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<document id="042A6355100E59A02C136001AAC84C87" ID-CLB-Dataset="21955" ID-DOI="10.5852/ejt.2017.368" ID-GBIF-Dataset="7d707e31-1104-4ed3-8493-ef9302b98796" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838412" ID-ZooBank="E4AEAA16-A546-48B8-99BA-2E0152C8C6F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590087673171" checkinUser="carolina" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2017" docId="03E9996C145E4E6EB53E6EB48594DEDF" docLanguage="en" docName="ejt-2017-368.pdf.imf" docOrigin="European Journal of Taxonomy 368" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phtheochroa cantabriana Zlatkov &amp; Huemer 2017, sp. nov." docType="treatment" docUuid="AB5396E9-E6BF-487D-8CC3-D5E7F2B1674E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="20" masterDocId="FFD0E114144F4E7DB7706C11845EDC35" masterDocTitle="Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="18" updateTime="1698840706466" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="811408356552788EC1B2F9C1F59706F3">Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)</mods:title>
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<mods:name id="B4A7BA3921F9B26B022D2D743A090A96" type="personal">
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<mods:namePart id="61224ACB2DCB9A1080F81894FBB6CE2F">Zlatkov, Boyan</mods:namePart>
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<mods:affiliation id="7357917DF1AD4B5DE5C452D484113646">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; urn: lsid: zoobank. org: author: 3538 D 3 F 5 - 6822 - 403 F-B 9 E 6 - E 8 A 95 E 884404 &amp; Corresponding author: bzlatkov @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="1C597A07CC395D74C40B3847980ABC62" type="email">bzlatkov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="82013C4ABB95F1A3D976669E731A195F">Huemer, Peter</mods:namePart>
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<mods:affiliation id="22E7C7E176DAB9C25E7AF1B9A2A28AFE">Tiroler Landesmuseen Betriebsgesellschaft m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria. &amp; Email: p. huemer @ tiroler-landesmuseen. at &amp; urn: lsid: zoobank. org: author: 4 E 6 F 001 E-DB 9 D- 460 C-AB 39 - 0 CE 7 CC 773 B 59</mods:affiliation>
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<mods:title id="25C64771E94645191AD4888668D1B012">European Journal of Taxonomy</mods:title>
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<mods:date id="BBD22C6B9D33EADECFF1370016BFF396">2017</mods:date>
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<mods:number id="DAFD722148B63E28B3A67DBA1D8915D7">2017-11-21</mods:number>
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<treatment id="03E9996C145E4E6EB53E6EB48594DEDF" ID-DOI="http://doi.org/10.5281/zenodo.3852272" ID-GBIF-Taxon="164246397" ID-Zenodo-Dep="3852272" LSID="urn:lsid:zoobank.org:act:AB5396E9-E6BF-487D-8CC3-D5E7F2B1674E" httpUri="http://treatment.plazi.org/id/03E9996C145E4E6EB53E6EB48594DEDF" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C35A7BF1145E4E6CB53E6EB487B8DEF5" box="[590,998,677,704]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BFF287A145E4E6CB53E6EB487B8DEF5" blockId="17.[363,1225,677,774]" box="[590,998,677,704]" pageId="17" pageNumber="18">
<heading id="D0B79F16145E4E6CB53E6EB487B8DEF5" box="[590,998,677,704]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="4C4053F9145E4E6CB53E6EB487DADEF5" ID-CoL="B3ZCG" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[590,900,677,704]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F468145E4E6CB53E6EB487DADEF5" bold="true" box="[590,900,677,704]" italics="true" pageId="17" pageNumber="18">Phtheochroa cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913145E4E6CB4FC6EB787B8DEF5" box="[908,998,678,704]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C35A7BF1145E4E6CB61B6ED987A3DF33" pageId="17" pageNumber="18" type="description">
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<uri id="FFD12478145E4E6CB61B6ED98097DED6" box="[363,1225,712,739]" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BFF287A145E4E6CB5466EFD87A3DF33" blockId="17.[363,1225,677,774]" box="[566,1021,748,774]" pageId="17" pageNumber="18">
<figureCitation id="137B34FF145E4E6CB5466EFD8624DF33" box="[566,634,748,774]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="17" pageNumber="18">Figs 1</figureCitation>
I–J, 2F, 3F, 7C–D, 9, 10;
<tableCitation id="C6C21DC1145E4E6CB4D66EFD87A3DF33" box="[934,1021,748,774]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="17" pageNumber="18" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1145E4E6CB7CD6F2486A4D807" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BFF287A145E4E6CB7CD6F248568DF7A" blockId="17.[189,310,821,847]" box="[189,310,821,847]" pageId="17" pageNumber="18">
<heading id="D0B79F16145E4E6CB7CD6F248568DF7A" bold="true" box="[189,310,821,847]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B934F468145E4E6CB7CD6F248568DF7A" bold="true" box="[189,310,821,847]" pageId="17" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A145E4E6CB7CD6F7686A4D807" blockId="17.[189,1399,871,1074]" pageId="17" pageNumber="18">
Externally, this species is practically indistinguishable from
<taxonomicName id="4C4053F9145E4E6CB4F76F79804CDFB4" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[903,1042,871,897]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F468145E4E6CB4F76F79804CDFB4" box="[903,1042,871,897]" italics="true" pageId="17" pageNumber="18">P. frigidana</emphasis>
</taxonomicName>
; the same is valid for most of the genital characters, but the vesica is strikingly different. The general shape of the vesica resembles that of
<taxonomicName id="4C4053F9145E4E6CB6606FBF85E9DFFD" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[272,439,942,968]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F468145E4E6CB6606FBF85E9DFFD" box="[272,439,942,968]" italics="true" pageId="17" pageNumber="18">P. schawerdae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4053F9145E4E6CB6806FBF86C9DFF2" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[496,663,942,967]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F468145E4E6CB6806FBF86C9DFF2" box="[496,663,942,967]" italics="true" pageId="17" pageNumber="18">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913145E4E6CB5EE6FBF86A6DFFD" box="[670,760,942,968]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
but in lateral left view its main part looks angular (not ovoid), and the angle between the axes of the phallus and right diverticulum (in dorsal view) is usually larger (more than 130°). The wing upperside colouration of
<taxonomicName id="4C4053F9145E4E6CB4F46FE4806ED83B" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[900,1072,1012,1038]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F468145E4E6CB4F46FE4806ED83B" box="[900,1072,1012,1038]" italics="true" pageId="17" pageNumber="18">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913145E4E6CB3496FE580CAD83B" box="[1081,1172,1012,1038]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is much paler than that of
<taxonomicName id="4C4053F9145E4E6CB660680985E6D804" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[272,440,1047,1073]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F468145E4E6CB660680985E6D804" box="[272,440,1047,1073]" italics="true" pageId="17" pageNumber="18">P. schawerdae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4053F9145E4E6CB681680986C7D804" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[497,665,1048,1073]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F468145E4E6CB681680986C7D804" box="[497,665,1048,1073]" italics="true" pageId="17" pageNumber="18">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913145E4E6CB5D0680986A4D807" box="[672,762,1048,1074]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1145E4E6CB7CD6871814AD899" box="[189,1300,1120,1197]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BFF287A145E4E6CB7CD6871851AD84F" blockId="17.[189,324,1120,1146]" box="[189,324,1120,1146]" pageId="17" pageNumber="18">
<heading id="D0B79F16145E4E6CB7CD6871851AD84F" bold="true" box="[189,324,1120,1146]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B934F468145E4E6CB7CD6871851AD84F" bold="true" box="[189,324,1120,1146]" pageId="17" pageNumber="18">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A145E4E6CB7CD6883814AD899" blockId="17.[189,1300,1170,1197]" box="[189,1300,1170,1197]" pageId="17" pageNumber="18">The specific name is a feminine adjective derived from the name of the Cantabrian Mountains.</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1145E4E6CB7CD68CA8573DBF7" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BFF287A145E4E6CB7CD68CA85F4D8C0" blockId="17.[189,426,1243,1269]" box="[189,426,1243,1269]" pageId="17" pageNumber="18">
<heading id="D0B79F16145E4E6CB7CD68CA85F4D8C0" bold="true" box="[189,426,1243,1269]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B934F468145E4E6CB7CD68CA85F4D8C0" bold="true" box="[189,426,1243,1269]" pageId="17" pageNumber="18">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B282227145E4E6CB79A691F8129D9F1" ID-GBIF-Occurrence="2621725303" collectingDate="2012-11-07" collectionCode="TLMF" collectorName="PN Picos de Europa Espinama &amp; Huemer &amp; Lep &amp; Gen." country="Spain" elevation="1340" latitude="43.148666" location="Portillas de Poqueion" longLatPrecision="11" longitude="-4.7755" municipality="Picos de Europa Espinama" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Cantabria" typeStatus="holotype">
<paragraph id="8BFF287A145E4E6CB79A691F8502D91D" blockId="17.[234,348,1294,1320]" box="[234,348,1294,1320]" pageId="17" pageNumber="18">
<emphasis id="B934F468145E4E6CB79A691F8502D91D" bold="true" box="[234,348,1294,1320]" pageId="17" pageNumber="18">
<typeStatus id="54FB96D8145E4E6CB79A691F8502D91D" box="[234,348,1294,1320]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFF287A145E4E6CB7CD69518129D9F1" blockId="17.[189,1399,1344,1476]" pageId="17" pageNumber="18">
<collectingCountry id="F35768EA145E4E6CB7CD6951854AD96F" box="[189,276,1344,1370]" name="Spain" pageId="17" pageNumber="18">SPAIN</collectingCountry>
:
<specimenCount id="9D46E3F3145E4E6CB66F69508569D96E" box="[287,311,1345,1371]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
, pinned, with partially spread wings, slightly worn, four labels: // ESPANA, Prov.
<collectingRegion id="4984E698145E4E6CB27469518128D96F" box="[1284,1398,1344,1370]" country="Spain" name="Cantabria" pageId="17" pageNumber="18">Cantabria</collectingRegion>
<collectorName id="26B54DAC145E4E6CB7CD69758676D948" box="[189,552,1379,1406]" pageId="17" pageNumber="18">PN Picos de Europa Espinama</collectorName>
,
<location id="8E9F7EA1145E4E6CB5446972876DD948" LSID="urn:lsid:plazi:treatment:03E9996C145E4E6EB53E6EB48594DEDF:8E9F7EA1145E4E6CB5446972876DD948" box="[564,819,1379,1405]" country="Spain" latitude="43.148666" longLatPrecision="11" longitude="-4.7755" municipality="Picos de Europa Espinama" name="Portillas de Poqueion" pageId="17" pageNumber="18" stateProvince="Cantabria">Portillas de Poqueion</location>
,
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<elevation id="006DCF49145E4E6CB44F697287C6D94B" box="[831,920,1379,1406]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="17" pageNumber="18" unit="m" value="1340.0">1340 m</elevation>
</quantity>
<geoCoordinate id="EE744EBD145E4E6CB4D069708078D94B" box="[928,1062,1377,1406]" degrees="4" direction="west" minutes="46,53" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="-4.7755">4°46,53′ W</geoCoordinate>
,
<geoCoordinate id="EE744EBD145E4E6CB3426970809ED94B" box="[1074,1216,1377,1406]" degrees="43" direction="north" minutes="08,92" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="43.148666">43°08,92′ N</geoCoordinate>
<date id="FFFE0EBA145E4E6CB3B869728161D948" box="[1224,1343,1379,1405]" pageId="17" pageNumber="18" value="2012-11-07">
<collectingDate id="EFBAF752145E4E6CB3B869728161D948" box="[1224,1343,1379,1405]" pageId="17" pageNumber="18" value="2012-11-07">11.7.2012</collectingDate>
</date>
, leg.
<collectorName id="26B54DAC145E4E6CB7CD69968545D994" box="[189,283,1415,1441]" pageId="17" pageNumber="18">Huemer</collectorName>
TLMF 2012-011 // BC TLMF
<collectorName id="26B54DAC145E4E6CB5E1699686E1D994" box="[657,703,1415,1441]" pageId="17" pageNumber="18">Lep</collectorName>
08335 [green label] //
<collectorName id="26B54DAC145E4E6CB4A169968057D994" box="[977,1033,1415,1441]" pageId="17" pageNumber="18">Gen.</collectorName>
prep.
<specimenCount id="9D46E3F3145E4E6CB32569968035D994" box="[1109,1131,1415,1441]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
1/
<date id="FFFE0EBA145E4E6CB3F6699680A1D994" box="[1158,1279,1415,1441]" pageId="17" pageNumber="18" value="2012-11-07">
<collectingDate id="EFBAF752145E4E6CB3F6699680A1D994" box="[1158,1279,1415,1441]" pageId="17" pageNumber="18" value="2012-11-07">11.7.2012</collectingDate>
</date>
[genitalia slide number] //
<typeStatus id="54FB96D8145E4E6CB60B69BB8648D9F1" box="[379,534,1450,1476]" pageId="17" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C4053F9145E4E6CB56C69BB802CD9F1" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[540,1138,1450,1476]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cantabriana">
<emphasis id="B934F468145E4E6CB56C69BB8719D9F1" box="[540,839,1450,1476]" italics="true" pageId="17" pageNumber="18">Phtheochroa cantabriana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF145E4E6CB26669BB8135D9F1" box="[1302,1387,1450,1476]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="17" pageNumber="18">TLMF</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BFF287A145E4E6CB79A69E28539DA38" blockId="17.[234,359,1523,1549]" box="[234,359,1523,1549]" pageId="17" pageNumber="18">
<emphasis id="B934F468145E4E6CB79A69E28539DA38" bold="true" box="[234,359,1523,1549]" pageId="17" pageNumber="18">
<typeStatus id="54FB96D8145E4E6CB79A69E28539DA38" box="[234,359,1523,1549]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFF287A145E4E6CB7CD6A348573DBF7" blockId="17.[189,1399,1573,1986]" pageId="17" pageNumber="18">
<collectingCountry id="F35768EA145E4E6CB7CD6A34854ADA0A" box="[189,276,1573,1599]" name="Spain" pageId="17" pageNumber="18">SPAIN</collectingCountry>
:
<specimenCount id="9D46E3F3145E4E6CB66D6A348514DA75" box="[285,330,1573,1600]" pageId="17" pageNumber="18" type="male">1 ♂</specimenCount>
, pinned, with partially spread wings, slightly worn, four labels: //
<collectingCountry id="F35768EA145E4E6CB3346A3480E9DA0A" box="[1092,1207,1573,1599]" name="Spain" pageId="17" pageNumber="18">ESPANA</collectingCountry>
, Prov.
<collectingRegion id="4984E698145E4E6CB2746A348128DA0A" box="[1284,1398,1573,1599]" country="Spain" name="Cantabria" pageId="17" pageNumber="18">Cantabria</collectingRegion>
PN Picos de Europa Espinama, Portillas de Poqueion,
<quantity id="4CB8859F145E4E6CB44F6A5987C6DA56" box="[831,920,1608,1635]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="17" pageNumber="18" unit="m" value="1340.0">1340 m</quantity>
<geoCoordinate id="EE744EBD145E4E6CB4D06A578078DA56" box="[928,1062,1606,1635]" degrees="4" direction="west" minutes="46,53" orientation="longitude" pageId="17" pageNumber="18" precision="9" value="-4.7755">4°46,53′ W</geoCoordinate>
,
<geoCoordinate id="EE744EBD145E4E6CB3426A57809EDA56" box="[1074,1216,1606,1635]" degrees="43" direction="north" minutes="08,92" orientation="latitude" pageId="17" pageNumber="18" precision="9" value="43.148666">43°08,92′ N</geoCoordinate>
<date id="FFFE0EBA145E4E6CB3B86A598161DA57" box="[1224,1343,1608,1634]" pageId="17" pageNumber="18" value="2012-11-07">11.7.2012</date>
, leg. Huemer TLMF 2012-011 // BC TLMF Lep 08336 [green label] // Gen. prep.
<specimenCount id="9D46E3F3145E4E6CB3256A7D8035DAB3" box="[1109,1131,1644,1670]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
2/
<date id="FFFE0EBA145E4E6CB3FA6A7D80A1DAB3" box="[1162,1279,1644,1670]" pageId="17" pageNumber="18" value="2012-11-07">11.7.2012</date>
[genitalia slide number] //
<typeStatus id="54FB96D8145E4E6CB60D6A9E8651DA9C" box="[381,527,1679,1705]" pageId="17" pageNumber="18" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C4053F9145E4E6CB5656A9E8031DA9C" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[533,1135,1679,1705]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cantabriana">
<emphasis id="B934F468145E4E6CB5656A9E871FDA9C" box="[533,833,1679,1705]" italics="true" pageId="17" pageNumber="18">Phtheochroa cantabriana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF145E4E6CB2656A9E8134DA9C" box="[1301,1386,1679,1705]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="17" pageNumber="18">TLMF</collectionCode>
);
<specimenCount id="9D46E3F3145E4E6CB7CD6AA384B5DAF9" box="[189,235,1714,1740]" pageId="17" pageNumber="18" type="male">1 ♂</specimenCount>
, pinned, with partially spread wings, slightly worn, three labels: // locality data label as preceding // BC TLMF Lep 08334 [green label] // Gen. prep.
<specimenCount id="9D46E3F3145E4E6CB58D6AC7874DDAC5" box="[765,787,1750,1776]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
3/
<date id="FFFE0EBA145E4E6CB4406AC487F8DADA" box="[816,934,1749,1775]" pageId="17" pageNumber="18" value="2012-11-07">11.7.2012</date>
[genitalia slide number] //
<typeStatus id="54FB96D8145E4E6CB3956AC48129DADA" box="[1253,1399,1749,1775]" pageId="17" pageNumber="18" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C4053F9145E4E6CB7CD6AE98778DB27" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[189,806,1784,1811]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cantabriana">
<emphasis id="B934F468145E4E6CB7CD6AE985B2DB27" box="[189,492,1784,1810]" italics="true" pageId="17" pageNumber="18">Phtheochroa cantabriana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF145E4E6CB4A86AE98073DB27" box="[984,1069,1784,1810]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="17" pageNumber="18">TLMF</collectionCode>
);
<specimenCount id="9D46E3F3145E4E6CB3336AE9802ADB26" box="[1091,1140,1784,1811]" pageId="17" pageNumber="18" type="male">1 ♂</specimenCount>
, well-preserved, with spread wings, five labels: //
<collectingCountry id="F35768EA145E4E6CB5796B0A8611DB00" box="[521,591,1819,1845]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
,
<collectingRegion id="4984E698145E4E6CB52B6B0A8693DB00" box="[603,717,1819,1845]" country="Spain" name="Cantabria" pageId="17" pageNumber="18">Cantabria</collectingRegion>
Puerto San Glorio Picos [de] Europa,
<quantity id="4CB8859F145E4E6CB3E76B0A80AFDB03" box="[1175,1265,1819,1846]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="17" pageNumber="18" unit="m" value="1610.0">1610 m</quantity>
<date id="FFFE0EBA145E4E6CB3896B0A8128DB00" box="[1273,1398,1819,1845]" pageId="17" pageNumber="18" value="1988-06-10">10.vi.1988</date>
P. Stadel Nielsen [hand written] // ZMUC
<taxonomicName id="4C4053F9145E4E6CB5C26B2E8059DB6C" authority="Spanish Revision" authorityName="Spanish Revision" box="[690,1031,1855,1881]" class="Insecta" family="Tortricidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Tortricidae Spanish Revision</taxonomicName>
specimen # 1341 [green label] // Coll. ZMUC, Copenhagen
<collectingCountry id="F35768EA145E4E6CB56E6B7386D5DB49" box="[542,651,1890,1916]" name="Denmark" pageId="17" pageNumber="18">Denmark</collectingCountry>
[yellow label] // DK Copenhagen Zool. Museum
<specimenCount id="9D46E3F3145E4E6CB39B6B73815FDB49" box="[1259,1281,1890,1916]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
Genitalia slide No. 1/
<date id="FFFE0EBA145E4E6CB6336B9485E3DBAA" box="[323,445,1925,1951]" pageId="17" pageNumber="18" value="1988-10-06">10.6.1988</date>
//
<typeStatus id="54FB96D8145E4E6CB6AE6B94862EDBAA" box="[478,624,1925,1951]" pageId="17" pageNumber="18" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C4053F9145E4E6CB5086B948085DBAA" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[632,1243,1925,1952]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cantabriana">
<emphasis id="B934F468145E4E6CB5086B9487F8DBAA" box="[632,934,1925,1951]" italics="true" pageId="17" pageNumber="18">Phtheochroa cantabriana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF145E4E6CB7B56BB9857FDBF7" box="[197,289,1960,1986]" country="Denmark" name="Zoological Museum, University of Copenhagen" pageId="17" pageNumber="18">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1145D4E6EB7CD6D1A8007DDD3" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8BFF287A145D4E6FB7CD6D1A8510DD10" blockId="18.[189,334,267,293]" box="[189,334,267,293]" pageId="18" pageNumber="19">
<heading id="D0B79F16145D4E6FB7CD6D1A8510DD10" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B934F468145D4E6FB7CD6D1A8510DD10" bold="true" box="[189,334,267,293]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A145D4E6FB79A6D2B85F3DD61" blockId="18.[234,429,313,340]" box="[234,429,313,340]" pageId="18" pageNumber="19">
<emphasis id="B934F468145D4E6FB79A6D2B8574DD61" bold="true" box="[234,298,314,340]" pageId="18" pageNumber="19">Male</emphasis>
(
<figureCitation id="137B34FF145D4E6FB64A6D288525DD61" box="[314,379,313,340]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
I–J)
</paragraph>
<paragraph id="8BFF287A145D4E6FB7CD6D79850DDDFD" blockId="18.[189,1398,360,456]" pageId="18" pageNumber="19">HEAD. Frons beige, vertex creamy, labial palps long, pointed anterad, white dorsally, rust laterally and brown ventrally. Antennae with scapus and pedicellus dorsally rust, ventrally white, flagellum with white scales.</paragraph>
<paragraph id="8BFF287A145D4E6FB7CD6DE18658DE8F" blockId="18.[189,1399,496,698]" pageId="18" pageNumber="19">
THORAX. Dorsally with white and grey scales, tegula the same but laterally rust, ventrally whitish, fore- and middle legs brownish, hind legs whitish. Forewing long and narrow with pointed apex, length
<quantity id="4CB8859F145D4E6FB7CD6E278503DE64" box="[189,349,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.4" metricValueMin="8.8" pageId="18" pageNumber="19" unit="mm" value="9.6" valueMax="10.4" valueMin="8.8">8.8–10.4 mm</quantity>
(x=
<quantity id="4CB8859F145D4E6FB6E96E2785A4DE64" box="[409,506,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="18" pageNumber="19" unit="mm" value="9.6">9.6 mm</quantity>
, n =4), upperside pale yellow with rust base, sometimes with rust dorsal blotch and subterminal fascia, cilia white, underside pale grey, with white scales in costal and anal areas. Hindwing upperside pale grey, cilia white with pale grey basal part, underside white with pale grey scales along veins R and M.
</paragraph>
<paragraph id="8BFF287A145D4E6FB7CD6EF08524DECE" blockId="18.[189,378,737,764]" box="[189,378,737,764]" pageId="18" pageNumber="19">ABDOMEN. Grey.</paragraph>
<paragraph id="8BFF287A145D4E6FB7CD6F32803CD8F5" blockId="18.[189,1399,803,1216]" pageId="18" pageNumber="19">
GENITALIA (
<figureCitation id="137B34FF145D4E6FB6336F3285C1DF08" box="[323,415,803,829]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="18" pageNumber="19">Fig. 2F</figureCitation>
). Curvature of valva weak, rest of valva, uncus and socii very similar to those of
<taxonomicName id="4C4053F9145D4E6FB7CD6F56853DDF55" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[189,355,838,864]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F468145D4E6FB7CD6F56853DDF55" box="[189,355,838,864]" italics="true" pageId="18" pageNumber="19">P. schawerdae</emphasis>
</taxonomicName>
. Transtilla (
<figureCitation id="137B34FF145D4E6FB69C6F57861FDF55" box="[492,577,838,864]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="18" pageNumber="19">Fig. 3F</figureCitation>
) large, rectangular. Phallus (
<figureCitation id="137B34FF145D4E6FB4F96F578792DF55" box="[905,972,838,864]" captionStart="Fig" captionStartId="14.[189,232,1893,1919]" captionTargetBox="[250,1346,269,1849]" captionTargetId="figure@14.[238,1349,266,1851]" captionTargetPageId="14" captionText="Fig. 7. Phalli with vesica everted of Phtheochroa spp. A–B. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – C–D. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
C–D) curved ventrally, medioventral process slender, pointed, with apex slightly curved to the right. Vesica in general resembles that of
<taxonomicName id="4C4053F9145D4E6FB7CD6F9C853ADF93" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[189,356,908,934]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F468145D4E6FB7CD6F9C853ADF93" box="[189,356,908,934]" italics="true" pageId="18" pageNumber="19">P. schawerdae</emphasis>
</taxonomicName>
, with large main part protruded at right from sclerotized phallus and two large diverticula bearing long, robust cornuti. Small, conical undulating structure (diverticulum) emerges ventrally from main part. Gonopore located dorsally, surrounded by semicylindrical sclerotized plate with minute tubercles. Main part medially constricted by circular furrow, in which gonopore is sunk. Main part of vesica looks angular in lateral left view. Diverticula emerge close to each other from right side, right one straight, emerging from dorsal half of right part and pointed posterolaterally, left one curved and oriented laterally. In dorsal view, axes of right diverticulum and sclerotized phallus usually form angle of 120–140°. Cornuti aciculate, capitate, of equal length, left one slightly bent. Barely discernible acanthae present on left portion of main part of vesica and at end of diverticula.
</paragraph>
<caption id="DF3F78F2145D4E6FB7CD6B9B80CBDBF2" ID-DOI="http://doi.org/10.5281/zenodo.3838430" ID-Zenodo-Dep="3838430" httpUri="https://zenodo.org/record/3838430/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1930,1956]" targetBox="[190,1397,1281,1884]" targetPageId="18">
<paragraph id="8BFF287A145D4E6FB7CD6B9B80CBDBF2" blockId="18.[189,1399,1930,1992]" pageId="18" pageNumber="19">
<emphasis id="B934F468145D4E6FB7CD6B9B8554DB91" bold="true" box="[189,266,1930,1956]" pageId="18" pageNumber="19">Fig. 9.</emphasis>
Maximum likelihood tree (built with MEGA6) of cytochrome
<emphasis id="B934F468145D4E6FB49A6B9A87A9DB91" box="[1002,1015,1931,1956]" italics="true" pageId="18" pageNumber="19">c</emphasis>
oxidase subunit I (COI) barcode fragments. Values at the nodes are bootstrap support values based on 500 replicates.
</paragraph>
</caption>
<paragraph id="8BFF287A145C4E6EB79A6D1A851BDD10" blockId="19.[234,325,267,293]" box="[234,325,267,293]" pageId="19" pageNumber="20">
<emphasis id="B934F468145C4E6EB79A6D1A851BDD10" bold="true" box="[234,325,267,293]" pageId="19" pageNumber="20">Female</emphasis>
</paragraph>
<paragraph id="8BFF287A145C4E6EB7CD6D288567DD66" blockId="19.[189,313,313,339]" box="[189,313,313,339]" pageId="19" pageNumber="20">Unknown.</paragraph>
<paragraph id="8BFF287A145C4E6EB7CD6D6A861FDDA0" blockId="19.[189,577,379,405]" box="[189,577,379,405]" pageId="19" pageNumber="20">
<heading id="D0B79F16145C4E6EB7CD6D6A8525DDA0" box="[189,379,379,405]" pageId="19" pageNumber="20">
<emphasis id="B934F468145C4E6EB7CD6D6A8525DDA0" bold="true" box="[189,379,379,405]" pageId="19" pageNumber="20">Molecular data</emphasis>
</heading>
(
<tableCitation id="C6C21DC1145C4E6EB6FB6D6A85BADDA0" box="[395,484,379,405]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="19" pageNumber="20" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
,
<figureCitation id="137B34FF145C4E6EB6826D6A8669DDA0" box="[498,567,379,405]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[190,1397,1281,1884]" captionTargetId="figure@18.[172,1412,1251,1902]" captionTargetPageId="18" captionText="Fig. 9. Maximum likelihood tree (built with MEGA6) of cytochrome c oxidase subunit I (COI) barcode fragments. Values at the nodes are bootstrap support values based on 500 replicates." figureDoi="http://doi.org/10.5281/zenodo.3838430" httpUri="https://zenodo.org/record/3838430/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A145C4E6EB7CD6DBB8007DDD3" blockId="19.[189,1398,425,487]" pageId="19" pageNumber="20">
BIN URI: BOLD:ACA9597. The maximum intraspecific divergence of the barcode region is 0.31% (n=3). The minimum distance to the nearest neighbour,
<taxonomicName id="4C4053F9145C4E6EB4226DDC8782DDD2" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[850,988,461,487]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F468145C4E6EB4226DDC8782DDD2" box="[850,988,461,487]" italics="true" pageId="19" pageNumber="20">P. frigidana</emphasis>
</taxonomicName>
, is 3.69%.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1145C4E6EB7CD6E1F8692DE4F" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="8BFF287A145C4E6EB7CD6E1F857EDE1D" blockId="19.[189,288,526,552]" box="[189,288,526,552]" pageId="19" pageNumber="20">
<heading id="D0B79F16145C4E6EB7CD6E1F857EDE1D" bold="true" box="[189,288,526,552]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B934F468145C4E6EB7CD6E1F857EDE1D" bold="true" box="[189,288,526,552]" pageId="19" pageNumber="20">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A145C4E6EB7CD6E2D8692DE4F" blockId="19.[189,1399,572,634]" pageId="19" pageNumber="20">
Preimaginal stages and larval host plant are unknown. The moths fly in June–July. Habitat: the
<typeStatus id="54FB96D8145C4E6EB2346E2C8129DE62" box="[1348,1399,573,599]" pageId="19" pageNumber="20">type</typeStatus>
material was collected in montane meadows.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1145C4E6EB7CD6EB08594DEDF" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BFF287A145C4E6EB7CD6EB0859ADE8E" blockId="19.[189,452,673,699]" box="[189,452,673,699]" pageId="19" pageNumber="20">
<heading id="D0B79F16145C4E6EB7CD6EB08508DE8E" box="[189,342,673,699]" pageId="19" pageNumber="20">
<emphasis id="B934F468145C4E6EB7CD6EB08508DE8E" bold="true" box="[189,342,673,699]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
(
<figureCitation id="137B34FF145C4E6EB6166EB085E5DE8E" box="[358,443,673,699]" captionStart="Fig" captionStartId="20.[189,232,1990,2016]" captionTargetBox="[189,1398,1296,1948]" captionTargetId="figure@20.[183,1404,1290,1954]" captionTargetPageId="20" captionText="Fig. 10. Distribution of the Phtheochroa frigidana s. lat. species complex based on examined material." figureDoi="http://doi.org/10.5281/zenodo.3838432" httpUri="https://zenodo.org/record/3838432/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A145C4E6EB7CD6EC18594DEDF" blockId="19.[189,458,720,746]" box="[189,458,720,746]" pageId="19" pageNumber="20">Cantabrian Mountains.</paragraph>
</subSubSection>
</treatment>
</document>
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