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<document id="8B8C12EF318309791E90B684EAEBB173" ID-DOI="10.5281/zenodo.178756" ID-GBIF-Dataset="f9ea8775-9b2d-4364-b20a-4738aab6a54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="178756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920987951" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2007" docId="03E987D1FFA4FFE6FF178C173132FCE1" docLanguage="en" docName="zt01604p020.pdf" docOrigin="Zootaxa 1604" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpalus potanini Tschitscherine 1906" docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FFD0FFA9FFB5FFF4FF8088153517FFD3" masterDocTitle="Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="18" updateTime="1698229235843" updateUser="plazi">
<mods:mods id="1ECF202018AEA622C3DEFA0497F179C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BAE1EC0CC26774ECE3ECD8B83B9D1116">
<mods:title id="83057165C40808F1BBFE1D1F541EF254">Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="CEE059E3338F150128DBFE8E694A3018">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="9651913F3A48F4C5EEB232311368B3CA">Liang, Hongbin</mods:namePart>
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<mods:title id="BFBC9A9B48DE97E67030E824179DCA0A">Zootaxa</mods:title>
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<mods:date id="B2A522767EA9C74131F098F53E12A22E">2007</mods:date>
<mods:detail id="B649219389B608C752D16EF0C8D9F9B1" type="volume">
<mods:number id="6B04C375FB22F65597120781B9A0419E">1604</mods:number>
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<mods:classification id="602B9737792C2B30525E0914C5821275">journal article</mods:classification>
<mods:identifier id="A43C9836E63E954E0404ADA3A8DB664F" type="DOI">10.5281/zenodo.178756</mods:identifier>
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<mods:identifier id="2892F4D4F5F298F97DF77D6815C9B7AE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D2E783C8B590A89EB99E130B467CADA9" type="Zenodo-Dep">178756</mods:identifier>
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<treatment id="03E987D1FFA4FFE6FF178C173132FCE1" ID-DOI="http://doi.org/10.5281/zenodo.6250252" ID-GBIF-Taxon="119373725" ID-Zenodo-Dep="6250252" LSID="urn:lsid:plazi:treatment:03E987D1FFA4FFE6FF178C173132FCE1" httpUri="http://treatment.plazi.org/id/03E987D1FFA4FFE6FF178C173132FCE1" lastPageId="18" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="C35A654CFFA4FFE5FF178C17343AFBEC" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BFF36C7FFA4FFE5FF178C17376EFBCF" blockId="17.[151,633,1026,1087]" box="[151,633,1026,1052]" pageId="17" pageNumber="18">
<heading id="D0B781ABFFA4FFE5FF178C17376EFBCF" bold="true" box="[151,633,1026,1052]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="4C404D44FFA4FFE5FF178C17376EFBCF" ID-CoL="3JSPS" authority="Tschitscherine, 1906" authorityName="Tschitscherine" authorityYear="1906" box="[151,633,1026,1052]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FF178C17376EFBCF" bold="true" box="[151,633,1026,1052]" pageId="17" pageNumber="18">
<emphasis id="B934EAD5FFA4FFE5FF178C173462FBCF" bold="true" box="[151,373,1026,1052]" italics="true" pageId="17" pageNumber="18">Harpalus potanini</emphasis>
<bibRefCitation id="EFD14B36FFA4FFE5FEFC8C17376EFBCF" author="Tschitscherine" box="[380,633,1026,1052]" pageId="17" pageNumber="20" refString="Tschitscherine, T. (1906) Notes detachees sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37 (3 / 4), 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFF36C7FFA4FFE5FF178C30343AFBEC" blockId="17.[151,633,1026,1087]" box="[151,301,1061,1087]" pageId="17" pageNumber="18">
(
<figureCitation id="137B2A42FFA4FFE5FF1F8C303432FBEC" box="[159,293,1061,1087]" captionStart="FIGURES 54 – 55" captionStartId="18.[151,269,1954,1978]" captionTargetBox="[420,1148,871,1953]" captionTargetId="figure@18.[384,1202,838,1958]" captionTargetPageId="18" captionText="FIGURES 54 – 55. Harpalus potanini (Sichuan, Xiaojin Valley, lectotype), median lobe of aedeagus. 54, lateral aspect; 55, dorsal aspect. Scale = 1.0 mm." httpUri="https://zenodo.org/record/178764/files/figure.png" pageId="17" pageNumber="18">Figs 54–55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFA4FFE5FF178C60300BFB5F" box="[151,1308,1141,1164]" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BFF36C7FFA4FFE5FF178C60300BFB5F" blockId="17.[151,1308,1141,1164]" box="[151,1308,1141,1164]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB5011E9FFA4FFE5FF178C60300BFB5F" box="[151,1308,1141,1164]" pageId="17" pageNumber="18">
<taxonomicName id="4C404D44FFA4FFE5FF178C603760FB5F" ID-CoL="3JSPS" authority="Tschitscherine, 1906: 255" authorityName="Tschitscherine" authorityPageNumber="255" authorityYear="1906" box="[151,631,1141,1164]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FF178C60344CFB5F" box="[151,347,1141,1164]" italics="true" pageId="17" pageNumber="18">Harpalus potanini</emphasis>
<treatmentCitation id="0AE110D6FFA4FFE5FEE28C603760FB5F" author="Tschitscherine" box="[354,631,1141,1164]" page="255" pageId="17" pageNumber="18" year="1906">
<bibRefCitation id="EFD14B36FFA4FFE5FEE28C603755FB5F" author="Tschitscherine" box="[354,578,1141,1164]" pageId="17" pageNumber="20" refString="Tschitscherine, T. (1906) Notes detachees sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37 (3 / 4), 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
: 255
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54FB8865FFA4FFE5FD018C6037A1FB5F" box="[641,694,1141,1164]" pageId="17" pageNumber="18">Type</typeStatus>
locality: Xiaojin Valley, Sichuan,
<collectingCountry id="F3577657FFA4FFE5FBA58C603172FB5F" box="[1061,1125,1141,1164]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
(here restricted).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFA4FFE5FF178CA83110FA14" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BFF36C7FFA4FFE5FF178CA83691FA9C" blockId="17.[151,1437,1212,1999]" pageId="17" pageNumber="18">
<emphasis id="B934EAD5FFA4FFE5FF178CA83453FB04" bold="true" box="[151,324,1213,1239]" pageId="17" pageNumber="18">
<typeStatus id="54FB8865FFA4FFE5FF178CA835C5FB04" box="[151,210,1213,1239]" pageId="17" pageNumber="18">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54FB8865FFA4FFE5FED48CA834DEFB04" box="[340,457,1213,1239]" pageId="17" pageNumber="18" type="lectotype">Lectotype</typeStatus>
(present designation): ɗ, labelled &quot;Syao-Chzhin-kho (= Xiaojin) Valley,
<date id="FFFE1007FFA4FFE5FAA28CA83081FB04" box="[1314,1430,1213,1239]" pageId="17" pageNumber="18" value="1993-07-27">27.VII.93</date>
, Potanin&quot; [in Cyrillic], &quot;
<taxonomicName id="4C404D44FFA4FFE5FE3E8CF03734FB2D" box="[446,547,1253,1278]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="19" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FE3E8CF03734FB2D" box="[446,547,1253,1278]" italics="true" pageId="17" pageNumber="18">Potanini</emphasis>
</taxonomicName>
m., typ, Tschtscherin det.&quot;, “
<typeStatus id="54FB8865FFA4FFE5FC188CF03137FB2C" box="[920,1056,1253,1279]" pageId="17" pageNumber="18" type="lectotype">Lectotypus</typeStatus>
,
<taxonomicName id="4C404D44FFA4FFE5FBB28CF035FAFAF4" authority="Tschit., Z. Mlynar" authorityName="Tschit., Z. Mlynar" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FBB28CF03007FB2D" box="[1074,1296,1253,1278]" italics="true" pageId="17" pageNumber="18">Harpalus potanini</emphasis>
Tschit., Z. Mlynář
</taxonomicName>
det. 1976&quot; (
<collectionCode id="ED51AE02FFA4FFE5FE068D1834DFFAF4" box="[390,456,1293,1319]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
).
<typeStatus id="54FB8865FFA4FFE5FE5F8D183798FAF4" box="[479,655,1293,1319]" pageId="17" pageNumber="18" type="paralectotype">Paralectotypes</typeStatus>
: 3 ɗ, 2 Ψ, same data as
<typeStatus id="54FB8865FFA4FFE5FC3D8D18313FFAF4" box="[957,1064,1293,1319]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="ED51AE02FFA4FFE5FBBC8D183169FAF4" box="[1084,1150,1293,1319]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
); 5 Ψ, &quot;Fu-byan-kho (= Fubian) Valley,
<date id="FFFE1007FFA4FFE5FED18D203733FA9C" box="[337,548,1333,1359]" pageId="17" pageNumber="18" value="1993-07-31" valueMax="1993-08-01" valueMin="1993-07-31">31.VII.–1.VIII.93</date>
, Potanin&quot; [in Cyrillic] (
<collectionCode id="ED51AE02FFA4FFE5FCB88D20366DFA9C" box="[824,890,1333,1359]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
).
</paragraph>
<paragraph id="8BFF36C7FFA4FFE5FF478D483110FA14" blockId="17.[151,1437,1212,1999]" pageId="17" pageNumber="18">
<emphasis id="B934EAD5FFA4FFE5FF478D483754FAA4" bold="true" box="[199,579,1373,1399]" pageId="17" pageNumber="18">Additional material examined</emphasis>
.
<collectingCountry id="F3577657FFA4FFE5FDD48D483789FAA4" box="[596,670,1373,1399]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
. SICHUAN: 1 ɗ, Luding Co.,
<date id="FFFE1007FFA4FFE5FB978D4831A5FAA4" box="[1047,1202,1373,1399]" pageId="17" pageNumber="18" value="1981-08-12">12.VIII.1981</date>
, Shang Jinwen leg. (
<collectionCode id="ED51AE02FFA4FFE5FF208D9035C3FA4C" box="[160,212,1413,1439]" pageId="17" pageNumber="18">IZB</collectionCode>
); 1 ɗ, Kangding Co.,
<quantity id="4CB89B22FFA4FFE5FE5B8D903721FA4C" box="[475,566,1413,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="18" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FFFE1007FFA4FFE5FDC48D9037F7FA4C" box="[580,736,1413,1439]" pageId="17" pageNumber="18" value="1981-08-18">18.VIII.1981</date>
, Shang Jinwen leg. (
<collectionCode id="ED51AE02FFA4FFE5FC578D90311CFA4C" box="[983,1035,1413,1439]" pageId="17" pageNumber="18">IZB</collectionCode>
); 2 ɗ, 1 Ψ, same data but
<quantity id="4CB89B22FFA4FFE5FAC28D90308BFA4C" box="[1346,1436,1413,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="m" value="1500.0">1500 m</quantity>
(
<collectionCode id="ED51AE02FFA4FFE5FF208DB835C3FA14" box="[160,212,1453,1479]" pageId="17" pageNumber="18">IZB</collectionCode>
); 1 ɗ, Kangding Co.,
<quantity id="4CB89B22FFA4FFE5FE548DB83727FA14" box="[468,560,1453,1479]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="17" pageNumber="18" unit="m" value="1980.0">1980 m</quantity>
,
<date id="FFFE1007FFA4FFE5FDBC8DB837D3FA14" box="[572,708,1453,1479]" pageId="17" pageNumber="18" value="1989-06-22">22.VI.1989</date>
, Wang Shuyong leg. (
<collectionCode id="ED51AE02FFA4FFE5FC458DB836EDFA14" box="[965,1018,1453,1479]" pageId="17" pageNumber="18">IZB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFA4FFE5FF478DC036DAF9EC" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BFF36C7FFA4FFE5FF478DC036DAF9EC" blockId="17.[151,1437,1212,1999]" pageId="17" pageNumber="18">
<emphasis id="B934EAD5FFA4FFE5FF478DC03477FA3C" bold="true" box="[199,352,1493,1519]" pageId="17" pageNumber="18">Distribution</emphasis>
. This species has originally been described from the series collected in two localities of the Central Sichuan, both to north of Kangding (Xiaojin Valley and Fubian Valley). According to the new examined material,
<taxonomicName id="4C404D44FFA4FFE5FEBF8E3034D2F9ED" box="[319,453,1573,1598]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FEBF8E3034D2F9ED" box="[319,453,1573,1598]" italics="true" pageId="17" pageNumber="18">H. potanini</emphasis>
</taxonomicName>
seems to be endemic to the central Sichuan.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFA4FFE6FF478E583132FCE1" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BFF36C7FFA4FFE6FF478E583132FCE1" blockId="17.[151,1437,1212,1999]" lastBlockId="18.[151,1437,152,818]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B934EAD5FFA4FFE5FF478E58342EF9B4" bold="true" box="[199,313,1613,1639]" pageId="17" pageNumber="18">Remarks</emphasis>
.
<taxonomicName id="4C404D44FFA4FFE5FEC78E5834DAF9B5" box="[327,461,1613,1638]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FEC78E5834DAF9B5" box="[327,461,1613,1638]" italics="true" pageId="17" pageNumber="18">H. potanini</emphasis>
</taxonomicName>
is very remarkable species and due to the rather flat body with mat dorsum, widely rounded basal angles of pronotum and rather slender legs slightly similar in appearance to some species of the genus
<taxonomicName id="4C404D44FFA4FFE5FF638E8834BCF964" authority="Bonelli" authorityName="Bonelli" box="[227,427,1693,1719]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAD5FFA4FFE5FF638E88345AF965" box="[227,333,1693,1718]" italics="true" pageId="17" pageNumber="18">Calathus</emphasis>
Bonelli
</taxonomicName>
belonging to the
<emphasis id="B934EAD5FFA4FFE5FDFB8E8837CEF965" box="[635,729,1693,1718]" italics="true" pageId="17" pageNumber="18">fuscipes</emphasis>
group. The species was described without comparison with any known species and we did not find additional information about its taxonomy or geographical distribution published since the original description. In our opinion,
<taxonomicName id="4C404D44FFA4FFE5FCA88EF836B9F8D5" box="[808,942,1773,1798]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA4FFE5FCA88EF836B9F8D5" box="[808,942,1773,1798]" italics="true" pageId="17" pageNumber="18">H. potanini</emphasis>
</taxonomicName>
is indeed dissimilar to any known species of the genus and seems to represent a separate species–group within
<taxonomicName id="4C404D44FFA4FFE5FC448F003124F8FD" box="[964,1075,1813,1838]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B934EAD5FFA4FFE5FC448F003124F8FD" box="[964,1075,1813,1838]" italics="true" pageId="17" pageNumber="18">Harpalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757AEFFA4FFE5FBBB8F00316EF8FC" box="[1083,1145,1813,1839]" pageId="17" pageNumber="18" sensu="stricto">s. str.</taxonomicNameLabel>
with the following most important distinctive character states: the body length 10.8–14.0 mm; the coloration dark, without metallic tinge on dorsum; the head, pronotum, elytra, abdominal sternites and tarsi dorsally glabrous; the pronotal basal margin not ciliate; the elytral basal border glabrous; the elytral preapical sinuation rather deep, with traces of obtuse denticle at its base; the elytral third interval at most with one discal pore; the wings completely reduced; the metepisterna wider than long; the legs comparatively long and slender, with long and narrow, almost parallel–sided metatarsomeres; the protibia with one ventroapical spine and three preapical spines on outer margin isolated from spines on ventral surface; the median lobe of aedeagus with rather long terminal lamella, with oblique horseshoe–shaped apical capitulum and distal orifice shifted to left; the internal sac of aedeagus with two large separate teeth, with a small left spiny patch medially and a larger right spiny patch apically (
<figureCitation id="137B2A42FFA7FFE6FE838975349AFEA9" box="[259,397,352,378]" captionStart="FIGURES 54 – 55" captionStartId="18.[151,269,1954,1978]" captionTargetBox="[420,1148,871,1953]" captionTargetId="figure@18.[384,1202,838,1958]" captionTargetPageId="18" captionText="FIGURES 54 – 55. Harpalus potanini (Sichuan, Xiaojin Valley, lectotype), median lobe of aedeagus. 54, lateral aspect; 55, dorsal aspect. Scale = 1.0 mm." httpUri="https://zenodo.org/record/178764/files/figure.png" pageId="18" pageNumber="19">Figs 54–55</figureCitation>
). Based on the external morphology and especially the male genitalia with characteristic structure of the internal sac,
<taxonomicName id="4C404D44FFA7FFE6FE67899D377AFE72" box="[487,621,392,417]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA7FFE6FE67899D377AFE72" box="[487,621,392,417]" italics="true" pageId="18" pageNumber="19">H. potanini</emphasis>
</taxonomicName>
probably belongs to the
<emphasis id="B934EAD5FFA7FFE6FC11899D36EFFE72" box="[913,1016,392,417]" italics="true" pageId="18" pageNumber="19">honestus</emphasis>
phylogenetic stock (lineage) which includes in addition to the
<emphasis id="B934EAD5FFA7FFE6FE5989A53728FE1A" box="[473,575,432,457]" italics="true" pageId="18" pageNumber="19">honestus</emphasis>
,
<emphasis id="B934EAD5FFA7FFE6FDD089A537B4FE1A" box="[592,675,432,457]" italics="true" pageId="18" pageNumber="19">atratus</emphasis>
,
<taxonomicName id="4C404D44FFA7FFE6FD3489A53657FE1A" box="[692,832,432,457]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B934EAD5FFA7FFE6FD3489A53657FE1A" box="[692,832,432,457]" italics="true" pageId="18" pageNumber="19">aeneipennis</emphasis>
</taxonomicName>
,
<emphasis id="B934EAD5FFA7FFE6FCD089A536A7FE1A" box="[848,944,432,457]" italics="true" pageId="18" pageNumber="19">laevipes</emphasis>
and
<emphasis id="B934EAD5FFA7FFE6FC6D89A53124FE1A" box="[1005,1075,432,457]" italics="true" pageId="18" pageNumber="19">major</emphasis>
groups distributed in the East Palaearctic (Kataev in
<bibRefCitation id="EFD14B36FFA7FFE6FE2489CD37CCFE21" author="Kryzhanovskij" box="[420,731,472,498]" pageId="18" pageNumber="20" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the ground-beetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers: Sofia - Moscow, 272 pp." type="book" year="1995">Kryzhanovskij et al. 1995</bibRefCitation>
: 140) also some species groups endemic to the West Palaearctic. Within this phylogenetic stock,
<taxonomicName id="4C404D44FFA7FFE6FDD78A1537CBFDCA" box="[599,732,512,537]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA7FFE6FDD78A1537CBFDCA" box="[599,732,512,537]" italics="true" pageId="18" pageNumber="19">H. potanini</emphasis>
</taxonomicName>
seems to be related to the
<taxonomicName id="4C404D44FFA7FFE6FB8E8A15318DFDCA" box="[1038,1178,512,537]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B934EAD5FFA7FFE6FB8E8A15318DFDCA" box="[1038,1178,512,537]" italics="true" pageId="18" pageNumber="19">aeneipennis</emphasis>
</taxonomicName>
group which includes two apterous species from the West Caucasus:
<taxonomicName id="4C404D44FFA7FFE6FD4E8A3D313DFD91" authority="Faldermann" authorityName="Faldermann" box="[718,1066,552,578]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B934EAD5FFA7FFE6FD4E8A3D3697FD92" box="[718,896,552,577]" italics="true" pageId="18" pageNumber="19">H. aeneipennis</emphasis>
(Faldermann)
</taxonomicName>
and
<taxonomicName id="4C404D44FFA7FFE6FBE88A3D308CFD91" authority="Reitter. The" authorityName="Reitter. The" box="[1128,1435,552,578]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chrysopus">
<emphasis id="B934EAD5FFA7FFE6FBE88A3D3012FD92" box="[1128,1285,552,577]" italics="true" pageId="18" pageNumber="19">H. chrysopus</emphasis>
Reitter. The
</taxonomicName>
species of the
<taxonomicName id="4C404D44FFA7FFE6FEBD8A4534DEFDBA" box="[317,457,592,617]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B934EAD5FFA7FFE6FEBD8A4534DEFDBA" box="[317,457,592,617]" italics="true" pageId="18" pageNumber="19">aeneipennis</emphasis>
</taxonomicName>
group share with
<taxonomicName id="4C404D44FFA7FFE6FD1D8A453634FDBA" box="[669,803,592,617]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA7FFE6FD1D8A453634FDBA" box="[669,803,592,617]" italics="true" pageId="18" pageNumber="19">H. potanini</emphasis>
</taxonomicName>
almost all the distinctive characters of the latter listed above, differing mainly in the smaller and more convex body, the shorter legs and the presence of prominent humeral denticle on elytra. The widely separated distributions of these taxa seem to reflect their relict character. It is known that the species of the
<taxonomicName id="4C404D44FFA7FFE6FDD98ADD37F2FD32" box="[601,741,712,737]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B934EAD5FFA7FFE6FDD98ADD37F2FD32" box="[601,741,712,737]" italics="true" pageId="18" pageNumber="19">aeneipennis</emphasis>
</taxonomicName>
group live mainly in deciduous broad–leaved forest that is unusual and probably relict feature for
<taxonomicName id="4C404D44FFA7FFE6FDF98AE537FFFCDA" box="[633,744,752,777]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAD5FFA7FFE6FDF98AE537FFFCDA" box="[633,744,752,777]" italics="true" pageId="18" pageNumber="19">Harpalus</emphasis>
</taxonomicName>
most representatives of which occur on open territories. Unfortunately, there is no available information about ecology of
<taxonomicName id="4C404D44FFA7FFE6FC188B0D310AFCE2" box="[920,1053,792,817]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B934EAD5FFA7FFE6FC188B0D310AFCE2" box="[920,1053,792,817]" italics="true" pageId="18" pageNumber="19">H. potanini</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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