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<document id="D57C77A19B8C7A53E5C5E5BC70D41E51" ID-DOI="10.5281/zenodo.198060" ID-GBIF-Dataset="ed312cce-318a-473e-9e60-a9b097373666" ID-ISSN="1175-5326" ID-Zenodo-Dep="198060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460283733665" checkinUser="plazi" docAuthor="Zhao, Zony-Yi &amp; Zhou, Hong-Zhang" docDate="2010" docId="03EF87FABD18FFE59DD3DD82FBDF71F3" docLanguage="en" docName="zt02619p038.pdf" docOrigin="Zootaxa 2619" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Indoquedius Blackwelder 1952" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFD6FF82BD19FFE49D44DF6FFFF3742C" masterDocTitle="Taxonomy of the genus Indoquedius Blackwelder (Coleoptera: Staphylinidae: Staphylininae) of China with description of four new species" masterLastPageNumber="38" masterPageNumber="27" pageNumber="28" updateTime="1698249979089" updateUser="plazi">
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<mods:title id="253A617F5C1BB80F1C8D834BA4946323">Taxonomy of the genus Indoquedius Blackwelder (Coleoptera: Staphylinidae: Staphylininae) of China with description of four new species</mods:title>
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<mods:namePart id="B2799D11B766104A927BEF3BC7B52E4B">Zhao, Zony-Yi</mods:namePart>
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<mods:namePart id="866DA0F4E718FC9B62704BB13945C5D4">Zhou, Hong-Zhang</mods:namePart>
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<subSubSection id="C35C6567BD18FFE59DD3DD82FBDF71F3" pageId="1" pageNumber="28" type="nomenclature">
<paragraph id="8BF936ECBD18FFE59DD3DD82FD83772B" blockId="1.[151,624,749,775]" box="[151,624,749,775]" pageId="1" pageNumber="28">
<heading id="D0B18180BD18FFE59DD3DD82FD83772B" bold="true" box="[151,624,749,775]" fontSize="11" level="1" pageId="1" pageNumber="28" reason="1">
<emphasis id="B932EAFEBD18FFE59DD3DD82FD83772B" bold="true" box="[151,624,749,775]" pageId="1" pageNumber="28">
Genus
<taxonomicName id="4C464D6FBD18FFE59DAADD82FD83772B" ID-CoL="8VYHC" authority="Blackwelder, 1952" authorityName="Blackwelder" authorityYear="1952" box="[238,624,749,775]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B932EAFEBD18FFE59DAADD82FE75772B" bold="true" box="[238,390,749,775]" italics="true" pageId="1" pageNumber="28">Indoquedius</emphasis>
<bibRefCitation id="EFD74B1DBD18FFE59CC8DD82FD83772B" author="Blackwelder" box="[396,624,749,775]" pageId="1" pageNumber="28" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, i - iv, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936ECBD18FFE59DD3DC5AFB8A77A0" blockId="1.[151,1436,821,908]" pageId="1" pageNumber="28">
<bibRefCitation id="EFD74B1DBD18FFE59DD3DC5AFEB27760" author="Cameron" box="[151,321,821,844]" pageId="1" pageNumber="28" refString="Cameron, M. (1932) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. III. Taylor and Francis, London, xiii + 1 - 443 pp." type="book chapter" year="1932">
<collectingCountry id="F351767CBD18FFE59DD3DC5AFF0F7760" box="[151,252,821,844]" name="Cameroon" pageId="1" pageNumber="28">Cameron</collectingCountry>
, 1932
</bibRefCitation>
: 281, 300 (subgenus of
<taxonomicName id="4C464D6FBD18FFE59F0EDC5AFD507760" box="[586,675,821,844]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B932EAFEBD18FFE59F0EDC5AFD507760" box="[586,675,821,844]" italics="true" pageId="1" pageNumber="28">Quedius</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFD74B1DBD18FFE59FF8DC5AFC797760" author="Blackwelder" box="[700,906,821,844]" pageId="1" pageNumber="28" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, i - iv, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
: 199 (designation of the
<typeStatus id="54FD884EBD18FFE599D8DC5AFB397760" box="[1180,1226,821,844]" pageId="1" pageNumber="28">type</typeStatus>
species);
<bibRefCitation id="EFD74B1DBD18FFE5987DDC5AFEF37740" author="Smetana" pageId="1" pageNumber="28" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24 (2), 163 - 464." type="journal article" year="1988">Smetana, 1988</bibRefCitation>
: 300;
<bibRefCitation id="EFD74B1DBD18FFE59C04DC3AFE117740" author="Smetana" box="[320,482,853,876]" pageId="1" pageNumber="28" refString="Smetana, A. (1995) Revision of the tribes Quediini and Tanygnathinini. Part III. Taiwan. (Coleoptera: Staphylinidae). National Museum of Natural Science, Special Publication Number 6, 1 - 145." type="book chapter" year="1995">Smetana, 1995</bibRefCitation>
: 112;
<bibRefCitation id="EFD74B1DBD18FFE59F64DC3AFD317740" author="Smetana" box="[544,706,853,876]" pageId="1" pageNumber="28" refString="Smetana, A. (2004) Staphylinidae, subfamilies Omaliinae - Dasycerinae, Phloeocharinae - Apatecinae, Piestinae - Staphylininae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. II. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 237 - 272, 329 - 495, 505 - 698." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 656;
<bibRefCitation id="EFD74B1DBD18FFE59E46DC3AFC597740" author="Herman" box="[770,938,853,876]" pageId="1" pageNumber="28" refString="Herman, L. H. (2001 a) Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History, 264, 1 - 83." type="journal article" year="2001" yearSuffix="a">Herman, 2001a</bibRefCitation>
: 9;
<bibRefCitation id="EFD74B1DBD18FFE59E94DC3AFB8A7740" author="Herman" box="[976,1145,853,876]" pageId="1" pageNumber="28" refString="Herman, L. H. (2001 b) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Parts I - VII. Bulletin of the American Museum of Natural History, 265, 1 - 4218." type="journal article" year="2001" yearSuffix="b">Herman, 2001b</bibRefCitation>
: 3079;
<bibRefCitation id="EFD74B1DBD18FFE59982DC3AFA907740" author="Hayashi" box="[1222,1379,853,876]" pageId="1" pageNumber="28" refString="Hayashi, Y. (2009) Notes on the genus Indoquedius, with description of a new species from Malaysia. The Japanese Society of Systematic Entomology, 15 (1), 147 - 151." type="journal article" year="2009">Hayashi, 2009</bibRefCitation>
: 147.
<emphasis id="B932EAFEBD18FFE59DD3DC1BFED077A0" bold="true" box="[151,291,884,908]" pageId="1" pageNumber="28">
<typeStatus id="54FD884EBD18FFE59DD3DC1BFF3C77A0" box="[151,207,884,908]" pageId="1" pageNumber="28">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C464D6FBD18FFE59C76DC1AFDCE77A0" authority="Fauvel" authorityName="Fauvel" box="[306,573,885,908]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B932EAFEBD18FFE59C76DC1AFE1877A0" box="[306,491,885,908]" italics="true" pageId="1" pageNumber="28">Quedius oculatus</emphasis>
Fauvel
</taxonomicName>
, fixed by original designation in
<bibRefCitation id="EFD74B1DBD18FFE59ED9DC1AFB8677A0" author="Blackwelder" box="[925,1141,885,908]" pageId="1" pageNumber="28" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, i - iv, 1 - 483." type="journal article" year="1952">Blackwelder (1952)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF936ECBD18FFE59DD3DCD5FAA47108" blockId="1.[151,1437,954,1503]" pageId="1" pageNumber="28">
The genus
<taxonomicName id="4C464D6FBD18FFE59C5FDCD5FE5C77FF" box="[283,431,954,979]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B932EAFEBD18FFE59C5FDCD5FE5C77FF" box="[283,431,954,979]" italics="true" pageId="1" pageNumber="28">Indoquedius</emphasis>
</taxonomicName>
can be easily recognized by the following characters:
<emphasis id="B932EAFEBD18FFE59904DCD5FB7877FF" box="[1088,1163,954,979]" italics="true" pageId="1" pageNumber="28">Head:</emphasis>
dorsal surface of head (and pronotum) very smooth and glossy, lacking microsculpture, two or three setiferous punctures between anterior and posterior frontal setiferous punctures along medial side of eye; labial palpomere II bearing numerous long setae on medial side forming a setal brush (
<figureCitation id="137D2A69BD18FFE59E2DDB45FC397068" box="[873,970,1066,1092]" captionStart="FIGURE 1. A C" captionStartId="2.[151,255,1057,1081]" captionTargetBox="[178,1411,234,1002]" captionTargetId="figure@2.[151,1436,197,1033]" captionTargetPageId="2" captionText="FIGURE 1. A C, Indoquedius baliyo Smetana: A, labial palpus; B, antenna; C, posterior ventral plate; D, Indoquedius nonparallelus sp. nov.: posterior ventral plate. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198061/files/figure.png" pageId="1" pageNumber="28">Fig. 1A</figureCitation>
), last maxillary and labial palpomeres sparsely setose; antennomeres IIII only bearing sparse large setae, lacking dense pubescence, all antennomeres obviously longer than wide (
<figureCitation id="137D2A69BD18FFE59FF2DB1AFCEA70A3" box="[694,793,1141,1167]" captionStart="FIGURE 1. A C" captionStartId="2.[151,255,1057,1081]" captionTargetBox="[178,1411,234,1002]" captionTargetId="figure@2.[151,1436,197,1033]" captionTargetPageId="2" captionText="FIGURE 1. A C, Indoquedius baliyo Smetana: A, labial palpus; B, antenna; C, posterior ventral plate; D, Indoquedius nonparallelus sp. nov.: posterior ventral plate. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198061/files/figure.png" pageId="1" pageNumber="28">Fig. 1B</figureCitation>
).
<emphasis id="B932EAFEBD18FFE59E76DB1AFC6570A2" box="[818,918,1141,1166]" italics="true" pageId="1" pageNumber="28">Thorax:</emphasis>
pronotum usually with two, rarely three, setiferous punctures in each dorsal row, one setiferous puncture and sometimes another less obvious and smaller puncture in each sublateral row; prothorax with additional posterior ventral plate, which divided into two pieces by median longitudinal suture (
<figureCitation id="137D2A69BD18FFE59FDFDB8AFD0B70D3" box="[667,760,1253,1279]" captionStart="FIGURE 1. A C" captionStartId="2.[151,255,1057,1081]" captionTargetBox="[178,1411,234,1002]" captionTargetId="figure@2.[151,1436,197,1033]" captionTargetPageId="2" captionText="FIGURE 1. A C, Indoquedius baliyo Smetana: A, labial palpus; B, antenna; C, posterior ventral plate; D, Indoquedius nonparallelus sp. nov.: posterior ventral plate. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198061/files/figure.png" pageId="1" pageNumber="28">Fig. 1C</figureCitation>
), or entire with anterior median longitudinal ridge (
<figureCitation id="137D2A69BD18FFE59828DB8AFF517108" captionStart="FIGURE 1. A C" captionStartId="2.[151,255,1057,1081]" captionTargetBox="[178,1411,234,1002]" captionTargetId="figure@2.[151,1436,197,1033]" captionTargetPageId="2" captionText="FIGURE 1. A C, Indoquedius baliyo Smetana: A, labial palpus; B, antenna; C, posterior ventral plate; D, Indoquedius nonparallelus sp. nov.: posterior ventral plate. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198061/files/figure.png" pageId="1" pageNumber="28">Fig. 1</figureCitation>
D).
<emphasis id="B932EAFEBD18FFE59D97DA64FEE77108" box="[211,276,1291,1316]" italics="true" pageId="1" pageNumber="28">Legs:</emphasis>
protarsomeres IIV strongly dilated, with ventral surface covered with dense adhesive setae.
</paragraph>
<paragraph id="8BF936ECBD18FFE59D81DA5FFBDF71F3" blockId="1.[151,1437,954,1503]" pageId="1" pageNumber="28">
The genus
<taxonomicName id="4C464D6FBD18FFE59C00DA5FFE267165" box="[324,469,1328,1353]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B932EAFEBD18FFE59C00DA5FFE267165" box="[324,469,1328,1353]" italics="true" pageId="1" pageNumber="28">Indoquedius</emphasis>
</taxonomicName>
is distributed in the Oriental and the Eastern Palaearctic regions (
<figureCitation id="137D2A69BD18FFE59996DA5FFAE97166" box="[1234,1306,1328,1354]" captionStart="FIGURE 2" captionStartId="3.[151,259,1251,1275]" captionTargetBox="[157,1431,216,1213]" captionTargetId="figure@3.[151,1436,197,1230]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of the genus Indoquedius Blackwelder: 1, Indoquedius aberrans Cameron; 2, I. baliyo Smetana; 3, I. bing Smetana; 4, I. bipunctatus Eppelsheim; 5, I. chuen Smetana; 6, I. daai Smetana; 7, I. filicornis Eppelsheim; 8, I. formosae Cameron; 9, I. grandiceps Kraatz; 10, I. hisamatsui Hayashi; 11, I. juno Sharp; 12, I. lasti Rougement; 13, I. liuae Zheng et Wang; 14, I. oculatus Fauvel; 15, I. praeditus Sharp; 16, I. saathi Smetana; 17, I. shibatai Smetana; 18, I. sikkimensis Cameron; 19, I. aculeus sp. nov.; 20, I. bicornutus sp. nov.; 21, I. nonparallelus sp. nov.; 22, I. radius sp. nov.." httpUri="https://zenodo.org/record/198062/files/figure.png" pageId="1" pageNumber="28">Fig. 2</figureCitation>
). In
<collectingCountry id="F351767CBD18FFE5980ADA5FFA647166" box="[1358,1431,1328,1354]" name="China" pageId="1" pageNumber="28">China</collectingCountry>
, six species (numbers 2, 8, 13, 19, 20 and 21) are present in the southwestern area of the country near the Himalayan region, and four (numbers 3, 5, 8 and 17) in the Island of
<collectingCountry id="F351767CBD18FFE59EBDDA15FBA071B8" box="[1017,1107,1402,1428]" name="Taiwan" pageId="1" pageNumber="28">Taiwan</collectingCountry>
(of them three species are endemic). No species of this genus are known from a large territory of North-Eastern and Southern
<collectingCountry id="F351767CBD18FFE59809DACFFA6B7196" box="[1357,1432,1440,1466]" name="China" pageId="1" pageNumber="28">China</collectingCountry>
. Only one species (number 22), described here, occurs in South-Eastern
<collectingCountry id="F351767CBD18FFE59E9ADAAAFBDB71F3" box="[990,1064,1477,1503]" name="China" pageId="1" pageNumber="28">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>