DEAF5AC68FFC80BA9652EE253FEF2" bold="true" box="[189,342,264,290]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636E7AC68FFC80BA9651EE610FE4A" blockId="30.[189,1398,312,410]" pageId="30" pageNumber="31">
Apparently widely distributed in north-eastern
<collectingCountry id="F34E7677AC68FFC809F2651EE046FE82" box="[742,835,312,338]" name="Mexico" pageId="30" pageNumber="31">Mexico</collectingCountry>
, ranging into southern Texas (
<figureCitation id="13622A62AC68FFC80FB2651EE7EBFE82" box="[1190,1262,312,338]" captionStart="Fig" captionStartId="11.[189,232,1822,1848]" captionTargetBox="[189,1398,730,1773]" captionTargetId="figure-150@11.[189,1398,730,1773]" captionTargetPageId="11" captionText="Fig. 2. Known distributions of ‘true’ Pholcophora Banks, 1896 (orange marks) and of presumably misplaced Caribbean ‘Pholcophora’ and of Papiamenta Huber, 2000 (red marks). Dominican amber species currently placed in Pholcophora are represented by an “F” in a pink square. Our molecular data suggest that Caribbean ‘Pholcophora’ are more closely related with the genus Papiamenta than with true Pholcophora. Barcoded specimens of Pholcophora americana Banks, 1896 are marked with an “x” and accompanied by the vial code (see Table 1 for details)." figureDoi="http://doi.org/10.5281/zenodo.8155478" httpUri="https://zenodo.org/record/8155478/files/figure.png" pageId="30" pageNumber="31">Fig. 2</figureCitation>
). However, the map in
<figureCitation id="13622A62AC68FFC80A53657AE28BFEA6" box="[327,398,348,374]" captionStart="Fig" captionStartId="11.[189,232,1822,1848]" captionTargetBox="[189,1398,730,1773]" captionTargetId="figure-150@11.[189,1398,730,1773]" captionTargetPageId="11" captionText="Fig. 2. Known distributions of ‘true’ Pholcophora Banks, 1896 (orange marks) and of presumably misplaced Caribbean ‘Pholcophora’ and of Papiamenta Huber, 2000 (red marks). Dominican amber species currently placed in Pholcophora are represented by an “F” in a pink square. Our molecular data suggest that Caribbean ‘Pholcophora’ are more closely related with the genus Papiamenta than with true Pholcophora. Barcoded specimens of Pholcophora americana Banks, 1896 are marked with an “x” and accompanied by the vial code (see Table 1 for details)." figureDoi="http://doi.org/10.5281/zenodo.8155478" httpUri="https://zenodo.org/record/8155478/files/figure.png" pageId="30" pageNumber="31">Fig. 2</figureCitation>
includes four records that are based exclusively on females, i.e., the identifications should be verified (e.g., by collecting males or by sequencing specimens from these localities).
</paragraph>
</subSubSection>
<subSubSection id="C343656CAC68FFC80BA965E4E23AFDE0" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="8BE636E7AC68FFC80BA965E4E27BFE0C" blockId="30.[189,382,450,476]" box="[189,382,450,476]" pageId="30" pageNumber="31">
<heading id="D0AE818BAC68FFC80BA965E4E27BFE0C" bold="true" box="[189,382,450,476]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B92DEAF5AC68FFC80BA965E4E27BFE0C" bold="true" box="[189,382,450,476]" pageId="30" pageNumber="31">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636E7AC68FFC80BA965D7E23AFDE0" blockId="30.[189,1399,497,560]" pageId="30" pageNumber="31">
The newly collected specimens were found under rocks on an arid hillside set with low thorn scrub (
<figureCitation id="13622A62AC68FFC80BD26633E234FDE0" box="[198,305,533,560]" captionStart="Fig" captionStartId="44.[189,232,1892,1918]" captionTargetBox="[189,1398,1014,1845]" captionTargetId="figure-172@44.[189,1398,1014,1845]" captionTargetPageId="44" captionText="Fig. 34. Representative sample of habitats of Pholcophora Banks, 1896 in Mexico. A. Guerrero, 2 km N of Mazatlán (type locality of P. mazatlan Huber sp. nov.). B. Hidalgo, 2.5 km SW of Jacala (P. texana Gertsch, 1935). C. Guerrero, 5 km S of Papanoa (type locality of P. papanoa Huber sp. nov.; with collecting tray). D. Puebla, 35 km SE of Tehuacan (type locality of P. tehuacan Huber sp. nov.)." figureDoi="http://doi.org/10.5281/zenodo.8155578" httpUri="https://zenodo.org/record/8155578/files/figure.png" pageId="30" pageNumber="31">Fig. 34B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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