127 lines
14 KiB
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127 lines
14 KiB
XML
<document id="916CCC8ECB5E73CD3844D4F2358960C7" ID-CLB-Dataset="24935" ID-DOI="10.11646/zootaxa.4695.3.1" ID-GBIF-Dataset="256591b2-6d7b-465d-a818-9a158017ca09" ID-ISSN="1175-5326" ID-Zenodo-Dep="3532515" ID-ZooBank="32E17629-F29A-47C8-ACCB-69A32C17B147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573197328991" checkinUser="plazi" docAuthor="Gunter, Nicole L. & Weir, Thomas A." docDate="2019" docId="03ED0167FF87FFBEFF1DF644FB6FFECF" docLanguage="en" docName="zootaxa.4695.3.1.pdf" docOrigin="Zootaxa 4695 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus australis Matthews 1974" docType="treatment" docVersion="9" lastPageNumber="204" masterDocId="FFD4791FFF85FFBDFF8AF379FF9DFFA8" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups" masterLastPageNumber="252" masterPageNumber="201" pageNumber="203" updateTime="1698764404900" updateUser="ExternalLinkService">
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<mods:mods id="201202CBF4AA8EF76075713E8DC3585D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2E5D7DB2B8A98F38C58597A2BA409F4F">
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<mods:title id="3F5E98008642FC65B95387B25AA6A773">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups</mods:title>
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</mods:titleInfo>
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<mods:name id="79B8766E572A4678C8C7B0C14632A0A0" type="personal">
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<mods:roleTerm id="81B3EFC8063CBE1920A1797A767E0D71">Author</mods:roleTerm>
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<mods:namePart id="DCA11569DF96EEE13F48C27C992AD58D">Gunter, Nicole L.</mods:namePart>
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</mods:name>
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<mods:name id="DB3AE402FC86AFF836F68C952F583794" type="personal">
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<mods:role id="3F41EEB45B44CF080AD23B1E019DBFC1">
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<mods:roleTerm id="4D028C772ED5E6D11871F5F39706673D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A6B607EF3F534649B3CF0DE95E5553C7">Weir, Thomas A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A4F05D47DB65F72E877B780E089157FF">text</mods:typeOfResource>
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<mods:titleInfo id="F4D577BEBEA156BD17DE707D532B1657">
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<mods:title id="BA67627712869F1F7B812104FC903C93">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B37ACC843657330C9FEB247ADB43AEEF">
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<mods:date id="40DB30EDBBDD4E110C94EA06F892E75E">2019</mods:date>
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<mods:detail id="7F05668DD85FA8A08C3A7E6D72556076" type="pubDate">
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<mods:number id="84999DC82086954D3A94EBF092B08800">2019-11-08</mods:number>
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<mods:detail id="7DAD97D541B63B7CE3018580FA694126" type="volume">
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<mods:number id="898F362BE8FE2D79157052F5A6AEF051">4695</mods:number>
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</mods:detail>
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<mods:detail id="99CC00D5230DB20303F7182D2A4788E7" type="issue">
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<mods:number id="D2A28AD2C3D73BC450D6FD59A9801B7A">3</mods:number>
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<mods:classification id="17A8B608E0180D1DDAA988EB5E4EE28F">journal article</mods:classification>
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<mods:identifier id="021F36D65A04AAD792218AFE23B12E04" type="CLB-Dataset">24935</mods:identifier>
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<mods:identifier id="7AF1A1D48AE855DC41E6ED3C24CE18EB" type="DOI">10.11646/zootaxa.4695.3.1</mods:identifier>
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<mods:identifier id="41A87A075959E6A1586CEE5F941A1DED" type="GBIF-Dataset">256591b2-6d7b-465d-a818-9a158017ca09</mods:identifier>
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<mods:identifier id="66F7DBAF14EAD540B6FC8EBB9A7170AD" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="648C961EE7085C55B96696D9CFDB4BC8" type="Zenodo-Dep">3532515</mods:identifier>
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<mods:identifier id="1C0E901AB0F0CD4AAC1C7EE04EA3E820" type="ZooBank">32E17629-F29A-47C8-ACCB-69A32C17B147</mods:identifier>
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<treatment id="03ED0167FF87FFBEFF1DF644FB6FFECF" ID-DOI="http://doi.org/10.5281/zenodo.5942873" ID-GBIF-Taxon="160218008" ID-Zenodo-Dep="5942873" LSID="urn:lsid:plazi:treatment:03ED0167FF87FFBEFF1DF644FB6FFECF" httpUri="http://treatment.plazi.org/id/03ED0167FF87FFBEFF1DF644FB6FFECF" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
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<subSubSection id="C35EE3FAFF87FFBFFF1DF644FD83FAFF" box="[151,542,1341,1368]" pageId="2" pageNumber="203" type="nomenclature">
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<paragraph id="8BFBB071FF87FFBFFF1DF644FD83FAFF" blockId="2.[151,542,1341,1368]" box="[151,542,1341,1368]" pageId="2" pageNumber="203">
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<heading id="D0B3071DFF87FFBFFF1DF644FD83FAFF" bold="true" box="[151,542,1341,1368]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
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<emphasis id="B9306C63FF87FFBFFF1DF644FD83FAFF" bold="true" box="[151,542,1341,1368]" pageId="2" pageNumber="203">
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<taxonomicName id="4C44CBF2FF87FFBFFF1DF644FEF2FAF0" ID-CoL="6P7WD" authorityName="Matthews" authorityYear="1974" box="[151,367,1341,1368]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B9306C63FF87FFBFFF1DF644FEF2FAF0" bold="true" box="[151,367,1341,1368]" italics="true" pageId="2" pageNumber="203">Lepanus australis</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35EE3FAFF87FFBEFF1DF6FDFB6FFECF" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203" type="diagnosis">
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<paragraph id="8BFBB071FF87FFBFFF1DF6FDFCC1FA6B" blockId="2.[151,1437,1412,2016]" pageId="2" pageNumber="203">
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<emphasis id="B9306C63FF87FFBFFF1DF6FDFE8FFA37" bold="true" box="[151,274,1412,1439]" pageId="2" pageNumber="203">Diagnosis.</emphasis>
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Species within the
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<taxonomicName id="4C44CBF2FF87FFBFFE61F6FFFDF7FA37" authorityName="Matthews" authorityYear="1974" box="[491,618,1413,1439]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B9306C63FF87FFBFFE61F6FFFE66FA37" box="[491,507,1414,1439]" italics="true" pageId="2" pageNumber="203">L</emphasis>
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.
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<emphasis id="B9306C63FF87FFBFFD82F6FCFDF7FA37" box="[520,618,1413,1439]" italics="true" pageId="2" pageNumber="203">australis</emphasis>
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</taxonomicName>
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species group share the following characteristics, with occasional exceptions noted in brackets. Note, the species group is paraphyletic.
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</paragraph>
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<paragraph id="8BFBB071FF87FFBFFF4DF6B4FDC6F9DF" blockId="2.[151,1437,1412,2016]" pageId="2" pageNumber="203">
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<emphasis id="B9306C63FF87FFBFFF4DF6B4FE96FA4F" box="[199,267,1485,1511]" italics="true" pageId="2" pageNumber="203">Males</emphasis>
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. Head: Surface smooth and nitid. Broadly U-shaped between clypeal teeth that are short and not upturned (very broad U-shape in
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<taxonomicName id="4C44CBF2FF87FFBFFE2EF68BFDDCF9A3" authority="Gunter & Weir, 2019" authorityName="Gunter & Weir" authorityYear="2019" box="[420,577,1521,1547]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="dukungarri" status="sp. nov.">
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<emphasis id="B9306C63FF87FFBFFE2EF68BFDDCF9A3" box="[420,577,1521,1547]" italics="true" pageId="2" pageNumber="203">L. dukungarri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203D118FF87FFBFFDC3F688FD4EF9A3" box="[585,723,1521,1547]" pageId="2" pageNumber="203" rank="species">new species</taxonomicNameLabel>
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, clypeal teeth slightly upturned in
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<taxonomicName id="4C44CBF2FF87FFBFFBD0F68BFB21F9A3" authority="Gunter & Weir, 2019" authorityName="Gunter & Weir" authorityYear="2019" box="[1114,1212,1521,1547]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="oxleyi" status="sp. nov.">
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<emphasis id="B9306C63FF87FFBFFBD0F68BFB21F9A3" box="[1114,1212,1521,1547]" italics="true" pageId="2" pageNumber="203">L. oxleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203D118FF87FFBFFB49F688FACDF9A4" box="[1219,1360,1521,1548]" pageId="2" pageNumber="203" rank="species">new species</taxonomicNameLabel>
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). Edge of head completely bordered; clypeal edge normal, without projection lateral to teeth; margin very slightly angulate at clypeogenal suture; genal angle rounded; margin of head rounded behind the eyes; mentum not bordered, without striations. Dorsal part of the eye narrow.
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</paragraph>
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<paragraph id="8BFBB071FF87FFBFFF4DF5F8FC6CF917" blockId="2.[151,1437,1412,2016]" pageId="2" pageNumber="203">Prothorax: Basal angles distinct. Lateral edges margined. Punctures along the basal margin similar to those posteriorly on disc, not extending to basal margin. Hypomeral striae present.</paragraph>
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<paragraph id="8BFBB071FF87FFBFFF4DF5B0FCA8F94B" blockId="2.[151,1437,1412,2016]" box="[199,821,1737,1763]" pageId="2" pageNumber="203">Elytra: Surface smooth with variously impressed striae.</paragraph>
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<paragraph id="8BFBB071FF87FFBFFF4DF594FE84F83F" blockId="2.[151,1437,1412,2016]" pageId="2" pageNumber="203">
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Legs: Protibiae with 2 teeth on outer edge. Front edge of protibiae with apical digit but apical spur absent; apical digit long and parallel sided with slight inner apical angle (except
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<taxonomicName id="4C44CBF2FF87FFBFFC79F46BFBE9F883" authority="Gunter & Weir, 2019" authorityName="Gunter & Weir" authorityYear="2019" box="[1011,1140,1809,1835]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="eungella" status="sp. nov.">
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<emphasis id="B9306C63FF87FFBFFC79F46BFBE9F883" box="[1011,1140,1809,1835]" italics="true" pageId="2" pageNumber="203">L. eungella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203D118FF87FFBFFBF5F468FA90F883" box="[1151,1293,1809,1835]" pageId="2" pageNumber="203" rank="species">new species</taxonomicNameLabel>
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, apical digit short and triangular). Metatibiae not modified, without extension (metatibiae of
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<taxonomicName id="4C44CBF2FF87FFBFFB92F44FFB29F8E7" authority="Gunter & Weir, 2019" authorityName="Gunter & Weir" authorityYear="2019" box="[1048,1204,1845,1871]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="dukungarri" status="sp. nov.">
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<emphasis id="B9306C63FF87FFBFFB92F44FFB29F8E7" box="[1048,1204,1845,1871]" italics="true" pageId="2" pageNumber="203">L. dukungarri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203D118FF87FFBFFB31F44CFADCF8E7" box="[1211,1345,1845,1871]" pageId="2" pageNumber="203" rank="species">new species</taxonomicNameLabel>
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slightly curved). Metatarsi with tarsomere 2 the longest. Tarsomere 1 normal, not lobed on inner edge. Tarsal claws small and simple.
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</paragraph>
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<paragraph id="8BFBB071FF87FFBFFF4DF4D8FE46F877" blockId="2.[151,1437,1412,2016]" pageId="2" pageNumber="203">
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Abdomen: Pygidium simple, finely punctate (more coarsely punctate in some specimens of
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<taxonomicName id="4C44CBF2FF87FFBFFB41F4DBFAFAF813" authority="Gunter & Weir, 2019" authorityName="Gunter & Weir" authorityYear="2019" box="[1227,1383,1953,1979]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="dukungarri" status="sp. nov.">
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<emphasis id="B9306C63FF87FFBFFB41F4DBFAFAF813" box="[1227,1383,1953,1979]" italics="true" pageId="2" pageNumber="203">L. dukungarri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203D118FF87FFBFFAE4F4D8FF77F848" pageId="2" pageNumber="203" rank="species">new species</taxonomicNameLabel>
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) and slightly convex.
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</paragraph>
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<paragraph id="8BFBB071FF86FFBEFF4DF3E0FCC7FF1B" blockId="3.[151,1436,153,360]" box="[199,858,153,179]" pageId="3" pageNumber="204">Pterothorax: Meso-metasternal suture very slightly arched.</paragraph>
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<paragraph id="8BFBB071FF86FFBEFF4DF3C4FD88FF7F" blockId="3.[151,1436,153,360]" box="[199,533,189,215]" pageId="3" pageNumber="204">Hind wings: Fully developed.</paragraph>
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<paragraph id="8BFBB071FF86FFBEFF4DF398FD81FEB7" blockId="3.[151,1436,153,360]" pageId="3" pageNumber="204">
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Genitalia: Aedeagus with parameres asymmetrical, right paramere a little longer and with apex variously bent downwards and expanded (
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<figureCitation id="137FACF4FF86FFBEFE43F27CFD93FEB7" box="[457,526,261,287]" captionStart="FIGURE 3" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[264,1312,179,1899]" captionTargetId="figure@11.[262,1324,166,1904]" captionTargetPageId="11" captionText="FIGURE 3. Left, dorsal, and right view of aedeagi of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532519" httpUri="https://zenodo.org/record/3532519/files/figure.png" pageId="3" pageNumber="204">Fig. 3</figureCitation>
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).
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</paragraph>
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<paragraph id="8BFBB071FF86FFBEFF4DF250FB6FFECF" blockId="3.[151,1436,153,360]" pageId="3" pageNumber="204">
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<emphasis id="B9306C63FF86FFBEFF4DF250FEBBFEEB" box="[199,294,297,323]" italics="true" pageId="3" pageNumber="204">Females</emphasis>
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. Protibiae with long apical spur and shorter apical digit (except for
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<taxonomicName id="4C44CBF2FF86FFBEFBB0F253FA9FFEEB" authority="Gunter & Weir, 2019" authorityName="Gunter & Weir" authorityYear="2019" box="[1082,1282,297,323]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="eungella" status="sp. nov.">
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<emphasis id="B9306C63FF86FFBEFBB0F253FA9FFEEB" box="[1082,1282,297,323]" italics="true" pageId="3" pageNumber="204">Lepanus eungella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203D118FF86FFBEFA86F250FA05FEEB" box="[1292,1432,297,323]" pageId="3" pageNumber="204" rank="species">new species</taxonomicNameLabel>
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, where apical digit is absent on truncate protibia). Abdominal ventrite 6 slightly longer than in male.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |