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<document id="E0A5F56251C33772467F6A35E56ECCDD" ID-DOI="10.11646/zootaxa.3746.2.3" ID-GBIF-Dataset="112d8c10-eade-4f43-9f8d-3f8042ec2fd2" ID-ISSN="1175-5326" ID-Zenodo-Dep="248936" ID-ZooBank="86A7ED83-1A8E-4227-9BD6-5949E57C392F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460707396962" checkinUser="plazi" docAuthor="Malipatil, M. B. &amp; Blacket, M. J." docDate="2013" docId="03F0879DE66D8E0D798CA0CECF6AF81B" docLanguage="en" docName="zt03746p300.pdf" docOrigin="Zootaxa 3746 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Germalus littoralis Malipatil, sp. nov." docType="treatment" docVersion="8" lastPageNumber="285" masterDocId="FFC9FFE5E6768E11791BA059CE5FFFFA" masterDocTitle="Review and revision of Australian Germalus Stål, with new genera and further new species of Australian Geocorinae (Hemiptera: Heteroptera: Geocoridae)" masterLastPageNumber="300" masterPageNumber="257" pageNumber="284" updateTime="1698357443482" updateUser="plazi">
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<mods:title id="F2548430756C85A8DD55D3E183C5E61E">Review and revision of Australian Germalus Stål, with new genera and further new species of Australian Geocorinae (Hemiptera: Heteroptera: Geocoridae)</mods:title>
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<mods:namePart id="9AB5C1D695BCD02A13731D5EFC031C1C">Malipatil, M. B.</mods:namePart>
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<mods:namePart id="0F748F4E4B9580B621F4F132A43C7728">Blacket, M. J.</mods:namePart>
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<mods:date id="D8458CA4765EA414A70074000C37102A">2013</mods:date>
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<treatment id="03F0879DE66D8E0D798CA0CECF6AF81B" ID-DOI="http://doi.org/10.5281/zenodo.6147031" ID-GBIF-Taxon="119584219" ID-Zenodo-Dep="6147031" LSID="urn:lsid:plazi:treatment:03F0879DE66D8E0D798CA0CECF6AF81B" httpUri="http://treatment.plazi.org/id/03F0879DE66D8E0D798CA0CECF6AF81B" lastPageId="28" lastPageNumber="285" pageId="27" pageNumber="284">
<subSubSection id="C3436500E66D8E0A798CA0CECF64FF29" pageId="27" pageNumber="284" type="nomenclature">
<paragraph id="8BE6368BE66D8E0A798CA0CECC3EFF48" blockId="27.[151,609,151,211]" box="[151,609,151,178]" pageId="27" pageNumber="284">
<heading id="D0AE81E7E66D8E0A798CA0CECC3EFF48" bold="true" box="[151,609,151,178]" fontSize="11" level="1" pageId="27" pageNumber="284" reason="1">
<emphasis id="B92DEA99E66D8E0A798CA0CECC3EFF48" bold="true" box="[151,609,151,178]" pageId="27" pageNumber="284">
<taxonomicName id="4C594D08E66D8E0A798CA0CECFA9FF48" ID-CoL="8QP9R" authority="Malipatil" authorityName="Malipatil" box="[151,502,151,178]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="284" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="B92DEA99E66D8E0A798CA0CECF25FF4B" bold="true" box="[151,378,151,177]" italics="true" pageId="27" pageNumber="284">Germalus littoralis</emphasis>
Malipatil
</taxonomicName>
,
<taxonomicNameLabel id="A21E57E2E66D8E0A7B1FA0C1CC3EFF48" box="[516,609,152,178]" pageId="27" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE6368BE66D8E0A798CA0E3CF64FF29" blockId="27.[151,609,151,211]" box="[151,315,186,211]" pageId="27" pageNumber="284">
(
<figureCitation id="13622A0EE66D8E0A7984A0E3CF6CFF29" box="[159,307,186,211]" captionStart-0="FIGURES 98 100" captionStart-1="FIGURES 101 105" captionStart-2="FIGURES 106 115" captionStartId-0="27.[151,269,1605,1629]" captionStartId-1="29.[151,269,1975,1999]" captionStartId-2="30.[151,269,1468,1492]" captionTargetBox-0="[152,1435,554,1578]" captionTargetBox-1="[207,1380,196,1951]" captionTargetBox-2="[152,1436,195,1446]" captionTargetId-0="figure@27.[151,1436,554,1580]" captionTargetId-1="figure@29.[205,1381,195,1951]" captionTargetId-2="figure@30.[151,1436,194,1447]" captionTargetPageId-0="27" captionTargetPageId-1="29" captionTargetPageId-2="30" captionText-0="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." captionText-1="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." captionText-2="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri-0="https://zenodo.org/record/248954/files/figure.png" httpUri-1="https://zenodo.org/record/248955/files/figure.png" httpUri-2="https://zenodo.org/record/248956/files/figure.png" pageId="27" pageNumber="284">Figs. 98115</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3436500E66D8E0A798CA15BCBDCFE99" pageId="27" pageNumber="284" type="materials_examined">
<paragraph id="8BE6368BE66D8E0A798CA15BCBDCFE99" blockId="27.[151,1436,258,499]" pageId="27" pageNumber="284">
<emphasis id="B92DEA99E66D8E0A798CA15BCF03FEE1" bold="true" box="[151,348,258,283]" pageId="27" pageNumber="284">
<typeStatus id="54E28829E66D8E0A798CA15BCE8DFEE1" box="[151,210,258,283]" pageId="27" pageNumber="284">Type</typeStatus>
specimens:
</emphasis>
<typeStatus id="54E28829E66D8E0A787FA15BCF93FEE1" box="[356,460,258,283]" pageId="27" pageNumber="284" type="holotype">Holotype</typeStatus>
female, Queensland, 7941 Yule Point, Cairns-Mossman Road,
<date id="FFE7104BE66D8E0A7D81A15BCB52FEE1" box="[1178,1293,258,283]" pageId="27" pageNumber="284" value="1966-10-27">27.x.1966</date>
, EB Britton, near mangroves, in ANIC.
<typeStatus id="54E28829E66D8E0A78C1A17ECC18FEC5" box="[474,583,295,319]" pageId="27" pageNumber="284" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D5FFD02E66D8E0A7B48A17ECCEFFEC5" box="[595,688,294,319]" pageId="27" pageNumber="284" type="male">3 males</specimenCount>
,
<specimenCount id="9D5FFD02E66D8E0A7BDAA17ECD75FEC5" box="[705,810,294,319]" pageId="27" pageNumber="284" type="female">1 female</specimenCount>
, same data as
<typeStatus id="54E28829E66D8E0A7AC6A17FCA1DFEC5" box="[989,1090,294,319]" pageId="27" pageNumber="284" type="holotype">holotype</typeStatus>
, in ANIC;
<specimenCount id="9D5FFD02E66D8E0A7DD0A17ECB41FEC5" box="[1227,1310,294,319]" pageId="27" pageNumber="284" type="male">1 male</specimenCount>
,
<specimenCount id="9D5FFD02E66D8E0A7C34A17ECBC7FEC5" box="[1327,1432,294,319]" pageId="27" pageNumber="284" type="female">1 female</specimenCount>
, Mackay, mangroves,
<date id="FFE7104BE66D8E0A789DA113CFADFE99" box="[390,498,330,355]" pageId="27" pageNumber="284" value="1967-01-19">19.i.1967</date>
, D McAlpine &amp; G Holloway coll, in AM;
<specimenCount id="9D5FFD02E66D8E0A7ACBA112CA42FE99" box="[976,1053,330,355]" pageId="27" pageNumber="284" type="male">1 male</specimenCount>
, Cairns dist., AM Lea, in SAM.
</paragraph>
</subSubSection>
<subSubSection id="C3436500E66D8E0D79DCA137CC8FF964" lastPageId="28" lastPageNumber="285" pageId="27" pageNumber="284" type="description">
<paragraph id="8BE6368BE66D8E0A79DCA137CC18FE09" blockId="27.[151,1436,258,499]" pageId="27" pageNumber="284">
<emphasis id="B92DEA99E66D8E0A79DCA137CF07FE7D" bold="true" box="[199,344,366,391]" pageId="27" pageNumber="284">Description.</emphasis>
<emphasis id="B92DEA99E66D8E0A787DA137CFB0FE7D" box="[358,495,366,391]" italics="true" pageId="27" pageNumber="284">Colouration</emphasis>
: Body pale stramineous (
<figureCitation id="13622A0EE66D8E0A7A3CA137CD20FE7D" box="[807,895,366,391]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="27" pageNumber="284">Fig. 98</figureCitation>
) with punctures on pronotum, scutellum and hemelytra slightly fuscous; abdominal dorsum, below hemelytra and membrane, with median narrow fuscous patch for entire length; apical segments of labium brown; tip of head with median line on clypeus, and inner lines demarcating jugae, fuscous (
<figureCitation id="13622A0EE66D8E0A78C3A183CC65FE09" box="[472,570,474,499]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="27" pageNumber="284">Fig. 100</figureCitation>
).
</paragraph>
<caption id="DF266603E66D8E0A798CA61CCF37F986" httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="27" pageNumber="284" targetBox="[152,1435,554,1578]" targetPageId="27">
<paragraph id="8BE6368BE66D8E0A798CA61CCF37F986" blockId="27.[151,1436,1604,1660]" pageId="27" pageNumber="284">
<emphasis id="B92DEA99E66D8E0A798CA61CCF32F9A7" bold="true" box="[151,365,1605,1629]" pageId="27" pageNumber="284">FIGURES 98100.</emphasis>
<taxonomicName id="4C594D08E66D8E0A786EA61DCC1CF9A6" box="[373,579,1604,1628]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="284" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="B92DEA99E66D8E0A786EA61DCC1CF9A6" box="[373,579,1604,1628]" italics="true" pageId="27" pageNumber="284">Germalus littoralis</emphasis>
</taxonomicName>
, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged.
</paragraph>
</caption>
<paragraph id="8BE6368BE66D8E0A79DCA6FECC5AF918" blockId="27.[151,1436,1701,2015]" pageId="27" pageNumber="284">
<emphasis id="B92DEA99E66D8E0A79DCA6FECF36F944" box="[199,361,1703,1726]" italics="true" pageId="27" pageNumber="284">Measurements</emphasis>
: are of
<typeStatus id="54E28829E66D8E0A78DDA6FCCC77F944" box="[454,552,1701,1726]" pageId="27" pageNumber="284" type="holotype">holotype</typeStatus>
female,
<typeStatus id="54E28829E66D8E0A7B95A6FECCB1F945" box="[654,750,1703,1727]" pageId="27" pageNumber="284" type="paratype">paratype</typeStatus>
male, followed by ranges of remaining
<typeStatus id="54E28829E66D8E0A7DA5A6FECB41F945" box="[1214,1310,1703,1727]" pageId="27" pageNumber="284" type="paratype">paratype</typeStatus>
specimens measured within round brackets.
</paragraph>
<paragraph id="8BE6368BE66D8E0A79DCA6B4CF7EF8D0" blockId="27.[151,1436,1701,2015]" pageId="27" pageNumber="284">
<emphasis id="B92DEA99E66D8E0A79DCA6B4CF5EF8FC" box="[199,257,1773,1798]" italics="true" pageId="27" pageNumber="284">Body</emphasis>
: Narrow, elongate, parallel-sided (
<figureCitation id="13622A0EE66D8E0A7B89A6B4CCB6F8FC" box="[658,745,1773,1798]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="27" pageNumber="284">Fig. 98</figureCitation>
); length 5.42, 4.83 (4.835.30); maximum width 1.65, 1.44 (1.401.63).
</paragraph>
<paragraph id="8BE6368BE66D8E0D79DCA76CCFC7FE48" blockId="27.[151,1436,1701,2015]" lastBlockId="28.[151,1436,151,2018]" lastPageId="28" lastPageNumber="285" pageId="27" pageNumber="284">
<emphasis id="B92DEA99E66D8E0A79DCA76CCF5BF8B4" box="[199,260,1845,1870]" italics="true" pageId="27" pageNumber="284">Head</emphasis>
: Surface impunctate; anterior of eyes almost parallel-sided (
<figureCitation id="13622A0EE66D8E0A7AA9A76CCA49F8B4" box="[946,1046,1845,1870]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="27" pageNumber="284">Fig. 100</figureCitation>
); head width across eyes subequal to body width; eyes stylate with antennifers slightly projecting outward (
<figureCitation id="13622A0EE66D8E0A7AD6A700CA6FF888" box="[973,1072,1881,1906]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="27" pageNumber="284">Fig. 100</figureCitation>
); bucculae abruptly diminishing behind flaps portion almost to surface of head just posterior of antennifers, continuing as parallel lines before joining in V-form, thereafter continuing as a fine line to base of head; gular area broadly strongly flattened, demarcated, covered with long dense fine setae. Length of head 0.85, 80 (0.780.82); width across eyes 1.42, 1.38 (1.341.45); interocular space 0.80, 0.75 (0.730.82); interocellar space 0.32, 0.27 (0.320.34); eye-ocellar space 0.24, 0.23 (0.200.24); eye length 0.46, 0.46 (0.440.46); eye width 0.27, 0.25 (0.240.27). Labium robust, extending to mid coxae, 1st segment almost reaching base of head; length of segments: I 0.57, 0.48 (0.500.57); II 0.48, 0.46 (0.400.48); III 0.43, 0.45 (0.430.46); IV 0.43, 0.40 (0.400.44). Labrum extending to about ½ length of 2nd labial segment. Antennae with 1st segment exceeding clypeus by about 1/3, generally short and stout; 4th segment incrassate, thicker than other segments; basal segment with bristles, remainder with pubescence only. Length of antennal segments: I 0.34, 0.35 (0.340.40); II 0.59, 0.64 (0.590.62); III 0.30, 0.30 (0.300.32); IV 0.57, 0.52 (0.520.60).
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA1E4CA94FC34" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">
<emphasis id="B92DEA99E66A8E0D79DCA1E4CF47FE2C" box="[199,280,445,470]" italics="true" pageId="28" pageNumber="285">Thorax</emphasis>
: Pronotum quadrate (
<figureCitation id="13622A0EE66A8E0D7B11A1E4CC03FE2C" box="[522,604,445,470]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="28" pageNumber="285">Fig. 99</figureCitation>
), narrowly carinate on lateral margins, slightly concave about middle, calli impunctate, posterior margin straight, sparsely but distinctly punctuate all over except on calli and anterior and posterior margins. Median length 0.98, 0.92 (0.991.03); width at posterior margin 1.38, 1.26 (1.291.40). Thoracic pleura densely punctate with dark punctures. Scent gland opening narrow and directed posteriorly, evaporative area narrow. Legs particularly femora shiny, fore femora more incrassate than other femora; all legs covered with minute black spicules irregularly distributed on tibiae and femora, particularly on posterior (outer) and dorsal (upper) surfaces and on apical ½ of mid and hind femora (thus surfaces rubbing against body devoid of spicules). [Note these spicules are present in many other species examined in the present study, but the spicules appear to be more conspicuous by their black colour on a pale background and their density, particularly in this species and in
<taxonomicName id="4C594D08E66A8E0D7853A35BCC16FCE3" box="[328,585,770,793]" class="Insecta" family="Geocoridae" genus="Unicageocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="285" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="B92DEA99E66A8E0D7853A35BCC16FCE3" box="[328,585,770,793]" italics="true" pageId="28" pageNumber="285">Unicageocoris griseus</emphasis>
</taxonomicName>
]. Scutellum length 0.65, 0.62 (0.600.67); width 0.69, 0.66 (0.660.71). Hemelytra parallel-sided (
<figureCitation id="13622A0EE66A8E0D78DAA37CCC47FCC4" box="[449,536,805,830]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="28" pageNumber="285">Fig. 98</figureCitation>
); corium with almost three complete rows of punctures: inner row continuing brokenly onto posterior margin, one submarginal row almost complete to tip, and one middle row; clavus parallelsided with faint punctures in two rows: one incomplete row along inner margin continuing onto commissural area, one almost complete row along outer margin becoming indistinct in apical third. Length of hemelytra 3.86, 3.35 (3.353.68); length of corium 2.53, 2.16 (2.252.50); claval commissure 0.46, 0.41 (0.430.46).
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA380CA20FB14" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">
<emphasis id="B92DEA99E66A8E0D79DCA380CF6EFC08" box="[199,305,985,1010]" italics="true" pageId="28" pageNumber="285">Abdomen</emphasis>
: Narrow and parallel-sided. Inner laterotergites narrow, almost absent. Dorsal scent gland scars between terga IVV and VVI subequal in width; sutures moderately curved posteriorly from margin to meson (
<figureCitation id="13622A0EE66A8E0D7984A478CF50FBC0" box="[159,271,1057,1082]" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Figs. 101</figureCitation>
,
<figureCitation id="13622A0EE66A8E0D783BA478CF11FBC0" box="[288,334,1057,1082]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">106</figureCitation>
). Trichobothria on tergum III reduced, in sublinear to triangular series; those on IV reduced, arranged in linear series. Female with tergum almost fully membranous, not sclerotised except for narrow median light fuscous band, as in
<figureCitation id="13622A0EE66A8E0D78A4A430CC47FB78" box="[447,536,1129,1154]" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Fig.101</figureCitation>
; abdominal sternum not sclerotised. uniformly covered with short decumbent setae (
<figureCitation id="13622A0EE66A8E0D79F9A4D4CF17FB5C" box="[226,328,1165,1190]" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Fig. 102</figureCitation>
). Male apodemes on anterior margin of sternum V with a pair of broad flaps on either side of midline (
<figureCitation id="13622A0EE66A8E0D79E6A4E8CF3FFB30" box="[253,352,1201,1226]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Fig. 107</figureCitation>
); those on anterior margin of VII with one large flap in middle extending to about anterior ¼ of sternum VI; sternum III with a median pair of minute flaps on anterior margin (
<figureCitation id="13622A0EE66A8E0D7D0BA48CCA2CFB14" box="[1040,1139,1237,1262]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Fig. 107</figureCitation>
).
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA4A0CA47FA84" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">
<emphasis id="B92DEA99E66A8E0D79DCA4A0CFD5FAE8" box="[199,394,1273,1298]" italics="true" pageId="28" pageNumber="285">Female genitalia</emphasis>
: Spermathecal bulb (
<figureCitation id="13622A0EE66A8E0D7B9DA4A0CCB5FAE8" box="[646,746,1273,1298]" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Fig. 103</figureCitation>
) almost spherical, with a flange near base, sclerotised and almost evenly expanded; duct part behind flange short and tubular, with no twists or coils; basal part with duct short and narrow. Ovipositor thin and elongate, 1st ramus traversing to almost 5/6 length of 1st gonapophysis (
<figureCitation id="13622A0EE66A8E0D7C75A518CE9AFA84" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Fig. 104</figureCitation>
); 2nd gonapophysis narrow, spatulate; 2nd gonocoxae crescentic (
<figureCitation id="13622A0EE66A8E0D7AB2A53CCA54FA84" box="[937,1035,1381,1406]" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Fig. 105</figureCitation>
).
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA5D0CC8FF964" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">
<emphasis id="B92DEA99E66A8E0D79DCA5D0CF2FFA58" box="[199,368,1417,1442]" italics="true" pageId="28" pageNumber="285">Male genitalia</emphasis>
: Pygophore as in
<figureCitation id="13622A0EE66A8E0D7B51A5D0CCF1FA58" box="[586,686,1417,1442]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Fig. 108</figureCitation>
, with brushy long setae. Paramere (
<figureCitation id="13622A0EE66A8E0D7D4DA5D0CAA4FA58" box="[1110,1275,1417,1442]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Figs. 113115</figureCitation>
) with acutely pointed sickle-shaped blade, dorsal lobe pointed, ventral lobe indistinct; blade above dorsal lobes with a tuft of long setae. Aedeagus as in (
<figureCitation id="13622A0EE66A8E0D78D5A588CC2EFA10" box="[462,625,1489,1514]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Figs. 109111</figureCitation>
). Phallotheca moderately sclerotised in a band form near base on ventral aspect; dorsally membranous except for two ill defined heavily sclerotised plates: one small near base, the other large elongate towards apical area near base of vesica. Ejaculatory reservoir (
<figureCitation id="13622A0EE66A8E0D7AE1A640CA06F9C8" box="[1018,1113,1561,1586]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Fig. 112</figureCitation>
) with neck, body, and wings well developed, body and neck not produced posteriorly. Vesica membranous; helicoid process moderately sclerotised, with 1½ coils, then gradually narrowed. Gonoporal process robust, with 23 coils, ending in a blunt unusually enlarged and flared gonopore (
<figureCitation id="13622A0EE66A8E0D7B79A6DCCC9BF964" box="[610,708,1669,1694]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Fig. 112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3436500E66A8E0D79DCA6F0CF6AF81B" pageId="28" pageNumber="285" type="distribution">
<paragraph id="8BE6368BE66A8E0D79DCA6F0CA93F938" blockId="28.[151,1436,151,2018]" box="[199,1228,1705,1730]" pageId="28" pageNumber="285">
<emphasis id="B92DEA99E66A8E0D79DCA6F0CF3FF938" bold="true" box="[199,352,1705,1730]" pageId="28" pageNumber="285">Distribution.</emphasis>
Only northern Queensland, commonly collected in the vicinity of mangroves.
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA694CF5BF8F0" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">
<emphasis id="B92DEA99E66A8E0D79DCA694CF4EF91C" bold="true" box="[199,273,1741,1766]" pageId="28" pageNumber="285">Notes.</emphasis>
The species name alludes to the “shore (littus or littoris) habitat from which the
<typeStatus id="54E28829E66A8E0D7DF7A697CB43F91C" box="[1260,1308,1742,1766]" pageId="28" pageNumber="285">type</typeStatus>
series was collected.
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA74CCD2AF860" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">
This species can be readily distinguished from other species examined in the present study by the narrow, elongate, parallel-sided body (
<figureCitation id="13622A0EE66A8E0D78E8A760CC16F8A8" box="[499,585,1849,1874]" captionStart="FIGURES 98 100" captionStartId="27.[151,269,1605,1629]" captionTargetBox="[152,1435,554,1578]" captionTargetId="figure@27.[151,1436,554,1580]" captionTargetPageId="27" captionText="FIGURES 98 100. Germalus littoralis, female, dorsal view: 98, habitus; 99, head, pronotum and scutellum enlarged; 100, head enlarged." httpUri="https://zenodo.org/record/248954/files/figure.png" pageId="28" pageNumber="285">Fig. 98</figureCitation>
), dorsally impunctate head, broad flattened and pilose gular area, and the quadrate pronotum. Further, the near spherical spermathecal bulb with its short non-coiled duct (
<figureCitation id="13622A0EE66A8E0D7DC8A704CB6AF88C" box="[1235,1333,1885,1910]" captionStart="FIGURES 101 105" captionStartId="29.[151,269,1975,1999]" captionTargetBox="[207,1380,196,1951]" captionTargetId="figure@29.[205,1381,195,1951]" captionTargetPageId="29" captionText="FIGURES 101 105. Germalus littoralis, female: 101, abdominal dorsum; 102, abdominal venter; 103, spermatheca; 104, 105, ovipositor." httpUri="https://zenodo.org/record/248955/files/figure.png" pageId="28" pageNumber="285">Fig. 103</figureCitation>
), and the flared gonoporal process (
<figureCitation id="13622A0EE66A8E0D78A0A7D8CC44F860" box="[443,539,1921,1946]" captionStart="FIGURES 106 115" captionStartId="30.[151,269,1468,1492]" captionTargetBox="[152,1436,195,1446]" captionTargetId="figure@30.[151,1436,194,1447]" captionTargetPageId="30" captionText="FIGURES 106 115. Germalus littoralis, male: 106, abdominal dorsum; 107, abdominal venter; 108, pygophore, dorsal view; 109 111, aedeagus, ventral, lateral and dorsal views; 112, ejaculatory reservoir and helicoid process enlarged; 113 115 paramere, different views." httpUri="https://zenodo.org/record/248956/files/figure.png" pageId="28" pageNumber="285">Fig. 112</figureCitation>
), help to diagnose
<emphasis id="B92DEA99E66A8E0D7BF5A7DBCD2AF860" box="[750,885,1921,1946]" italics="true" pageId="28" pageNumber="285">
<taxonomicName id="4C594D08E66A8E0D7BF5A7DBCD2EF860" box="[750,881,1921,1946]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="285" phylum="Arthropoda" rank="species" species="littoralis">G. littoralis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BE6368BE66A8E0D79DCA7FCCF6AF81B" blockId="28.[151,1436,151,2018]" pageId="28" pageNumber="285">The Mackay specimens have fuscous brown irregular areas or spots on the antennae and the humeral angles of the pronotum.</paragraph>
</subSubSection>
</treatment>
</document>