264 lines
47 KiB
XML
264 lines
47 KiB
XML
<document id="39692B575C449F1A1420BF1BB62A6824" ID-CLB-Dataset="8332" ID-DOI="10.11646/zootaxa.4920.3.2" ID-GBIF-Dataset="42a1a8bc-7979-4354-925e-9c8604ce1c7d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4478051" ID-ZooBank="DDB87C26-2371-4F1C-BCE5-339C5FA5B6AC" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1611899581582" checkinUser="plazi" docAuthor="Grehan, John R., Mielke, Carlos G. C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong" docDate="2021" docId="03E987879B25DA5DA39AFB669C56FC1D" docLanguage="en" docName="zootaxa.4920.3.2.pdf" docOrigin="Zootaxa 4920 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Magnificus Yan 2000" docType="treatment" docVersion="8" lastPageNumber="353" masterDocId="FFD0FFFF9B29DA53A30DFFCD987DFFF0" masterDocTitle="Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)" masterLastPageNumber="358" masterPageNumber="339" pageNumber="351" updateTime="1699980886292" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="10D556927DC713F8A87D172683D58D46">Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae)</mods:title>
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<mods:namePart id="5FE5B4C1F96802587B278EC9F4A36E98">Grehan, John R.</mods:namePart>
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<mods:affiliation id="A8F189E1A8E1C8B10114A438FB338D02">Research Associate, McGuire Center for Lepidoptera & Biodiversity, Gainesville, FL 32611, USA</mods:affiliation>
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<mods:namePart id="A15296242A40753D4C9E78ACB83AD928">Mielke, Carlos G. C.</mods:namePart>
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<mods:affiliation id="924858AB0EAF2FBB4EF23992D220D6EE">Caixa Postal 1206, 84.145 ‾ 000 Carambeí, Paraná, Brazil</mods:affiliation>
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<mods:namePart id="1F1B078EF86C29F6E095B4AB49DA69E6">Minet, Joël</mods:namePart>
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<mods:affiliation id="9EA657B4840D42A632E12E26FC0256C8">Muséum National d’Histoire Naturelle (ISYEB, UMR 7205), Entomologie, 45, rue Buffon, F- 75005 Paris, France minet @ mnhn. fr; https: // orcid. org / 0000 - 0002 - 4404 - 1635</mods:affiliation>
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<mods:namePart id="F3185513486DEB8C068C86F25E53DEAC">Ignatev, Nikolai</mods:namePart>
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<mods:affiliation id="A283CBC373D35E9CDB018944C2BEA862">Institute of Entomology, Biology Centre of the Czech Academy of Science, České Budějovice, The Czech Republic; Faculty of Sciences, University of South Bohemia, České Budějovice, Czech Republic</mods:affiliation>
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<mods:namePart id="7C0BD3722E591AD2D85775C096975709">Buchsbaum, Ulf</mods:namePart>
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<mods:affiliation id="15B3D3F975A273DB6D722CFB3140301E">Zoologische Staatssammlung M ̡ nchen, M ̡ nchhausenstrasse 21, D- 81247 M ̡ nchen, Germany</mods:affiliation>
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<mods:namePart id="66DA4FD06DF72611230D950695E22483">Xue, Dayong</mods:namePart>
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<mods:affiliation id="44C2C524AE4E10662A34868F359D6B30">Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chaoyang District, Beijing, 100101 China xuedy @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 4404 - 1635</mods:affiliation>
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<mods:title id="EA60BA72C709F38779ADFFC21C872481">Zootaxa</mods:title>
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<mods:date id="4CE87ECC7B19712D4974AE83DCE4683B">2021</mods:date>
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<mods:number id="7D61691AB05FBB60EE831648587B1F37">2021-01-29</mods:number>
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<mods:number id="7FAEF7BCB528721B6CCE87468379AE10">4920</mods:number>
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<treatment id="03E987879B25DA5DA39AFB669C56FC1D" ID-DOI="http://doi.org/10.5281/zenodo.4478069" ID-GBIF-Taxon="177316881" ID-Zenodo-Dep="4478069" LSID="urn:lsid:plazi:treatment:03E987879B25DA5DA39AFB669C56FC1D" httpUri="http://treatment.plazi.org/id/03E987879B25DA5DA39AFB669C56FC1D" lastPageId="14" lastPageNumber="353" pageId="12" pageNumber="351">
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<emphasis id="B934EA839B25DA5FA39AFB6699B4FB35" bold="true" box="[151,457,1195,1221]" pageId="12" pageNumber="351">
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Monophyly of
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<taxonomicName id="4C404D129B25DA5FA248FB6699B4FB35" authorityName="Yan" authorityYear="2000" box="[325,457,1195,1221]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">
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<emphasis id="B934EA839B25DA5FA248FB6699B4FB35" bold="true" box="[325,457,1195,1221]" italics="true" pageId="12" pageNumber="351">Magnificus</emphasis>
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</emphasis>
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<subSubSection id="C35A651A9B25DA5DA3CAFB029C56FC1D" lastPageId="14" lastPageNumber="353" pageId="12" pageNumber="351" type="description">
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<paragraph id="8BFF36919B25DA5FA3CAFB02999FF9DD" blockId="12.[151,1437,1195,1581]" pageId="12" pageNumber="351">
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Although the range FW patterns of
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<taxonomicName id="4C404D129B25DA5FA159FB029AAFFB19" authorityName="Yan" authorityYear="2000" box="[596,722,1231,1257]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">
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<emphasis id="B934EA839B25DA5FA159FB029AAFFB19" box="[596,722,1231,1257]" italics="true" pageId="12" pageNumber="351">Magnificus</emphasis>
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species are all distinct from other Eurasian genera, this contrast is not definitive as there is no unequivocal difference in wing pattern for the genus as a whole with respect to all other
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<taxonomicName id="4C404D129B25DA5FA3D5FADA9928FAC1" baseAuthorityName="Grehan" baseAuthorityYear="2012" box="[216,341,1303,1329]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
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. For example, the female FW pattern of
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<taxonomicName id="4C404D129B25DA5FA017FADA9BA9FAC1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[794,980,1303,1329]" class="Insecta" family="Hepialidae" genus="Hepialus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="humuli">
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<emphasis id="B934EA839B25DA5FA017FADA9BA9FAC1" box="[794,980,1303,1329]" italics="true" pageId="12" pageNumber="351">Hepialus humuli</emphasis>
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</taxonomicName>
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(
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<figureCitation id="137B2A149B25DA5FA0EEFADA9C58FAC1" box="[995,1061,1303,1329]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1759,1783]" captionTargetBox="[270,1318,181,1736]" captionTargetId="figure-19@3.[270,1318,181,1736]" captionTargetPageId="3" captionText="FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhidu- oensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev." figureDoi="http://doi.org/10.5281/zenodo.4478053" httpUri="https://zenodo.org/record/4478053/files/figure.png" pageId="12" pageNumber="351">Fig. 8</figureCitation>
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) is superficially similar to that of
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<taxonomicName id="4C404D129B25DA5FA39AFAF1993DFAA5" authorityName="Yan" authorityYear="2000" box="[151,320,1339,1365]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="zhiduoensis">
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<emphasis id="B934EA839B25DA5FA39AFAF1993DFAA5" box="[151,320,1339,1365]" italics="true" pageId="12" pageNumber="351">M. zhiduoensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="137B2A149B25DA5FA25DFAF699E9FAA6" box="[336,404,1339,1366]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1759,1783]" captionTargetBox="[270,1318,181,1736]" captionTargetId="figure-19@3.[270,1318,181,1736]" captionTargetPageId="3" captionText="FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhidu- oensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev." figureDoi="http://doi.org/10.5281/zenodo.4478053" httpUri="https://zenodo.org/record/4478053/files/figure.png" pageId="12" pageNumber="351">Fig. 6</figureCitation>
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) for the submarginal band, while that of
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<taxonomicName id="4C404D129B25DA5FA06FFAF69D5BFAA5" authority="(Christoph, 1888)" baseAuthorityName="Christoph" baseAuthorityYear="1888" box="[866,1318,1339,1365]" class="Insecta" family="Hepialidae" genus="Zenophassus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="schamyl">
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<emphasis id="B934EA839B25DA5FA06FFAF69C2AFAA5" box="[866,1111,1339,1365]" italics="true" pageId="12" pageNumber="351">Zenophassus schamyl</emphasis>
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(
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<bibRefCitation id="EFD14B609B25DA5FA76BFAF69D60FAA5" author="Christoph, H." box="[1126,1309,1339,1365]" pageId="12" pageNumber="351" pagination="308 - 314" refId="ref10471" refString="Christoph, H. (1888) Diagnosen zu einigen neuen Lepidoptera des palaearctischen Faunengebietes. Horae Societatuas Entomologicae Rossicae, 22, 308 - 314." type="journal article" year="1888">Christoph, 1888</bibRefCitation>
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)
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</taxonomicName>
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(
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<figureCitation id="137B2A149B25DA5FA63BFAF69D07FAA5" box="[1334,1402,1339,1365]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1759,1783]" captionTargetBox="[270,1318,181,1736]" captionTargetId="figure-19@3.[270,1318,181,1736]" captionTargetPageId="3" captionText="FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhidu- oensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev." figureDoi="http://doi.org/10.5281/zenodo.4478053" httpUri="https://zenodo.org/record/4478053/files/figure.png" pageId="12" pageNumber="351">Fig. 9</figureCitation>
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) is reminiscent of
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<taxonomicName id="4C404D129B25DA5FA24EFA929A65FA89" authorityName="Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue" authorityYear="2021" baseAuthorityName="Bang-Haas" baseAuthorityYear="1939" box="[323,536,1375,1401]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="species" species="dirschi">
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<emphasis id="B934EA839B25DA5FA24EFA929A65FA89" box="[323,536,1375,1401]" italics="true" pageId="12" pageNumber="351">Magnificus dirschi</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="137B2A149B25DA5FA124FA929A0DFA89" box="[553,624,1375,1401]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1759,1783]" captionTargetBox="[270,1318,181,1736]" captionTargetId="figure-19@3.[270,1318,181,1736]" captionTargetPageId="3" captionText="FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhidu- oensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev." figureDoi="http://doi.org/10.5281/zenodo.4478053" httpUri="https://zenodo.org/record/4478053/files/figure.png" pageId="12" pageNumber="351">Fig. 2</figureCitation>
|
||
). But two features are identified here as unambiguously supporting the monophyly of
|
||
<emphasis id="B934EA839B25DA5FA232FA4E99B9FA6D" box="[319,452,1411,1437]" italics="true" pageId="12" pageNumber="351">
|
||
<taxonomicName id="4C404D129B25DA5FA232FA4E99BDFA6D" authorityName="Yan" authorityYear="2000" box="[319,448,1411,1437]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">Magnificus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
i.e. they do not conflict with other derived character similarities within the
|
||
<taxonomicName id="4C404D129B25DA5FA62EFA4E9DE1FA6D" baseAuthorityName="Grehan" baseAuthorityYear="2012" box="[1315,1436,1411,1437]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
and appear to be unique to the genus. A third characteristic is also potentially informative pending future studies. The three features are: (i) shape of the abdominal tergosternal sclerite, (ii) the structure of the pseudoteguminal arms of the male genitalia and (iii) the shape of the lamella antevaginalis of the female genitalia. These features are individually discussed below.
|
||
</paragraph>
|
||
<paragraph id="8BFF36919B25DA5FA39AF99699F1F82D" blockId="12.[151,1437,1627,2013]" pageId="12" pageNumber="351">
|
||
<emphasis id="B934EA839B25DA5FA39AF99699BEF985" bold="true" box="[151,451,1627,1653]" pageId="12" pageNumber="351">(i) Tergosternal sclerite.</emphasis>
|
||
The anterio-abdominal tergosternal sclerite is a triple junction between the anterior lateral margins of sternite II, the junction of Tergites I and II, and the posterior edge of the metathorax (
|
||
<figureCitation id="137B2A149B25DA5FA662F9B29886F94E" captionStart="FIGURE 23" captionStartId="11.[151,250,1794,1818]" captionTargetBox="[164,1427,641,1760]" captionTargetId="figure-242@11.[151,1435,633,1770]" captionTargetPageId="11" captionText="FIGURE 23. Magnificus abdominal sclerites: (a) M. regius tergite II, (MWM), photo by Nikolai Ignatev, (b) M. bouvieri holotype SII (photo by Joël Minet), (c) Magnificus sp. sternites VII-VIII, photo by Nikolai Ignatev), (d) M. bouvieri holotype sternites VII-VIII, photo by Joël Minet), (e) M. regius sternites VII-VIII, photo by Nikolai Ignatev, (f) M. regius and (g) M. roseus, photos by Ulf Buchsbaum, (h) M. bouvieri holotype abdomen, photo by Joël Minet, (i) Magnificus sp. tergosternal sclerite, (j) Magnificus regius tergosternal sclerite, photos by Nikolai Ignatev, (k) Hepialus humuli tsclerite, (l) Zenophassus schamyl tergosternal sclerite, photos by John Grehan." figureDoi="http://doi.org/10.5281/zenodo.4478061" httpUri="https://zenodo.org/record/4478061/files/figure.png" pageId="12" pageNumber="351">Fig. 23i</figureCitation>
|
||
). The sclerite comprises a dorsal ‘brace’ that forms a right-angled bend between the anterior tergal brace connecting with the posterior thorax, and a dorsal brace that is aligned along the anterior margin of tergite II. Ventral to the brace is a central-posterior ‘intermediate zone’ that varies in size from a broad triangular region to being indistinct from the tergosternal bar that extends between the lateral arms of sternum II to the anterior or central region of the lateral posterior to the first spiracle of the abdomen (
|
||
<figureCitation id="137B2A149B25DA5FA717F8FE9C08F8BE" box="[1050,1141,1843,1870]" captionStart="FIGURE 23" captionStartId="11.[151,250,1794,1818]" captionTargetBox="[164,1427,641,1760]" captionTargetId="figure-242@11.[151,1435,633,1770]" captionTargetPageId="11" captionText="FIGURE 23. Magnificus abdominal sclerites: (a) M. regius tergite II, (MWM), photo by Nikolai Ignatev, (b) M. bouvieri holotype SII (photo by Joël Minet), (c) Magnificus sp. sternites VII-VIII, photo by Nikolai Ignatev), (d) M. bouvieri holotype sternites VII-VIII, photo by Joël Minet), (e) M. regius sternites VII-VIII, photo by Nikolai Ignatev, (f) M. regius and (g) M. roseus, photos by Ulf Buchsbaum, (h) M. bouvieri holotype abdomen, photo by Joël Minet, (i) Magnificus sp. tergosternal sclerite, (j) Magnificus regius tergosternal sclerite, photos by Nikolai Ignatev, (k) Hepialus humuli tsclerite, (l) Zenophassus schamyl tergosternal sclerite, photos by John Grehan." figureDoi="http://doi.org/10.5281/zenodo.4478061" httpUri="https://zenodo.org/record/4478061/files/figure.png" pageId="12" pageNumber="351">Fig. 23i</figureCitation>
|
||
). The tergosternal bar may merge with the intermediate zone or be partially distinct. In
|
||
<taxonomicName id="4C404D129B25DA5FA060F89A9B96F881" authorityName="Yan" authorityYear="2000" box="[877,1003,1879,1905]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="351" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B25DA5FA060F89A9B96F881" box="[877,1003,1879,1905]" italics="true" pageId="12" pageNumber="351">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
the tergosternal sclerites was observed for only two species (
|
||
<figureCitation id="137B2A149B25DA5FA2C9F8B69A67F866" box="[452,538,1915,1942]" captionStart="FIGURES 21–22" captionStartId="10.[151,264,1449,1473]" captionTargetBox="[151,1435,281,1425]" captionTargetId="figure-64@10.[151,1435,273,1425]" captionTargetPageId="10" captionText="FIGURES 21–22. Magnificus thoracic appendages: 21, wing venation a, Magnificus sp. female, b, M. regius male. Diagrams by John Grehan; 22, legs: left to right – pro. meso, meta (a) Magnificus sp. female (MWM), photo by Nikolai Ignatev, (b) Magnificus regius male (ZSBS), (c) Magnificus roseus male, photos by Ulf Buchsbaum." figureDoi="http://doi.org/10.5281/zenodo.4478059" httpUri="https://zenodo.org/record/4478059/files/figure.png" pageId="12" pageNumber="351">Figs 22</figureCitation>
|
||
a-b), but these are similar to each other and they are considered representative of the genus as the shape of this structure is consistent for other recognized monophyletic genera or larger clades (
|
||
<bibRefCitation id="EFD14B609B25DA5FA3D6F80E99FDF82D" author="Grehan, J. R." box="[219,384,1987,2013]" pageId="12" pageNumber="351" pagination="33 - 62" refId="ref10908" refString="Grehan, J. R. (2012 a) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Museum of Science, 42, 33 - 62." type="journal article" year="2012">Grehan 2012a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3F66199B24DA5EA39AF8A19A16F858" ID-DOI="http://doi.org/10.5281/zenodo.4478063" ID-Zenodo-Dep="4478063" httpUri="https://zenodo.org/record/4478063/files/figure.png" pageId="13" pageNumber="352" startId="13.[151,250,1900,1924]" targetBox="[151,1435,181,1876]" targetPageId="13">
|
||
<paragraph id="8BFF36919B24DA5EA39AF8A19A16F858" blockId="13.[151,1436,1900,1960]" pageId="13" pageNumber="352">
|
||
<emphasis id="B934EA839B24DA5EA39AF8A1995CF874" bold="true" box="[151,289,1900,1924]" pageId="13" pageNumber="352">FIGURE 30.</emphasis>
|
||
<taxonomicName id="4C404D129B24DA5EA224F8A199B4F874" authority="FW" authorityName="FW" box="[297,457,1900,1924]" class="Magnoliopsida" family="Rosaceae" genus="Magnificus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="352" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B934EA839B24DA5EA224F8A199E6F874" box="[297,411,1900,1924]" italics="true" pageId="13" pageNumber="352">Magnificus</emphasis>
|
||
FW
|
||
</taxonomicName>
|
||
major banding patterns (coloured outlines). Wing veins—blue lines. FW illustrations 30g, h of limited quality. Higher resolution unavailable.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF36919B27DA5DA3CAFF5A9B7AFE06" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="353">
|
||
The tergosternal sclerite of
|
||
<taxonomicName id="4C404D129B27DA5DA2F2FF5A9A00FF41" box="[511,637,151,177]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA2F2FF5A9A00FF41" box="[511,637,151,177]" italics="true" pageId="14" pageNumber="353">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
comprises a sub-parallel, crescent shaped, tergosternal bar that is continuous with the intermediate zone and fused to the edge of the corner between the lateral and dorsal brace. The posterior margin of the intermediate zone comprises a distinct corner or angle between a straight posteriorly angled dorsal edge and an anteriorly oriented ventral edge extending to the lateral arm of sternum II. The dorsal edge has darker shading that may represent an internal extension of the structure resulting in a darker texture from the external view. Comparison with other genera (cf.
|
||
<bibRefCitation id="EFD14B609B27DA5DA72FFE869CB9FE95" author="Grehan, J. R." box="[1058,1220,331,357]" pageId="14" pageNumber="353" pagination="33 - 62" refId="ref10908" refString="Grehan, J. R. (2012 a) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Museum of Science, 42, 33 - 62." type="journal article" year="2012">Grehan 2012a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD14B609B27DA5DA7DFFE869967FE79" author="Grehan, J. R. & Mielke, C. G. C." pageId="14" pageNumber="353" pagination="45 - 59" refId="ref11340" refString="Grehan, J. R. & Mielke, C. G. C. (2018 a) Description of new species of Yleuxas Viette, 1951 from Peru and taxonomic notes on the genus (Lepidoptera: Hepialidae). The European Entomologist, 9, 45 - 59." type="journal article" year="2018">Grehan & Mielke 2018a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD14B609B27DA5DA22AFEA29910FE79" author="Grehan, J. R. & Mielke, C. G. C." box="[295,365,367,393]" pageId="14" pageNumber="353" pagination="1 - 23" refId="ref11468" refString="Grehan, J. R. & Mielke, C. G. C. (2019 a) Two new species of Palpifer Hampson, 1893 from South East Asia (Lepidoptera: Hepialidae). Bishop Museum Occasional Papers, 125, 1 - 23." type="journal article" year="2019">2019a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD14B609B27DA5DA274FEA299C8FE79" author="Grehan, J. R. & Mielke, C. G. C." box="[377,437,367,393]" pageId="14" pageNumber="353" pagination="1 - 37" refId="ref11572" refString="Grehan, J. R. & Mielke, C. G. C. (2020) Three new species and biogeography of Phassodes Bethune-Baker, 1905 (Lepidoptera: Hepialidae), ghost moth descendants of a subduction zone weed in the south-west Pacific. Bishop Museum Occasional Papers, 136, 1 - 37." type="journal article" year="2020">2020</bibRefCitation>
|
||
; Grehan
|
||
<emphasis id="B934EA839B27DA5DA116FEA29A31FE79" box="[539,588,367,393]" italics="true" pageId="14" pageNumber="353">et al</emphasis>
|
||
. 2019; Mielke &
|
||
<bibRefCitation id="EFD14B609B27DA5DA01BFEA29BCBFE79" author="Grehan, J. R. & Mielke, C. G. C." box="[790,950,367,393]" pageId="14" pageNumber="353" pagination="76 - 77" refId="ref11044" refString="Grehan, J. R. & Mielke, C. G. C. (2016 a) Sthenopis auratus (Grote, 1878), a junior and subjective synomym of Sthenopis pretiosus (Herrich-Schaffer, [1856]) (Lepidoptera: Hepialidae). News of the Lepidopterists' Society, 58, 76 - 77." type="journal article" year="2016">Grehan 2016a</bibRefCitation>
|
||
, 2017,
|
||
<bibRefCitation id="EFD14B609B27DA5DA704FEA29CA7FE79" author="Mielke, C. G. C. & Grehan, J. R. & Grados, J." box="[1033,1242,367,393]" pageId="14" pageNumber="353" pagination="335 - 342" refId="ref12391" refString="Mielke, C. G. C., Grehan, J. R. & Grados, J. (2019) Description of Walkeriella, gen. n., with a new species from Madre de Dios dept., southeastern Peru (Lepidoptera: Hepialoidea: Hepialidae). Zootaxa, 4609 (2), 335 - 342. https: // doi. org / 10.11646 / zootaxa. 4609.2.8" type="journal article" year="2019">
|
||
Mielke
|
||
<emphasis id="B934EA839B27DA5DA76DFEA29CECFE79" box="[1120,1169,367,393]" italics="true" pageId="14" pageNumber="353">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
) indicate that the tergosternal sclerite shape in
|
||
<taxonomicName id="4C404D129B27DA5DA115FE5E9AEBFE5D" box="[536,662,403,429]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA115FE5E9AEBFE5D" box="[536,662,403,429]" italics="true" pageId="14" pageNumber="353">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
is unique within the
|
||
<taxonomicName id="4C404D129B27DA5DA072FE5E9B85FE5D" baseAuthorityName="Grehan" baseAuthorityYear="2012" box="[895,1016,403,429]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
and therefore supports monophyly of the genus, and there is no evidence of any specialized shared similarity with the northern Eurasian sister group (as illustrated here for
|
||
<emphasis id="B934EA839B27DA5DA2DCFE169AEFFE05" box="[465,658,475,501]" italics="true" pageId="14" pageNumber="353">
|
||
<taxonomicName id="4C404D129B27DA5DA2DCFE169AF3FE05" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[465,654,475,501]" class="Insecta" family="Hepialidae" genus="Hepialus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="humuli">Hepialus humuli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<figureCitation id="137B2A149B27DA5DA194FE169A87FE06" box="[665,762,475,502]" captionStart="FIGURE 23" captionStartId="11.[151,250,1794,1818]" captionTargetBox="[164,1427,641,1760]" captionTargetId="figure-242@11.[151,1435,633,1770]" captionTargetPageId="11" captionText="FIGURE 23. Magnificus abdominal sclerites: (a) M. regius tergite II, (MWM), photo by Nikolai Ignatev, (b) M. bouvieri holotype SII (photo by Joël Minet), (c) Magnificus sp. sternites VII-VIII, photo by Nikolai Ignatev), (d) M. bouvieri holotype sternites VII-VIII, photo by Joël Minet), (e) M. regius sternites VII-VIII, photo by Nikolai Ignatev, (f) M. regius and (g) M. roseus, photos by Ulf Buchsbaum, (h) M. bouvieri holotype abdomen, photo by Joël Minet, (i) Magnificus sp. tergosternal sclerite, (j) Magnificus regius tergosternal sclerite, photos by Nikolai Ignatev, (k) Hepialus humuli tsclerite, (l) Zenophassus schamyl tergosternal sclerite, photos by John Grehan." figureDoi="http://doi.org/10.5281/zenodo.4478061" httpUri="https://zenodo.org/record/4478061/files/figure.png" pageId="14" pageNumber="353">Fig. 23k</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF36919B27DA5DA39AFE339C7CFCE6" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="353">
|
||
<emphasis id="B934EA839B27DA5DA39AFE33999DFDE9" bold="true" box="[151,480,510,537]" pageId="14" pageNumber="353">(ii) Pseudoteguminal arms.</emphasis>
|
||
At the posterio-ventral junction of the pseudotegumen margin either side of the anogenital field each pseudotegumen plate may also be produced into a ‘pseudoteguminal arm’ of variable length in the
|
||
<taxonomicName id="4C404D129B27DA5DA21AFD8A99E9FD91" baseAuthorityName="Grehan" baseAuthorityYear="2012" box="[279,404,583,609]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C404D129B27DA5DA2B6FD8A9A44FD91" box="[443,569,583,609]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA2B6FD8A9A44FD91" box="[443,569,583,609]" italics="true" pageId="14" pageNumber="353">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
the arms are connected across the median to form a broad, ventrally open tube or channel. This formation is not always visible in slide mounted material as with
|
||
<taxonomicName id="4C404D129B27DA5DA778FDA69C85FD75" authorityName="Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue" authorityYear="2021" baseAuthorityName="Oberthur" baseAuthorityYear="1913" box="[1141,1272,619,645]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="bouvieri">
|
||
<emphasis id="B934EA839B27DA5DA778FDA69C85FD75" box="[1141,1272,619,645]" italics="true" pageId="14" pageNumber="353">M. bouvieri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137B2A149B27DA5DA605FDA69D21FD75" box="[1288,1372,619,645]" captionStart="FIGURES 10–20" captionStartId="8.[151,264,1687,1711]" captionTargetBox="[177,1410,181,1664]" captionTargetId="figure-22@8.[177,1410,181,1664]" captionTargetPageId="8" captionText="FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev." figureDoi="http://doi.org/10.5281/zenodo.4478055" httpUri="https://zenodo.org/record/4478055/files/figure.png" pageId="14" pageNumber="353">Fig. 10</figureCitation>
|
||
). The structure is visible in an unmounted specimen of
|
||
<taxonomicName id="4C404D129B27DA5DA1F9FD429B1CFD58" authorityName="Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue" authorityYear="2021" baseAuthorityName="Oberthur" baseAuthorityYear="1911" box="[756,865,655,680]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="roseus">
|
||
<emphasis id="B934EA839B27DA5DA1F9FD429B1CFD58" box="[756,865,655,680]" italics="true" pageId="14" pageNumber="353">M. roseus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137B2A149B27DA5DA062FD429BB0FD5A" box="[879,973,655,682]" captionStart="FIGURES 10–20" captionStartId="8.[151,264,1687,1711]" captionTargetBox="[177,1410,181,1664]" captionTargetId="figure-22@8.[177,1410,181,1664]" captionTargetPageId="8" captionText="FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev." figureDoi="http://doi.org/10.5281/zenodo.4478055" httpUri="https://zenodo.org/record/4478055/files/figure.png" pageId="14" pageNumber="353">Fig. 13b</figureCitation>
|
||
) but not in the compressed
|
||
<typeStatus id="54FB88339B27DA5DA7F2FD429D1CFD59" box="[1279,1377,655,681]" pageId="14" pageNumber="353" type="holotype">holotype</typeStatus>
|
||
slide (
|
||
<figureCitation id="137B2A149B27DA5DA3D1FD7E9946FD3D" box="[220,315,691,717]" captionStart="FIGURES 10–20" captionStartId="8.[151,264,1687,1711]" captionTargetBox="[177,1410,181,1664]" captionTargetId="figure-22@8.[177,1410,181,1664]" captionTargetPageId="8" captionText="FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev." figureDoi="http://doi.org/10.5281/zenodo.4478055" httpUri="https://zenodo.org/record/4478055/files/figure.png" pageId="14" pageNumber="353">Fig. 13a</figureCitation>
|
||
). The channel is visible in the dissections of
|
||
<taxonomicName id="4C404D129B27DA5DA020FD7E9B8BFD3C" authorityName="Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue" authorityYear="2021" baseAuthorityName="Staudinger" baseAuthorityYear="1896" box="[813,1014,691,717]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="regius">
|
||
<emphasis id="B934EA839B27DA5DA020FD7E9B8BFD3C" box="[813,1014,691,717]" italics="true" pageId="14" pageNumber="353">Magnificus regius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137B2A149B27DA5DA709FD7E9CE0FD3D" box="[1028,1181,691,717]" captionStart="FIGURES 10–20" captionStartId="8.[151,264,1687,1711]" captionTargetBox="[177,1410,181,1664]" captionTargetId="figure-22@8.[177,1410,181,1664]" captionTargetPageId="8" captionText="FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev." figureDoi="http://doi.org/10.5281/zenodo.4478055" httpUri="https://zenodo.org/record/4478055/files/figure.png" pageId="14" pageNumber="353">Figs 12a. b. c</figureCitation>
|
||
). The structure may be inferred for the slide mount of
|
||
<taxonomicName id="4C404D129B27DA5DA123FD1A9ADFFD01" authorityName="Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue" authorityYear="2021" baseAuthorityName="Bang-Haas" baseAuthorityYear="1939" box="[558,674,727,753]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="dirschi">
|
||
<emphasis id="B934EA839B27DA5DA123FD1A9ADFFD01" box="[558,674,727,753]" italics="true" pageId="14" pageNumber="353">M. dirschi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137B2A149B27DA5DA1BCFD1A9B7EFD01" box="[689,771,727,753]" captionStart="FIGURES 10–20" captionStartId="8.[151,264,1687,1711]" captionTargetBox="[177,1410,181,1664]" captionTargetId="figure-22@8.[177,1410,181,1664]" captionTargetPageId="8" captionText="FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev." figureDoi="http://doi.org/10.5281/zenodo.4478055" httpUri="https://zenodo.org/record/4478055/files/figure.png" pageId="14" pageNumber="353">Fig. 11</figureCitation>
|
||
). Further study of this character is required. The only other genus we have seen anything similar is
|
||
<taxonomicName id="4C404D129B27DA5DA19DFD369A88FCE5" authorityName="Fabricius" authorityYear="1775" box="[656,757,763,789]" class="Insecta" family="Hepialidae" genus="Hepialus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA19DFD369A88FCE5" box="[656,757,763,789]" italics="true" pageId="14" pageNumber="353">Hepialus</emphasis>
|
||
</taxonomicName>
|
||
(see discussion below).
|
||
</paragraph>
|
||
<paragraph id="8BFF36919B27DA5DA39AFCD39C56FC1D" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="353">
|
||
<emphasis id="B934EA839B27DA5DA39AFCD3999DFCC8" bold="true" box="[151,480,798,825]" pageId="14" pageNumber="353">(iii) Lamella antevaginalis.</emphasis>
|
||
The female the shape of the lamella antevaginalis in
|
||
<taxonomicName id="4C404D129B27DA5DA720FCD29CD6FCC9" box="[1069,1195,799,825]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA720FCD29CD6FCC9" box="[1069,1195,799,825]" italics="true" pageId="14" pageNumber="353">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
appears to be distinct with respect to its dorso-ventrally shallow sub-rectangular shape without dorsal development of the lateral lobes and minimal development of the central lobe. However, the structural details are insufficiently clear in the available dissections, and published diagrams for two other species are too generalized to fully characterize the lamella antevaginalis of
|
||
<taxonomicName id="4C404D129B27DA5DA11EFC629AECFC39" box="[531,657,943,969]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA11EFC629AECFC39" box="[531,657,943,969]" italics="true" pageId="14" pageNumber="353">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
with respect to other
|
||
<taxonomicName id="4C404D129B27DA5DA084FC629C7BFC39" baseAuthorityName="Grehan" baseAuthorityYear="2012" box="[905,1030,943,969]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
. Full corroboration of this character state for
|
||
<taxonomicName id="4C404D129B27DA5DA23BFC1E99C9FC1D" box="[310,436,979,1005]" class="Insecta" family="Hepialidae" genus="Magnificus" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EA839B27DA5DA23BFC1E99C9FC1D" box="[310,436,979,1005]" italics="true" pageId="14" pageNumber="353">Magnificus</emphasis>
|
||
</taxonomicName>
|
||
will require more detailed documentation for the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |