191 lines
22 KiB
XML
191 lines
22 KiB
XML
<document id="4E79E404BFBE8F1132E6364AB43B5D27" ID-DOI="10.3853/j.0067-1975.61.2009.1518" ID-ISSN="2201-4349" ID-Zenodo-Dep="5297884" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629586993830" checkinUser="felipe" docAuthor="Price, Gilbert J., Zhao, Jian-xin, Feng, Yue-xing & Hocknull, Scott A." docDate="2009" docId="03EE2F75FFD9281CFC76B8FAFE7C218B" docLanguage="en" docName="RecAustMus.61.1.39-48.pdf" docOrigin="Records of the Australian Museum 61 (1)" docSource="http://dx.doi.org/10.3853/j.0067-1975.61.2009.1518" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Phascolarctidae Owen 1839" docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="FFD7570DFFDB2818FFC6B922FFBB2518" masterDocTitle="New Records of Plio-Pleistocene Koalas from Australia: Palaeoecological and Taxonomic Implications" masterLastPageNumber="48" masterPageNumber="39" pageNumber="41" updateTime="1699081618020" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="F583E126DC0BC6CF1DE70B0A57688F32">
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<mods:title id="8651110940F223D3C06E28BC3219CAA5">New Records of Plio-Pleistocene Koalas from Australia: Palaeoecological and Taxonomic Implications</mods:title>
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<mods:namePart id="3E1097693308D8F95D17EC7AB5FEC4C9">Price, Gilbert J.</mods:namePart>
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<mods:roleTerm id="FC318A4C3AA2A634D48D0A199DD7855B">Author</mods:roleTerm>
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<mods:namePart id="D767F1140AC11BD2AE45B2187C27146A">Zhao, Jian-xin</mods:namePart>
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<mods:namePart id="27B3FA006B1CD1E8907DC9DDA756BCA0">Feng, Yue-xing</mods:namePart>
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</mods:name>
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<mods:name id="B89210A62EBC928CC72925B1BE151B51" type="personal">
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<mods:roleTerm id="76F535AF83A0546F37451396E751AFF6">Author</mods:roleTerm>
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<mods:namePart id="4712125C7694DF6A97C4F7860B1D248E">Hocknull, Scott A.</mods:namePart>
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<mods:typeOfResource id="BE3732279398BA44990B72CF1765BB72">text</mods:typeOfResource>
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<mods:title id="10F3D555EBBEB0C3CEC79324380F6813">Records of the Australian Museum</mods:title>
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<mods:part id="19C5F8E9AE5DBFC931ED383CCD2F2650">
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<mods:date id="A74D16BC4718A358230D0C713FC175C6">2009</mods:date>
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<mods:number id="B19858D396DFF5245304C3042D00E97E">2009-05-27</mods:number>
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<mods:detail id="9F49C2B9D9E21497C33D75FB574D93E9" type="volume">
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<mods:number id="B67949160F7366C2CC4F3419B126C4BA">61</mods:number>
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<mods:start id="F67DDA175C0A6C730C501AC021BC3644">39</mods:start>
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<mods:url id="E66C4D348E926256C96128404E7E05F4">http://dx.doi.org/10.3853/j.0067-1975.61.2009.1518</mods:url>
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<mods:classification id="4D449F845135676774220E898D23BFC4">journal article</mods:classification>
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<mods:identifier id="7F785BEFDCAE124F7F60D860160CF25F" type="DOI">10.3853/j.0067-1975.61.2009.1518</mods:identifier>
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<mods:identifier id="1BB6A99C41DFE051EBD9DBE6BB90E84C" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="578700FC20FB9612E6C675F6B03DD1F1" type="Zenodo-Dep">5297884</mods:identifier>
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<treatment id="03EE2F75FFD9281CFC76B8FAFE7C218B" LSID="urn:lsid:plazi:treatment:03EE2F75FFD9281CFC76B8FAFE7C218B" httpUri="http://treatment.plazi.org/id/03EE2F75FFD9281CFC76B8FAFE7C218B" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="41">
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<subSubSection id="C35DCDE8FFD9281AFC76B8FAFAEA24EA" box="[944,1361,472,498]" pageId="2" pageNumber="41" type="nomenclature">
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<paragraph id="8BF89E63FFD9281AFC76B8FAFAEA24EA" blockId="2.[944,1361,472,498]" box="[944,1361,472,498]" pageId="2" pageNumber="41">
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<heading id="D0B0290FFFD9281AFC76B8FAFAEA24EA" box="[944,1361,472,498]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="41" reason="3">
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<emphasis id="B9334271FFD9281AFC76B8FAFAEA24EA" bold="true" box="[944,1361,472,498]" pageId="2" pageNumber="41">
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<taxonomicName id="4C47E5E0FFD9281AFC76B8FAFBC124EA" ID-CoL="6268B" authorityName="Owen" authorityYear="1839" box="[944,1146,472,498]" class="Mammalia" family="Phascolarctidae" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="family">Phascolarctidae</taxonomicName>
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gen. et sp. indet.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35DCDE8FFD9281AFB84BB2FFB04273F" box="[1090,1215,525,551]" pageId="2" pageNumber="41" type="description">
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<paragraph id="8BF89E63FFD9281AFB84BB2FFB04273F" blockId="2.[1090,1215,525,551]" box="[1090,1215,525,551]" pageId="2" pageNumber="41">
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<figureCitation id="137C82E6FFD9281AFB84BB2FFB35273F" box="[1090,1166,525,551]" captionStart="Fig" captionStartId="3.[318,353,1820,1841]" captionTargetBox="[319,1226,263,1803]" captionTargetId="figure-10@3.[319,1226,263,1803]" captionTargetPageId="3" captionText="Fig. 2. Phascolarctid gen et sp. indet. dentary (QMF52287) from site QML7, Chinchilla, eastern Australia. (A) External view, (B) Internal view, and (C) Occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5297888" httpUri="https://zenodo.org/record/5297888/files/figure.png" pageId="2" pageNumber="41">Figs 2</figureCitation>
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,
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<figureCitation id="137C82E6FFD9281AFB5DBB2FFB04273F" box="[1179,1215,525,551]" captionStart="Fig" captionStartId="4.[284,319,732,753]" captionTargetBox="[277,1357,166,706]" captionTargetId="figure-9@4.[265,1369,151,719]" captionTargetPageId="4" captionText="Fig. 3. Morphometrics of fossil koala specimens. (A) Depth versus width of dentaries of modern Phascolarctos cinereus from eastern Queensland (Appendix), fossil Cundokoala (?Ph.) yorkensis (SAMP24904) from Corra Lynn Cave (South Australia), and phascolarctid gen et. sp. indet (QMF52287) from Chinchilla, eastern Australia. Note that the depth of the Chinchilla koala dentary is a minimum measurement because the specimen is broken along the alveolar border (Fig. 2). (B) Anterior versus posterior width of Phascolarctos spp. M2 (See Appendix for list of modern Ph. cinereus specimens examined)." figureDoi="http://doi.org/10.5281/zenodo.5297890" httpUri="https://zenodo.org/record/5297890/files/figure.png" pageId="2" pageNumber="41">3A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35DCDE8FFD9281AFC81BB1EFC072796" pageId="2" pageNumber="41" type="materials_examined">
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<paragraph id="8BF89E63FFD9281AFC81BB1EFC072796" blockId="2.[839,1467,572,654]" pageId="2" pageNumber="41">
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<emphasis id="B9334271FFD9281AFC81BB1EFBA5274C" bold="true" box="[839,1054,572,596]" pageId="2" pageNumber="41">Referred material</emphasis>
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.
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<materialsCitation id="3B2F943EFFD9281AFBF6BB1EFC032796" accessionNumber="QMF52287" country="Australia" location="Chinchilla" pageId="2" pageNumber="41" specimenCount="1" stateProvince="Queensland">
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QMF52287, left dentary fragment, QML7,
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<location id="8E98C8B8FFD9281AFC6EBB78FB932769" LSID="urn:lsid:plazi:treatment:03EE2F75FFD9281CFC76B8FAFE7C218B:8E98C8B8FFD9281AFC6EBB78FB932769" box="[936,1064,602,625]" country="Australia" name="Chinchilla" pageId="2" pageNumber="41" stateProvince="Queensland">Chinchilla</location>
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, southeast
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<collectingRegion id="49835081FFD9281AFB74BB78FAFA2769" box="[1202,1345,602,625]" country="Australia" name="Queensland" pageId="2" pageNumber="41">Queensland</collectingRegion>
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,
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<collectingCountry id="F350DEF3FFD9281AFA89BB78FA002769" box="[1359,1467,602,625]" name="Australia" pageId="2" pageNumber="41">Australia</collectingCountry>
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(Pliocene)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35DCDE8FFD9281AFC81BB93FBBE217B" pageId="2" pageNumber="41" type="description">
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<paragraph id="8BF89E63FFD9281AFC81BB93FBBE217B" blockId="2.[839,1467,689,1123]" pageId="2" pageNumber="41">
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<emphasis id="B9334271FFD9281AFC81BB93FC7C27D1" bold="true" box="[839,967,689,713]" pageId="2" pageNumber="41">Description</emphasis>
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. Dentary fragment with horizontal ramus broken anterior to P
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<subScript id="17C39C26FFD9281AFC0FBBF9FC6A27F2" attach="left" box="[969,977,731,746]" fontSize="6" pageId="2" pageNumber="41">3</subScript>
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alveolus and posterior to M
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<subScript id="17C39C26FFD9281AFB3CBBF9FAB927F2" attach="left" box="[1274,1282,731,746]" fontSize="6" pageId="2" pageNumber="41">3</subScript>
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alveolus; surface bone slightly root-etched; cheek teeth crowns missing, roots present, alveolar border broken buccally; ventral border slightly in-turned lingually; mental foramen anteroventral to P
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<subScript id="17C39C26FFD9281AFCB1BA73FCC42678" attach="left" box="[887,895,849,864]" fontSize="6" pageId="2" pageNumber="41">3</subScript>
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anterior alveolus; posterior mental foramen well developed, ventral to position between posterior root of M
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<subScript id="17C39C26FFD9281AFA74BA4CFA012665" attach="left" box="[1458,1466,878,893]" fontSize="6" pageId="2" pageNumber="41">1</subScript>
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and anterior root of M
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<subScript id="17C39C26FFD9281AFBEFBAA9FB8A2682" attach="left" box="[1065,1073,907,922]" fontSize="6" pageId="2" pageNumber="41">2</subScript>
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; symphysis strongly fused, very deep, extended posteriorly to below posterior root of M
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<subScript id="17C39C26FFD9281AFA9ABA8BFADF26A0" attach="left" box="[1372,1380,937,952]" fontSize="6" pageId="2" pageNumber="41">1</subScript>
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, kinked such that posteroventral border extends below ventral border of horizontal ramus, inclined posteriorly at posteroventral margin, inclined anteriorly at a low angle (35°); genial pit, c.
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<quantity id="4CBF3386FFD9281AFC98BD30FC212131" box="[862,922,1042,1065]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="41" unit="mm" value="3.0">3 mm</quantity>
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largest diameter, present at posterior ventral portion of symphysis, similar symmetrical but broken pit present on opposing dentary.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35DCDE8FFD9281CFC81BDA5FE7C218B" lastPageId="4" lastPageNumber="43" pageId="2" pageNumber="41" type="discussion">
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<paragraph id="8BF89E63FFD9281CFC81BDA5FE7C218B" blockId="2.[839,1467,1159,2003]" lastBlockId="4.[166,794,1060,1171]" lastPageId="4" lastPageNumber="43" pageId="2" pageNumber="41">
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<emphasis id="B9334271FFD9281AFC81BDA5FC142187" bold="true" box="[839,943,1159,1183]" pageId="2" pageNumber="41">Remarks</emphasis>
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. The dentary fragment is considered to be adult based on the presence of well-developed P
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<subScript id="17C39C26FFD9281AFACCBD93FAA921D8" attach="left" box="[1290,1298,1201,1216]" fontSize="6" pageId="2" pageNumber="41">3</subScript>
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and M
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<subScript id="17C39C26FFD9281AFA9BBD93FADE21D8" attach="left" box="[1373,1381,1201,1216]" fontSize="6" pageId="2" pageNumber="41">3</subScript>
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alveoli, and a strongly fused symphysis. The dentary resembles phascolarctids in general morphology. However, QMF52287 does not appear to be referable to any phascolarctid where the dentary is known. It differs from
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<taxonomicName id="4C47E5E0FFD9281AFB03BC38FAFA2029" authorityName="Black & Archer" authorityYear="1997" box="[1221,1345,1306,1329]" class="Mammalia" family="Phascolarctidae" genus="Nimiokoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFB03BC38FAFA2029" box="[1221,1345,1306,1329]" italics="true" pageId="2" pageNumber="41">Nimiokoala</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C47E5E0FFD9281AFA8BBC38FA0F2029" authorityName="Stirton" authorityYear="1957" box="[1357,1460,1306,1329]" class="Mammalia" family="Phascolarctidae" genus="Perikoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFA8BBC38FA0F2029" box="[1357,1460,1306,1329]" italics="true" pageId="2" pageNumber="41">Perikoala</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C47E5E0FFD9281AFC81BC15FC7D2056" box="[839,966,1335,1358]" class="Mammalia" family="Phascolarctidae" genus="Madakoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFC81BC15FC7D2056" box="[839,966,1335,1358]" italics="true" pageId="2" pageNumber="41">Madakoala</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C47E5E0FFD9281AFBCABC15FAAD2056" baseAuthorityName="Bartholomai" baseAuthorityYear="1968" box="[1036,1302,1335,1358]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="cinereus">
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<emphasis id="B9334271FFD9281AFBCABC15FAAD2056" box="[1036,1302,1335,1358]" italics="true" pageId="2" pageNumber="41">Phascolarctos cinereus</emphasis>
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</taxonomicName>
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by: (
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<emphasis id="B9334271FFD9281AFA9CBC15FADC2056" box="[1370,1383,1335,1358]" italics="true" pageId="2" pageNumber="41">a</emphasis>
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) being significantly more robust in terms of depth and width (
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<figureCitation id="137C82E6FFD9281AFA49BC76FCD02091" captionStart="Fig" captionStartId="4.[284,319,732,753]" captionTargetBox="[277,1357,166,706]" captionTargetId="figure-9@4.[265,1369,151,719]" captionTargetPageId="4" captionText="Fig. 3. Morphometrics of fossil koala specimens. (A) Depth versus width of dentaries of modern Phascolarctos cinereus from eastern Queensland (Appendix), fossil Cundokoala (?Ph.) yorkensis (SAMP24904) from Corra Lynn Cave (South Australia), and phascolarctid gen et. sp. indet (QMF52287) from Chinchilla, eastern Australia. Note that the depth of the Chinchilla koala dentary is a minimum measurement because the specimen is broken along the alveolar border (Fig. 2). (B) Anterior versus posterior width of Phascolarctos spp. M2 (See Appendix for list of modern Ph. cinereus specimens examined)." figureDoi="http://doi.org/10.5281/zenodo.5297890" httpUri="https://zenodo.org/record/5297890/files/figure.png" pageId="2" pageNumber="41">Fig. 3A</figureCitation>
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); (
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<emphasis id="B9334271FFD9281AFC4FBC50FC2D2091" box="[905,918,1394,1417]" italics="true" pageId="2" pageNumber="41">b</emphasis>
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) possessing a more anteriorly positioned posterior mental foramen; (
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<emphasis id="B9334271FFD9281AFBD2BCADFB9B20BE" box="[1044,1056,1423,1446]" italics="true" pageId="2" pageNumber="41">c</emphasis>
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) having a straighter ventral border; and (
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<emphasis id="B9334271FFD9281AFC47BC8FFC3520DC" box="[897,910,1453,1476]" italics="true" pageId="2" pageNumber="41">d</emphasis>
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) by possessing a relatively deeper symphysis that has a lower anterior ascending angle. QMF52287 is more gracile in comparison to
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<taxonomicName id="4C47E5E0FFD9281AFBA6BCC5FAE920E6" box="[1120,1362,1511,1534]" class="Mammalia" family="Phascolarctidae" genus="Cundokoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="yorkensis">
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<emphasis id="B9334271FFD9281AFBA6BCC5FAE920E6" box="[1120,1362,1511,1534]" italics="true" pageId="2" pageNumber="41">Cundokoala yorkensis</emphasis>
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</taxonomicName>
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(a genus that is questionably distinct from
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<taxonomicName id="4C47E5E0FFD9281AFB6FBF27FAFA2304" authorityName="Blainville" authorityYear="1816" box="[1193,1345,1541,1564]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFB6FBF27FAFA2304" box="[1193,1345,1541,1564]" italics="true" pageId="2" pageNumber="41">Phascolarctos</emphasis>
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</taxonomicName>
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; see
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<bibRefCitation id="EFD6E392FFD9281AFAB0BF26FCC52321" author="Black, K" pageId="2" pageNumber="41" pagination="16 - 17" refId="ref7100" refString="Black, K., 1999. Diversity and relationships of living and extinct koalas (Phascolarctidae, Marsupialia). Australian Mammalogy 21: 16 - 17." type="journal article" year="1999">Black, 1999</bibRefCitation>
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) (
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<figureCitation id="137C82E6FFD9281AFC5EBF00FC4A2321" box="[920,1009,1570,1593]" captionStart="Fig" captionStartId="4.[284,319,732,753]" captionTargetBox="[277,1357,166,706]" captionTargetId="figure-9@4.[265,1369,151,719]" captionTargetPageId="4" captionText="Fig. 3. Morphometrics of fossil koala specimens. (A) Depth versus width of dentaries of modern Phascolarctos cinereus from eastern Queensland (Appendix), fossil Cundokoala (?Ph.) yorkensis (SAMP24904) from Corra Lynn Cave (South Australia), and phascolarctid gen et. sp. indet (QMF52287) from Chinchilla, eastern Australia. Note that the depth of the Chinchilla koala dentary is a minimum measurement because the specimen is broken along the alveolar border (Fig. 2). (B) Anterior versus posterior width of Phascolarctos spp. M2 (See Appendix for list of modern Ph. cinereus specimens examined)." figureDoi="http://doi.org/10.5281/zenodo.5297890" httpUri="https://zenodo.org/record/5297890/files/figure.png" pageId="2" pageNumber="41">Fig. 3A</figureCitation>
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), but like
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<taxonomicName id="4C47E5E0FFD9281AFBA1BF00FB4B2321" baseAuthorityName="Bartholomai" baseAuthorityYear="1968" box="[1127,1264,1570,1593]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="cinereus">
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<emphasis id="B9334271FFD9281AFBA1BF00FB4B2321" box="[1127,1264,1570,1593]" italics="true" pageId="2" pageNumber="41">Ph. cinereus</emphasis>
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</taxonomicName>
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, also differs from the former taxon in having a more anteriorly positioned posterior mental foramen, straighter ventral border, and deeper symphysis. The dentary of
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<taxonomicName id="4C47E5E0FFD9281AFB6DBF58FAAB2389" box="[1195,1296,1658,1681]" class="Mammalia" family="Phascolarctidae" genus="Litokoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFB6DBF58FAAB2389" box="[1195,1296,1658,1681]" italics="true" pageId="2" pageNumber="41">Litokoala</emphasis>
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</taxonomicName>
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is unknown, but on the basis of alveoli dimensions, is probably significantly smaller than QMF52287. QMF52287 differs from
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<taxonomicName id="4C47E5E0FFD9281AFA93BF97FA0023D4" authorityName="Stirton" authorityYear="1957" box="[1365,1467,1717,1740]" class="Mammalia" family="Phascolarctidae" genus="Perikoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFA93BF97FA0023D4" box="[1365,1467,1717,1740]" italics="true" pageId="2" pageNumber="41">Perikoala</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C47E5E0FFD9281AFCBFBFF0FC4D23F1" box="[889,1014,1746,1769]" class="Mammalia" family="Phascolarctidae" genus="Madakoala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFCBFBFF0FC4D23F1" box="[889,1014,1746,1769]" italics="true" pageId="2" pageNumber="41">Madakoala</emphasis>
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</taxonomicName>
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in that the symphysis is more strongly fused, and the diastema is relatively longer. Well-developed symmetrical genial pits are present at the posteroventral base of the symphysis of QMF52287, but are not expressed as strongly in other koala genera. Well-developed genial pits occur in wombats such as
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<taxonomicName id="4C47E5E0FFD9281AFB55BE47FB402264" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1803" box="[1171,1275,1893,1916]" class="Mammalia" family="Vombatidae" genus="Vombatus" kingdom="Animalia" order="Diprotodontia" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFD9281AFB55BE47FB402264" box="[1171,1275,1893,1916]" italics="true" pageId="2" pageNumber="41">Vombatus</emphasis>
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</taxonomicName>
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, but such pits are relatively larger, positioned relatively higher from the ventral border, and the pits commonly converge on each other on the left and right side of the symphyseal fusion forming a single large pit. Comparison of QMF52287 to other largesized phascolarctids such as
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<taxonomicName id="4C47E5E0FFDF281CFE08BD63FD882140" authorityName="Pledge" authorityYear="1987" box="[462,563,1089,1112]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="maris">
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<emphasis id="B9334271FFDF281CFE08BD63FD882140" box="[462,563,1089,1112]" italics="true" pageId="4" pageNumber="43">Ph. maris</emphasis>
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</taxonomicName>
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, or unusual koala-like marsupials such as
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<taxonomicName id="4C47E5E0FFDF281CFEA8BD7DFE05216E" authorityName="Archer & Wade" authorityYear="1976" box="[366,446,1119,1142]" class="Mammalia" family="Phascolarctidae" genus="Koobor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="4" pageNumber="43" phylum="Chordata" rank="genus">
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<emphasis id="B9334271FFDF281CFEA8BD7DFE05216E" box="[366,446,1119,1142]" italics="true" pageId="4" pageNumber="43">Koobor</emphasis>
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</taxonomicName>
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, is not possible because dentaries of those taxa are unknown.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |