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<document ID-DOI="10.1093/zoolinnean/zlab052" ID-ISSN="0024-4082" ID-Zenodo-Dep="6530695" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1652084412237" checkinUser="plazi" docAuthor="Slavenko, Alex, Tamar, Karin, Tallowin, Oliver J S, Kraus, Fred, Allison, Allen, Carranza, Salvador &amp; Meiri, Shai" docDate="2022" docId="03EF4E777C1ACA74FC12FC69242AFA3A" docLanguage="en" docName="zlab052.pdf" docOrigin="Zoological Journal of the Linnean Society 195 (1)" docSource="https://academic.oup.com/zoolinnean/article/195/1/220/6365875" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Lobulia Greer 1974" docType="treatment" docVersion="5" lastPageNumber="234" masterDocId="FFD6360F7C17CA7AFFB0FFB42066FFA4" masterDocTitle="Revision of the montane New Guinean skink genus Lobulia (Squamata: Scincidae), with the description of four new genera and nine new species" masterLastPageNumber="278" masterPageNumber="220" pageNumber="233" updateTime="1673543820768" updateUser="felipe">
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<mods:title>Revision of the montane New Guinean skink genus Lobulia (Squamata: Scincidae), with the description of four new genera and nine new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Slavenko, Alex</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tamar, Karin</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tallowin, Oliver J S</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kraus, Fred</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Allison, Allen</mods:namePart>
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<mods:namePart>Carranza, Salvador</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Meiri, Shai</mods:namePart>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-05-01</mods:number>
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<mods:number>195</mods:number>
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<mods:identifier type="DOI">10.1093/zoolinnean/zlab052</mods:identifier>
<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6536308" ID-Zenodo-Dep="6536308" LSID="urn:lsid:plazi:treatment:03EF4E777C1ACA74FC12FC69242AFA3A" httpUri="http://treatment.plazi.org/id/03EF4E777C1ACA74FC12FC69242AFA3A" lastPageId="14" lastPageNumber="234" pageId="13" pageNumber="233">
<subSubSection box="[930,1303,989,1014]" pageId="13" pageNumber="233" type="nomenclature">
<paragraph blockId="13.[930,1303,989,1014]" box="[930,1303,989,1014]" pageId="13" pageNumber="233">
<heading box="[930,1303,989,1014]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="233" reason="2">
<taxonomicName authority="GREER, 1974" authorityName="Greer" authorityYear="1974" box="[930,1185,989,1014]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis box="[930,1030,989,1012]" italics="true" pageId="13" pageNumber="233">LOBULIA</emphasis>
<bibRefCitation author="Greer AE" box="[1037,1185,989,1014]" pageId="13" pageNumber="233" pagination="1 - 67" refId="ref44557" refString="Greer AE. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 31: 1 - 67." type="book chapter" year="1974">GREER, 1974</bibRefCitation>
</taxonomicName>
(CLADE I)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="233" type="description">
<paragraph blockId="13.[819,1414,1027,1083]" pageId="13" pageNumber="233">
(
<figureCitation box="[827,945,1027,1052]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-10="Figure 15" captionStart-2="Figure 7" captionStart-3="Figure 8" captionStart-4="Figure 9" captionStart-5="Figure 10" captionStart-6="Figure 11" captionStart-7="Figure 12" captionStart-8="Figure 13" captionStart-9="Figure 14" captionStartId-0="14.[163,243,1109,1131]" captionStartId-1="15.[145,223,1530,1552]" captionStartId-10="24.[163,242,1447,1469]" captionStartId-2="16.[163,241,1520,1542]" captionStartId-3="17.[145,223,926,948]" captionStartId-4="18.[163,242,1594,1616]" captionStartId-5="19.[146,226,580,602]" captionStartId-6="20.[163,243,1575,1597]" captionStartId-7="21.[147,226,734,756]" captionStartId-8="22.[163,242,1550,1572]" captionStartId-9="23.[145,225,545,567]" captionTargetBox-0="[163,1443,196,1069]" captionTargetBox-1="[145,1425,195,1491]" captionTargetBox-10="[419,1187,195,1407]" captionTargetBox-2="[419,1184,195,1480]" captionTargetBox-3="[154,761,201,871]" captionTargetBox-4="[419,1187,195,1554]" captionTargetBox-5="[146,760,195,540]" captionTargetBox-6="[423,1182,196,1532]" captionTargetBox-7="[145,769,195,695]" captionTargetBox-8="[420,1187,196,1511]" captionTargetBox-9="[145,769,201,506]" captionTargetId-0="figure-329@14.[163,1443,195,1069]" captionTargetId-1="figure-158@15.[145,1425,195,1491]" captionTargetId-10="figure-187@24.[419,1187,195,1407]" captionTargetId-2="figure-26@16.[419,1187,195,1480]" captionTargetId-3="figure-775@17.[145,769,195,886]" captionTargetId-4="figure-119@18.[419,1187,195,1554]" captionTargetId-5="figure-732@19.[145,760,195,541]" captionTargetId-6="figure-87@20.[419,1187,192,1536]" captionTargetId-7="figure-743@21.[145,769,195,695]" captionTargetId-8="figure-169@22.[419,1187,195,1511]" captionTargetId-9="figure-857@23.[145,769,195,506]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-10="24" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="21" captionTargetPageId-8="22" captionTargetPageId-9="23" captionText-0="Figure 5. Representatives of each of the seven revised genera, not to scale: (A) Lobulia huonensis (BPBM 40322); (B) Prasinohaema flavipes (BPBM 40369); (C) Nubeoscincus glacialis (BPBM 14712); (D) Papuascincus lineage VII (BPBM 44749); (E) Palaia pulchra (BPBM 38789); (F) Alpinoscincus alpinus (BPBM 44218); (G) Ornithuroscincus pterophilus (BPBM 45705). Images by Allen Allison." captionText-1="Figure 6. Pictures in life of (A) Lobulia brongersmai (BPBM 34733); (B) Lobulia elegans (BPBM 18692); (C) Lobulia fortis (paratype; BPBM 41130); (D) Lobulia huonensis (holotype; BPBM 40322); (E) Lobulia lobulus (BPBM 47837); (F) Lobulia marmorata (holotype; BPBM 34150). Images A, B, F by Fred Kraus, images CE by Allen Allison." captionText-10="Figure 15. Lobulia vogelkopensis holotype (BPBM 6917), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-2="Figure 7. Lobulia lobulus holotype (MCZ R-47067), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Museum of Comparative Zoology, Harvard University." captionText-3="Figure 8. Lobulia lobulus (BPBM 47837) in life, (A) dorsal view and (B) ventral view. Images by Allen Allison." captionText-4="Figure 9. Lobulia fortis holotype (BPBM 41149), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-5="Figure 10. Lobulia fortis holotype (BPBM 41149) in life, (A) dorsal view and (B) ventral view. Images by Allen Allison." captionText-6="Figure 11. Lobulia huonensis holotype (BPBM 40322), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-7="Figure 12. Lobulia huonensis holotype (BPBM 40322), in life, in (A) dorsal view and (B) ventral view. Images by Allen Allison." captionText-8="Figure 13. Lobulia marmorata holotype (BPBM 34150), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-9="Figure 14. Lobulia marmorata holotype (BPBM 34150), ventral view in life. Image by Fred Kraus." figureDoi-0="http://doi.org/10.5281/zenodo.6530705" figureDoi-1="http://doi.org/10.5281/zenodo.6530711" figureDoi-10="http://doi.org/10.5281/zenodo.6530733" figureDoi-2="http://doi.org/10.5281/zenodo.6530713" figureDoi-3="http://doi.org/10.5281/zenodo.6530715" figureDoi-4="http://doi.org/10.5281/zenodo.6530717" figureDoi-5="http://doi.org/10.5281/zenodo.6530719" figureDoi-6="http://doi.org/10.5281/zenodo.6530721" figureDoi-7="http://doi.org/10.5281/zenodo.6530725" figureDoi-8="http://doi.org/10.5281/zenodo.6530729" figureDoi-9="http://doi.org/10.5281/zenodo.6530731" httpUri-0="https://zenodo.org/record/6530705/files/figure.png" httpUri-1="https://zenodo.org/record/6530711/files/figure.png" httpUri-10="https://zenodo.org/record/6530733/files/figure.png" httpUri-2="https://zenodo.org/record/6530713/files/figure.png" httpUri-3="https://zenodo.org/record/6530715/files/figure.png" httpUri-4="https://zenodo.org/record/6530717/files/figure.png" httpUri-5="https://zenodo.org/record/6530719/files/figure.png" httpUri-6="https://zenodo.org/record/6530721/files/figure.png" httpUri-7="https://zenodo.org/record/6530725/files/figure.png" httpUri-8="https://zenodo.org/record/6530729/files/figure.png" httpUri-9="https://zenodo.org/record/6530731/files/figure.png" pageId="13" pageNumber="233">FIGS 515</figureCitation>
; SUPPORTING INFORMATION,
<figureCitation box="[1280,1395,1027,1052]" captionStart="Figure 6" captionStartId="15.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1491]" captionTargetId="figure-158@15.[145,1425,195,1491]" captionTargetPageId="15" captionText="Figure 6. Pictures in life of (A) Lobulia brongersmai (BPBM 34733); (B) Lobulia elegans (BPBM 18692); (C) Lobulia fortis (paratype; BPBM 41130); (D) Lobulia huonensis (holotype; BPBM 40322); (E) Lobulia lobulus (BPBM 47837); (F) Lobulia marmorata (holotype; BPBM 34150). Images A, B, F by Fred Kraus, images CE by Allen Allison." figureDoi="http://doi.org/10.5281/zenodo.6530711" httpUri="https://zenodo.org/record/6530711/files/figure.png" pageId="13" pageNumber="233">FIGS S6S</figureCitation>
<figureCitation box="[1394,1408,1027,1052]" captionStart="Figure 7" captionStartId="16.[163,241,1520,1542]" captionTargetBox="[419,1184,195,1480]" captionTargetId="figure-26@16.[419,1187,195,1480]" captionTargetPageId="16" captionText="Figure 7. Lobulia lobulus holotype (MCZ R-47067), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Museum of Comparative Zoology, Harvard University." figureDoi="http://doi.org/10.5281/zenodo.6530713" httpUri="https://zenodo.org/record/6530713/files/figure.png" pageId="13" pageNumber="233">7</figureCitation>
;
<tableCitation box="[1065,1160,1058,1083]" captionStart="Table 1" captionStartId="9.[142,206,414,436]" captionTargetPageId="9" captionText="Table 1. Ranges of morphometrics and meristics for specimens of species treated in this study (mean values in parentheses). Only adult specimens were examined for morphometric measurements, but juveniles were included for meristics" httpUri="http://table.plazi.org/id/DF39AFE97C1ECA73FF3EFE2A2460FE75" pageId="13" pageNumber="233" tableUuid="DF39AFE97C1ECA73FF3EFE2A2460FE75">TABLE 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="233" type="reference_group">
<paragraph blockId="13.[809,1424,1098,1151]" pageId="13" pageNumber="233">
<taxonomicName authority="Greer, 1974" authorityName="Greer" authorityYear="1974" box="[809,1048,1098,1120]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis box="[809,901,1098,1119]" italics="true" pageId="13" pageNumber="233">Lobulia</emphasis>
<bibRefCitation author="Greer AE" box="[910,1048,1099,1120]" pageId="13" pageNumber="233" pagination="1 - 67" refId="ref44557" refString="Greer AE. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 31: 1 - 67." type="book chapter" year="1974">Greer, 1974</bibRefCitation>
</taxonomicName>
.
<emphasis italics="true" pageId="13" pageNumber="233">Australian Journal of Zoology Supplementary Series</emphasis>
(31): 167.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="233" type="type_taxon">
<paragraph blockId="13.[809,1425,1170,1223]" pageId="13" pageNumber="233">
<emphasis box="[809,959,1171,1192]" italics="true" pageId="13" pageNumber="233">
<typeStatus box="[809,864,1171,1192]" pageId="13" pageNumber="233">Type</typeStatus>
species:
</emphasis>
<taxonomicName authority="Boulenger, 1897" authorityName="Boulenger" authorityYear="1897" box="[973,1382,1170,1192]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="species" species="elegans">
<emphasis box="[973,1184,1170,1192]" italics="true" pageId="13" pageNumber="233">Lygosoma elegans</emphasis>
<bibRefCitation author="Boulenger GA" box="[1192,1382,1170,1192]" pageId="13" pageNumber="233" pagination="6 - 13" refId="ref43763" refString="Boulenger GA. 1897. Descriptions of new lizards and frogs from Mount Victoria, Owen Stanley Range, New Guinea, collected by Mr. A. S. Anthony. Annals and Magazine of Natural History Series 6 19: 6 - 13." type="journal article" year="1897">Boulenger, 1897</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="233" type="diagnosis">
<paragraph blockId="13.[809,1425,1264,1899]" pageId="13" pageNumber="233">
<emphasis box="[809,932,1264,1285]" italics="true" pageId="13" pageNumber="233">Diagnosis:</emphasis>
Medium-sized (adult SVL
<quantity box="[1252,1417,1264,1286]" metricMagnitude="-2" metricUnit="m" metricValue="5.29" metricValueMax="6.39" metricValueMin="4.1899999999999995" pageId="13" pageNumber="233" unit="mm" value="52.9" valueMax="63.9" valueMin="41.9">41.963.9 mm</quantity>
) terrestrial or semi-arboreal skinks with long limbs (forelimbs 32.947.6% of SVL, hindlimbs 41.6 54.8% of SVL); lobules present on anterior edge of ear opening; two pairs of chin shields in medial contact; two supralabials posterior to subocular supralabial; chin shields abutting infralabials; lower eyelid with semi-transparent window; standard three-scale temporal region; nasal scale undivided; frontoparietals either fused or unfused; viviparous; litter size 14.
</paragraph>
<paragraph blockId="13.[809,1425,1264,1899]" pageId="13" pageNumber="233">
<taxonomicName authorityName="Greer" authorityYear="1974" box="[833,925,1601,1622]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis box="[833,925,1601,1622]" italics="true" pageId="13" pageNumber="233">Lobulia</emphasis>
</taxonomicName>
differs from all other genera by its much longer limbs (forelimbs 32.947.6% vs. 27.739.8% of SVL, hindlimbs 41.654.8% vs. 29.949.6% of SVL). It further differs from
<taxonomicName authorityName="GREER" authorityYear="1974" box="[1050,1220,1693,1714]" class="Reptilia" family="Scincidae" genus="Prasinohaema" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis box="[1050,1220,1693,1714]" italics="true" pageId="13" pageNumber="233">Prasinohaema</emphasis>
</taxonomicName>
by lacking green blood serum and tissues (
<bibRefCitation author="Greer AE" box="[1093,1223,1724,1745]" pageId="13" pageNumber="233" pagination="1 - 67" refId="ref44557" refString="Greer AE. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 31: 1 - 67." type="book chapter" year="1974">Greer, 1974</bibRefCitation>
), a prehensile tail with a glandular tip and basally expanded subdigital lamellae. It differs from
<taxonomicName box="[1095,1253,1786,1807]" class="Reptilia" family="Scincidae" genus="Papuascincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis box="[1095,1253,1786,1807]" italics="true" pageId="13" pageNumber="233">Papuascincus</emphasis>
</taxonomicName>
by having two pairs of chin shields in medial contact (vs. one), an undivided (vs. divided) nasal scale and a viviparous (vs. oviparous) reproductive mode.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6530705" ID-Zenodo-Dep="6530705" httpUri="https://zenodo.org/record/6530705/files/figure.png" pageId="14" pageNumber="234" startId="14.[163,243,1109,1131]" targetBox="[163,1443,196,1069]" targetPageId="14">
<paragraph blockId="14.[163,1443,1109,1219]" pageId="14" pageNumber="234">
<emphasis bold="true" box="[163,270,1109,1131]" pageId="14" pageNumber="234">Figure 5.</emphasis>
Representatives of each of the seven revised genera, not to scale: (A)
<taxonomicName authorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" authorityYear="2022" box="[1033,1231,1109,1131]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="huonensis">
<emphasis box="[1033,1231,1109,1131]" italics="true" pageId="14" pageNumber="234">Lobulia huonensis</emphasis>
</taxonomicName>
(BPBM 40322); (B)
<taxonomicName baseAuthorityName="Parker" baseAuthorityYear="1936" box="[163,407,1138,1160]" class="Reptilia" family="Scincidae" genus="Prasinohaema" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="flavipes">
<emphasis box="[163,407,1138,1160]" italics="true" pageId="14" pageNumber="234">Prasinohaema flavipes</emphasis>
</taxonomicName>
(BPBM 40369); (C)
<taxonomicName authorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" authorityYear="2022" baseAuthorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" baseAuthorityYear="2005" box="[624,868,1138,1160]" class="Squamata" family="Scincidae" genus="Nubeoscincus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="glacialis">
<emphasis box="[624,868,1138,1160]" italics="true" pageId="14" pageNumber="234">Nubeoscincus glacialis</emphasis>
</taxonomicName>
(BPBM 14712); (D)
<taxonomicName box="[1086,1231,1138,1160]" class="Reptilia" family="Scincidae" genus="Papuascincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis box="[1086,1231,1138,1160]" italics="true" pageId="14" pageNumber="234">Papuascincus</emphasis>
</taxonomicName>
lineage VII (BPBM 44749); (E)
<taxonomicName authorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" authorityYear="2022" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[288,449,1168,1190]" class="Squamata" family="Scincidae" genus="Palaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="pulchra">
<emphasis box="[288,449,1168,1190]" italics="true" pageId="14" pageNumber="234">Palaia pulchra</emphasis>
</taxonomicName>
(BPBM 38789); (F)
<taxonomicName authorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" authorityYear="2022" baseAuthorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" baseAuthorityYear="2005" box="[667,905,1168,1190]" class="Squamata" family="Scincidae" genus="Alpinoscincus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="alpinus">
<emphasis box="[667,905,1168,1190]" italics="true" pageId="14" pageNumber="234">Alpinoscincus alpinus</emphasis>
</taxonomicName>
(BPBM 44218); (G)
<taxonomicName authorityName="Slavenko &amp; Tamar &amp; Tallowin &amp; Kraus &amp; Allison &amp; Carranza &amp; Meiri" authorityYear="2022" box="[1126,1443,1168,1190]" class="Squamata" family="Scincidae" genus="Ornithuroscincus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="pterophilus">
<emphasis box="[1126,1443,1168,1190]" italics="true" pageId="14" pageNumber="234">Ornithuroscincus pterophilus</emphasis>
</taxonomicName>
(BPBM 45705). Images by Allen Allison.
</paragraph>
</caption>
<subSubSection pageId="14" pageNumber="234" type="discussion">
<paragraph blockId="14.[163,780,1263,1346]" pageId="14" pageNumber="234">
<emphasis box="[163,360,1263,1284]" italics="true" pageId="14" pageNumber="234">Species included:</emphasis>
<taxonomicName authority="Zweifel, 1972" authorityName="Zweifel" authorityYear="1972" box="[374,773,1263,1285]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="brongersmai">
<emphasis box="[374,614,1263,1284]" italics="true" pageId="14" pageNumber="234">Lobulia brongersmai</emphasis>
<bibRefCitation author="Zweifel RG" box="[620,773,1263,1285]" pageId="14" pageNumber="234" pagination="530 - 539" refId="ref46955" refString="Zweifel RG. 1972. A new scincid lizard of the genus Leiolopisma from New Guinea. Zoologische Mededelingen 47: 530 - 539." type="journal article" year="1972">Zweifel, 1972</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="(Boulenger, 1897)" baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[163,575,1293,1315]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="elegans">
<emphasis box="[163,353,1293,1314]" italics="true" pageId="14" pageNumber="234">Lobulia elegans</emphasis>
(
<bibRefCitation author="Boulenger GA" box="[371,564,1293,1315]" pageId="14" pageNumber="234" pagination="6 - 13" refId="ref43763" refString="Boulenger GA. 1897. Descriptions of new lizards and frogs from Mount Victoria, Owen Stanley Range, New Guinea, collected by Mr. A. S. Anthony. Annals and Magazine of Natural History Series 6 19: 6 - 13." type="journal article" year="1897">Boulenger, 1897</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName authority="(Loveridge, 1945)" baseAuthorityName="Loveridge" baseAuthorityYear="1945" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="lobulus">
<emphasis box="[589,779,1293,1314]" italics="true" pageId="14" pageNumber="234">Lobulia lobulus</emphasis>
(
<bibRefCitation author="Loveridge A" box="[171,356,1324,1346]" pageId="14" pageNumber="234" pagination="47 - 52" refId="ref45207" refString="Loveridge A. 1945. New scincid lizards of the genera Tropidophorus and Lygosoma from New Guinea. Proceedings of the Biological Society of Washington 58: 47 - 52." type="journal article" year="1945">Loveridge, 1945</bibRefCitation>
)
</taxonomicName>
; and new species described below.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="234" type="distribution">
<paragraph blockId="14.[163,781,1387,1745]" pageId="14" pageNumber="234">
<emphasis box="[163,320,1387,1408]" italics="true" pageId="14" pageNumber="234">Distribution:</emphasis>
Members of
<taxonomicName authorityName="Greer" authorityYear="1974" box="[487,581,1387,1408]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis box="[487,581,1387,1408]" italics="true" pageId="14" pageNumber="234">Lobulia</emphasis>
</taxonomicName>
are widespread along most of the montane regions of New
<collectingCountry box="[685,773,1417,1439]" name="Guinea-Bissau" pageId="14" pageNumber="234">Guinea</collectingCountry>
, ranging from the central Owen Stanley Mountains in the Papuan Peninsula in the east (
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[610,739,1479,1500]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="elegans">
<emphasis box="[610,739,1479,1500]" italics="true" pageId="14" pageNumber="234">Lo. elegans</emphasis>
</taxonomicName>
) to the Arfak Mountains in the Vogelkop Peninsula in the west. Most species are montane, found at elevations up to
<quantity box="[192,278,1571,1592]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="14" pageNumber="234" unit="m" value="2700.0">2700 m</quantity>
a.s.l.; however,
<taxonomicName authorityName="Zweifel" authorityYear="1972" box="[461,647,1571,1592]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="brongersmai">
<emphasis box="[461,647,1571,1592]" italics="true" pageId="14" pageNumber="234">Lo. brongersmai</emphasis>
</taxonomicName>
is found in the lowlands and hill regions of the northern versant of New
<collectingCountry box="[249,332,1632,1654]" name="Guinea-Bissau" pageId="14" pageNumber="234">Guinea</collectingCountry>
(
<quantity box="[347,459,1632,1653]" metricMagnitude="2" metricUnit="m" metricValue="6.7" metricValueMax="13.4" metricValueMin="0.0" pageId="14" pageNumber="234" unit="m" value="670.0" valueMax="1340.0" valueMin="0.0">01340 m</quantity>
a.s.l.).
<taxonomicName authorityName="Zweifel" authorityYear="1972" box="[541,778,1632,1653]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="brongersmai">
<emphasis box="[541,778,1632,1653]" italics="true" pageId="14" pageNumber="234">Lobulia brongersmai</emphasis>
</taxonomicName>
is the only member of the genus to be found below elevations of
<quantity box="[311,382,1693,1715]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="234" unit="m" value="700.0">700 m</quantity>
, the lowest recorded locality for
<taxonomicName authority="(Kraus, 2020)" baseAuthorityName="Kraus" baseAuthorityYear="2020" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="elegans">
<emphasis italics="true" pageId="14" pageNumber="234">Lo. elegans</emphasis>
(
<bibRefCitation author="Kraus F" box="[261,399,1724,1745]" pageId="14" pageNumber="234" pagination="201 - 214" refId="ref45028" refString="Kraus F. 2020. A new species of Lobulia (Squamata: Scincidae) from Papua New Guinea. Zootaxa 4779: 201 - 214." type="journal article" year="2020">Kraus, 2020</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="234" type="discussion">
<paragraph blockId="14.[163,778,1786,1900]" lastBlockId="14.[827,1443,1262,1438]" pageId="14" pageNumber="234">
<emphasis box="[163,273,1786,1807]" italics="true" pageId="14" pageNumber="234">Remarks:</emphasis>
Molecular evidence suggests that at least two other species not already named or described herein occur in the genus: one in Woitape (BPBM 1868990; WGS 84:
<geoCoordinate box="[403,495,1878,1900]" degrees="8.545" direction="south" orientation="latitude" pageId="14" pageNumber="234" precision="55" value="-8.545">8.545°S</geoCoordinate>
,
<geoCoordinate box="[508,630,1878,1900]" degrees="147.251" direction="east" orientation="longitude" pageId="14" pageNumber="234" precision="55" value="147.251">147.251°E</geoCoordinate>
) and one on Mt Yakapi in the Muller Range (BPBM 34161; WGS 84:
<geoCoordinate box="[874,970,1293,1315]" degrees="5.666" direction="south" orientation="latitude" pageId="14" pageNumber="234" precision="55" value="-5.666">5.666°S</geoCoordinate>
,
<geoCoordinate box="[986,1116,1293,1315]" degrees="142.643" direction="east" orientation="longitude" pageId="14" pageNumber="234" precision="55" value="142.643">142.643°E</geoCoordinate>
). However, since both are only known from a few specimens each, and are not extremely morphologically distinct, we refrain from formally describing them until further material can be collected and examined.
</paragraph>
</subSubSection>
</treatment>
</document>