729 lines
73 KiB
XML
729 lines
73 KiB
XML
<document ID-DOI="10.1071/is21067" ID-GBIF-Dataset="8c0715c9-57f5-40db-8c48-99bf3719eedb" ID-Zenodo-Dep="6987297" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1660328621757" checkinUser="felipe" docAuthor="Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan & Matos-Maraví, Pável" docDate="2022" docId="03E5E9156515053AFF7AFBAC9AB92A01" docLanguage="en" docName="InvSyst.36.6.557-579.pdf" docOrigin="Invertebrate Systematics 36 (6)" docSource="http://dx.doi.org/10.1071/is21067" docTitle="Overbeckia subclavata Viehmeyer 1916" docType="treatment" docVersion="3" lastPageNumber="569" masterDocId="FFDC916D651D0536FFBCFFF69F1B2B6D" masterDocTitle="Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae)" masterLastPageNumber="288" masterPageNumber="277" pageNumber="565" updateTime="1660340715462" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klimeš, Petr</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, BraniŠovská 31, CZ- 370 05 České Budějovice, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Drescher, Jochen</mods:namePart>
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<mods:affiliation>Department of Animal Ecology, University of Göttingen, J. - F. - Blumenbach Institute for Zoology & Anthropology, Untere Karspüle 2, D- 37073 Göttingen, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Buchori, Damayanti</mods:namePart>
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<mods:affiliation>Department of Plant Protection, Faculty of Agriculture, Bogor Agricultural University, Jl. Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hidayat, Purnama</mods:namePart>
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<mods:affiliation>Department of Plant Protection, Faculty of Agriculture, Bogor Agricultural University, Jl. Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nazarreta, Rizky</mods:namePart>
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<mods:affiliation>Department of Plant Protection, Faculty of Agriculture, Bogor Agricultural University, Jl. Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Potocký, Pavel</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, BraniŠovská 31, CZ- 370 05 České Budějovice, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Rimandai, Maling</mods:namePart>
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<mods:affiliation>New Guinea Binatang Research Center, PO Box 604, Madang, Papua New Guinea</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Scheu, Stefan</mods:namePart>
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<mods:affiliation>Department of Animal Ecology, University of Göttingen, J. - F. - Blumenbach Institute for Zoology & Anthropology, Untere Karspüle 2, D- 37073 Göttingen, Germany & Centre of Biodiversity and Sustainable Land Use, University of Göttingen, Büsgenweg 1, D- 37077 Göttingen, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Matos-Maraví, Pável</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, BraniŠovská 31, CZ- 370 05 České Budějovice, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Invertebrate Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-25</mods:number>
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||
</mods:detail>
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<mods:detail type="volume">
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<mods:number>36</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>6</mods:number>
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||
</mods:detail>
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<mods:extent unit="page">
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<mods:start>277</mods:start>
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<mods:end>288</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1071/is21067</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.1071/is21067</mods:identifier>
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<mods:identifier type="GBIF-Dataset">8c0715c9-57f5-40db-8c48-99bf3719eedb</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6987297</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6987100" ID-Zenodo-Dep="6987100" LSID="urn:lsid:plazi:treatment:03E5E9156515053AFF7AFBAC9AB92A01" httpUri="http://treatment.plazi.org/id/03E5E9156515053AFF7AFBAC9AB92A01" lastPageId="12" lastPageNumber="569" pageId="8" pageNumber="565">
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<subSubSection box="[198,669,1114,1145]" pageId="8" pageNumber="565" type="nomenclature">
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||
<paragraph blockId="8.[198,669,1114,1145]" box="[198,669,1114,1145]" pageId="8" pageNumber="565">
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||
<heading bold="true" box="[198,669,1114,1145]" centered="true" fontSize="12" level="2" pageId="8" pageNumber="565" reason="0">
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||
<taxonomicName authority="Viehmeyer" authorityName="Viehmeyer" authorityYear="1916" box="[198,669,1114,1145]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="565" phylum="Arthropoda" rank="species" species="subclavata">
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||
<emphasis bold="true" box="[198,669,1114,1145]" pageId="8" pageNumber="565">
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||
<emphasis bold="true" box="[198,519,1114,1145]" italics="true" pageId="8" pageNumber="565">Overbeckia subclavata</emphasis>
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||
Viehmeyer
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection box="[377,491,1183,1207]" pageId="8" pageNumber="565" type="description">
|
||
<paragraph blockId="8.[377,491,1183,1207]" box="[377,491,1183,1207]" pageId="8" pageNumber="565">
|
||
<heading box="[377,491,1183,1207]" centered="true" fontSize="10" level="4" pageId="8" pageNumber="565" reason="4">
|
||
(
|
||
<figureCitation box="[386,451,1183,1207]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="8" pageNumber="565">Fig. 1</figureCitation>
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,
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<figureCitation box="[467,481,1183,1207]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="8" pageNumber="565">2</figureCitation>
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)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="8" pageNumber="565" type="etymology">
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||
<paragraph blockId="8.[98,245,1277,1305]" box="[98,245,1277,1305]" pageId="8" pageNumber="565">
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||
<heading bold="true" box="[98,245,1277,1305]" fontSize="11" level="3" pageId="8" pageNumber="565" reason="0">
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||
<emphasis bold="true" box="[98,245,1277,1305]" pageId="8" pageNumber="565">Etymology</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph blockId="8.[98,770,1328,1480]" pageId="8" pageNumber="565">
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||
<bibRefCitation author="Viehmeyer H" box="[98,306,1328,1352]" pageId="8" pageNumber="565" pagination="108 - 168" refId="ref20034" refString="Viehmeyer H (1916) Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Archiv fur Naturgeschichte - A 81 (8), 108 - 168." type="journal article" year="1916">Viehmeyer (1916)</bibRefCitation>
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does not explain the reasoning for the species name, but this likely refers to the club-shaped anterior part of antennae, mentioned in the original description (‘Geissel gegen Ende verdickt’: funinculus enlarged towards the tip) (
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<figureCitation box="[200,277,1455,1480]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="8" pageNumber="565">
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||
Fig. 2
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||
<emphasis bold="true" box="[264,277,1455,1479]" italics="true" pageId="8" pageNumber="565">h</emphasis>
|
||
</figureCitation>
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||
).
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="8" pageNumber="565" type="materials_examined">
|
||
<paragraph blockId="8.[98,771,1515,1794]" box="[98,356,1515,1543]" pageId="8" pageNumber="565">
|
||
<heading bold="true" box="[98,356,1515,1543]" fontSize="11" level="3" pageId="8" pageNumber="565" reason="0">
|
||
<emphasis bold="true" box="[98,356,1515,1543]" pageId="8" pageNumber="565">Material examined</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph blockId="8.[98,771,1515,1794]" pageId="8" pageNumber="565">
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||
<emphasis bold="true" box="[98,179,1561,1580]" italics="true" pageId="8" pageNumber="565">
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||
<typeStatus box="[98,179,1561,1580]" pageId="8" pageNumber="565">Syntypes</typeStatus>
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||
</emphasis>
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||
. All examined from photos (AntWeb codes):
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<materialsCitation collectedFrom="Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree" collectionCode="NHMB" collectorName="H. Overbeck" country="Singapore" determinerName="Viehmeyer" latitude="1.2930555" location="Singapore" longLatPrecision="1" longitude="103.855835" pageId="8" pageNumber="565" specimenCode="CASENT0101191, CASENT0101192" specimenCount="1" typeStatus="syntype">
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||
<specimenCode box="[606,767,1562,1581]" pageId="8" pageNumber="565">CASENT0101191</specimenCode>
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,
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<specimenCode box="[98,258,1588,1607]" pageId="8" pageNumber="565">CASENT0101192</specimenCode>
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, workers (
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<collectionCode box="[364,430,1588,1607]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="565">NHMB</collectionCode>
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||
;
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||
<collectorName box="[444,548,1588,1608]" pageId="8" pageNumber="565">F. Santschi</collectorName>
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collection);
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</materialsCitation>
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||
<materialsCitation collectedFrom="Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree" collectionCode="DEIC" collectorName="H. Overbeck" country="Singapore" determinerName="Viehmeyer" latitude="1.2930555" location="Singapore" longLatPrecision="1" longitude="103.855835" pageId="8" pageNumber="565" specimenCode="focol0130" specimenCount="1" specimenCount-worker="1" typeStatus="syntype">
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<specimenCode box="[673,767,1588,1607]" pageId="8" pageNumber="565">focol0130</specimenCode>
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,
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<specimenCount box="[98,164,1615,1634]" pageId="8" pageNumber="565" type="worker">worker</specimenCount>
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||
(
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<collectionCode box="[178,230,1615,1634]" collectionName="Germany, Muencheberg, Deutsches Entomologisches Institut im ZALF" pageId="8" pageNumber="565">DEIC</collectionCode>
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, marked as ‘
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<typeStatus box="[351,446,1615,1634]" pageId="8" pageNumber="565">Syntypus’</typeStatus>
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||
);
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||
</materialsCitation>
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||
<materialsCitation collectedFrom="Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree" collectionCode="DEIC" collectorName="H. Overbeck" country="Singapore" determinerName="Viehmeyer" latitude="1.2930555" location="Singapore" longLatPrecision="1" longitude="103.855835" pageId="8" pageNumber="565" specimenCode="focol2566" specimenCount="1" specimenCount-queen="1" typeStatus="syntype">
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<specimenCode box="[465,561,1615,1634]" pageId="8" pageNumber="565">focol2566</specimenCode>
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, alate
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<specimenCount box="[626,682,1615,1634]" pageId="8" pageNumber="565" type="queen">queen</specimenCount>
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||
(marked as ‘
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||
<typeStatus box="[140,207,1642,1661]" pageId="8" pageNumber="565">Typus’</typeStatus>
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||
),
|
||
</materialsCitation>
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||
<materialsCitation collectedFrom="Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree" collectionCode="ZMHB" collectorName="H. Overbeck" country="Singapore" determinerName="Viehmeyer" latitude="1.2930555" location="Singapore" longLatPrecision="1" longitude="103.855835" pageId="8" pageNumber="565" specimenCode="focol2567, focol2568, focol2564, focol2565" specimenCount="2" specimenCount-male="1" specimenCount-worker="1" typeStatus="syntype">
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||
<specimenCode box="[236,329,1641,1660]" pageId="8" pageNumber="565">focol2567</specimenCode>
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and
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||
<specimenCode box="[403,500,1641,1660]" pageId="8" pageNumber="565">focol2568</specimenCode>
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||
,
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||
<specimenCount box="[522,599,1641,1660]" pageId="8" pageNumber="565" type="worker">workers</specimenCount>
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||
,
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||
<specimenCode box="[622,716,1641,1660]" pageId="8" pageNumber="565">focol2564</specimenCode>
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||
and
|
||
<specimenCode box="[98,193,1668,1687]" pageId="8" pageNumber="565">focol2565</specimenCode>
|
||
,
|
||
<specimenCount box="[209,263,1668,1687]" pageId="8" pageNumber="565" type="male">males</specimenCount>
|
||
(
|
||
<collectionCode box="[281,345,1668,1687]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="8" pageNumber="565">ZMHB</collectionCode>
|
||
, all marked as ‘
|
||
<typeStatus box="[513,580,1668,1687]" pageId="8" pageNumber="565">Typus’</typeStatus>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation collectedFrom="Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree" collectionCode="MCZC" collectorName="H. Overbeck" country="Singapore" determinerName="Viehmeyer" latitude="1.2930555" location="Singapore" longLatPrecision="1" longitude="103.855835" pageId="8" pageNumber="565" specimenCode="CASENT0178502" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
|
||
<specimenCode box="[602,766,1668,1687]" pageId="8" pageNumber="565">CASENT0178502</specimenCode>
|
||
,
|
||
<specimenCount box="[98,166,1695,1714]" pageId="8" pageNumber="565" type="worker">worker</specimenCount>
|
||
(
|
||
<collectionCode box="[180,244,1695,1714]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="8" pageNumber="565">MCZC</collectionCode>
|
||
, marked as ‘
|
||
<typeStatus box="[367,455,1695,1714]" pageId="8" pageNumber="565">Paratype</typeStatus>
|
||
30741’)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectedFrom="Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree" collectionCode="MSGN" collectorName="H. Overbeck" country="Singapore" determinerName="Viehmeyer" latitude="1.2930555" location="Singapore" longLatPrecision="1" longitude="103.855835" pageId="8" pageNumber="565" specimenCode="CASENT0905180" specimenCount="1">
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||
<specimenCode box="[552,714,1695,1714]" pageId="8" pageNumber="565">CASENT0905180</specimenCode>
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||
, male (
|
||
<collectionCode box="[104,171,1721,1740]" collectionName="MSGN" pageId="8" pageNumber="565">MSGN</collectionCode>
|
||
)/coll.
|
||
<collectorName box="[236,361,1721,1741]" pageId="8" pageNumber="565">H. Overbeck</collectorName>
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||
, <1915/
|
||
<collectingCountry box="[447,547,1721,1740]" name="Singapore" pageId="8" pageNumber="565">Singapore</collectingCountry>
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||
/
|
||
<collectedFrom pageId="8" pageNumber="565">Nests in dry/narrow, hollow bamboo, colony in the hollow branch of a mango tree</collectedFrom>
|
||
/
|
||
<geoCoordinate box="[98,211,1774,1794]" degrees="1" direction="north" minutes="17" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="35.0" value="1.2930555">1°17′35.0″N</geoCoordinate>
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||
,
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||
<geoCoordinate box="[222,358,1774,1794]" degrees="103" direction="east" minutes="51" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="21.0" value="103.855835">103°51′21.0″E</geoCoordinate>
|
||
/det.
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||
<bibRefCitation author="Viehmeyer H" box="[405,574,1775,1794]" pageId="8" pageNumber="565" pagination="108 - 168" refId="ref20034" refString="Viehmeyer H (1916) Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Archiv fur Naturgeschichte - A 81 (8), 108 - 168." type="journal article" year="1916">
|
||
<determinerName box="[405,504,1775,1794]" pageId="8" pageNumber="565">Viehmeyer</determinerName>
|
||
(1916)
|
||
</bibRefCitation>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="8.[98,511,1838,1866]" box="[98,511,1838,1866]" pageId="8" pageNumber="565">
|
||
<heading bold="true" box="[98,511,1838,1866]" fontSize="11" level="3" pageId="8" pageNumber="565" reason="0">
|
||
<emphasis bold="true" box="[98,511,1838,1866]" pageId="8" pageNumber="565">Additional material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[98,771,1885,1931]" lastBlockId="8.[818,1491,1116,1457]" pageId="8" pageNumber="565">
|
||
<emphasis bold="true" box="[98,201,1885,1904]" pageId="8" pageNumber="565">
|
||
<collectingCountry box="[98,197,1885,1904]" name="Indonesia" pageId="8" pageNumber="565">Indonesia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount box="[208,301,1885,1904]" pageId="8" pageNumber="565" type="worker">5 workers</specimenCount>
|
||
and
|
||
<specimenCount box="[351,428,1885,1904]" pageId="8" pageNumber="565" type="queen">1 queen</specimenCount>
|
||
; all leg. J. Drescher, canopy fogging (det.
|
||
<collectionCode box="[160,177,1912,1931]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="565">R</collectionCode>
|
||
. Nazarreta & P. Klimes): Worker (
|
||
<collectionCode box="[571,636,1912,1931]" pageId="8" pageNumber="565">ZMHB</collectionCode>
|
||
): HJ3.1/
|
||
<date box="[732,865,1117,1931]" pageId="8" pageNumber="565" value="2013-11-05">5.xi. 2013</date>
|
||
/Harapan Rainforest/jungle rubber/dry season/
|
||
<geoCoordinate box="[1365,1486,1116,1136]" degrees="01" direction="south" minutes="50" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="57.7" value="-1.8493611">01°50′57.7″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[818,953,1143,1163]" degrees="103" direction="east" minutes="17" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="59.7" value="103.29992">103°17′59.7″E</geoCoordinate>
|
||
. Worker (
|
||
<collectionCode box="[1082,1131,1144,1163]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="565" type="Museum">MZB</collectionCode>
|
||
): BJ6.2/
|
||
<date box="[1229,1317,1144,1163]" pageId="8" pageNumber="565" value="2013-12-01">1.xii.2013</date>
|
||
/Bukit Duabelas National Park/jungle rubber/rainy season/
|
||
<geoCoordinate box="[1219,1340,1170,1190]" degrees="02" direction="south" minutes="01" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="48.4" value="-2.030111">02°01′48.4″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1352,1486,1170,1190]" degrees="102" direction="east" minutes="46" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="14.9" value="102.770805">102°46′14.9″E</geoCoordinate>
|
||
. Worker (
|
||
<collectionCode box="[905,958,1198,1217]" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="8" pageNumber="565" type="Other institutional type">IECA</collectionCode>
|
||
): BF3.1/
|
||
<date box="[1043,1119,1197,1216]" pageId="8" pageNumber="565" value="2013-10-05">5.x.2013</date>
|
||
/Bukit Duabelas National Park/jungle rubber/rainy season/
|
||
<geoCoordinate box="[1030,1150,1223,1243]" degrees="01" direction="south" minutes="56" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="34.1" value="-1.9428056">01°56′34.1″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1175,1310,1223,1243]" degrees="102" direction="east" minutes="34" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="51.5" value="102.58097">102°34′51.5″E</geoCoordinate>
|
||
. Worker (
|
||
<collectionCode box="[1431,1480,1224,1243]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="565" type="Museum">MZB</collectionCode>
|
||
): BF1.1/
|
||
<date box="[881,956,1251,1270]" pageId="8" pageNumber="565" value="2013-10-08">8.x.2013</date>
|
||
/Bukit Duabelas National Park/forest/dry season/
|
||
<geoCoordinate box="[818,939,1276,1296]" degrees="01" direction="south" minutes="59" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="41.4" value="-1.9948334">01°59′41.4″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[970,1104,1276,1296]" degrees="102" direction="east" minutes="45" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="08.5" value="102.75236">102°45′08.5″E</geoCoordinate>
|
||
. Worker (
|
||
<collectionCode box="[1236,1294,1278,1297]" pageId="8" pageNumber="565">CASC</collectionCode>
|
||
): CASENT0872838 (AntWeb)/HymFrm285
|
||
<date box="[1042,1130,1304,1323]" pageId="8" pageNumber="565" value="2013-12-01">1.xii.2013</date>
|
||
/Bukit Duabelas National Park/jungle rubber/rainy season/
|
||
<geoCoordinate box="[1013,1135,1330,1350]" degrees="02" direction="south" minutes="00" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="56.2" value="-2.0156112">02°00′56.2″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1146,1281,1330,1350]" degrees="102" direction="east" minutes="45" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="11.5" value="102.7532">102°45′11.5″E</geoCoordinate>
|
||
. Queen without wings (
|
||
<collectionCode box="[823,876,1358,1377]" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="8" pageNumber="565" type="Other institutional type">IECA</collectionCode>
|
||
): HF4.1/
|
||
<date box="[981,1075,1357,1376]" pageId="8" pageNumber="565" value="2013-09-12">12.ix.2013</date>
|
||
/Harapan Rainforest/forest/dry season/
|
||
<geoCoordinate box="[818,938,1383,1403]" degrees="02" direction="south" minutes="11" orientation="latitude" pageId="8" pageNumber="565" precision="1" seconds="15.4" value="-2.187611">02°11′15.4″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[958,1093,1383,1403]" degrees="103" direction="east" minutes="20" orientation="longitude" pageId="8" pageNumber="565" precision="1" seconds="34.7" value="103.34297">103°20′34.7″E</geoCoordinate>
|
||
.
|
||
<emphasis bold="true" box="[1112,1224,1384,1403]" pageId="8" pageNumber="565">
|
||
<collectingCountry box="[1112,1224,1384,1403]" name="Philippines" pageId="8" pageNumber="565">Philippines</collectingCountry>
|
||
</emphasis>
|
||
(photos): Acc. No. 3576 (
|
||
<collectionCode box="[823,887,1411,1430]" pageId="8" pageNumber="565">MCZC</collectionCode>
|
||
)/Lot Bu.of Sci., P.I./coll. M. Ramos (further details in
|
||
<bibRefCitation author="General DM & Alpert GD" pageId="8" pageNumber="565" pagination="1 - 111" refId="ref17845" refString="General DM, Alpert GD (2012) A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines. ZooKeys 200, 1 - 111. doi: 10.3897 / zookeys. 200.2447" type="journal article" year="2012">General and Alpert 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="565" type="diagnosis">
|
||
<paragraph blockId="8.[818,948,1505,1533]" box="[818,948,1505,1533]" pageId="8" pageNumber="565">
|
||
<heading bold="true" box="[818,948,1505,1533]" fontSize="11" level="3" pageId="8" pageNumber="565" reason="0">
|
||
<emphasis bold="true" box="[818,948,1505,1533]" pageId="8" pageNumber="565">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[818,1490,1555,1933]" pageId="8" pageNumber="565">
|
||
Compared to the other two
|
||
<taxonomicName box="[1156,1370,1555,1580]" pageId="8" pageNumber="565">
|
||
<emphasis bold="true" box="[1156,1278,1555,1579]" italics="true" pageId="8" pageNumber="565">Overbeckia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
described here,
|
||
<taxonomicName authority="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví, 2022" authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[887,1117,1588,1612]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="565" phylum="Arthropoda" rank="species" species="papuana" status="sp. nov.">
|
||
<emphasis bold="true" box="[887,1117,1588,1612]" italics="true" pageId="8" pageNumber="565">Overbeckia papuana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1131,1225,1588,1611]" pageId="8" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví, 2022" authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[1292,1444,1588,1612]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="565" phylum="Arthropoda" rank="species" species="jambiensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[1292,1444,1588,1612]" italics="true" pageId="8" pageNumber="565">O. jambiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="8" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, workers and queens in
|
||
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1161,1314,1620,1644]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="565" phylum="Arthropoda" rank="species" species="subclavata">
|
||
<emphasis bold="true" box="[1161,1314,1620,1644]" italics="true" pageId="8" pageNumber="565">O. subclavata</emphasis>
|
||
</taxonomicName>
|
||
are less hairy, with a lack of long erect hairs on dorsum of petiole, on thorax and on scape except the apical margin; head integument with fine dense punctation; no erect hairs extending past head margins in frontal view; frontal carinae approaching one another less anteriorly compared to those in
|
||
<taxonomicName authority="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví, 2022" authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[851,982,1812,1836]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="565" phylum="Arthropoda" rank="species" species="papuana" status="sp. nov.">
|
||
<emphasis bold="true" box="[851,982,1812,1836]" italics="true" pageId="8" pageNumber="565">O. papuana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[994,1086,1813,1836]" pageId="8" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<collectionCode box="[1103,1161,1812,1836]" pageId="8" pageNumber="565">FCDI</collectionCode>
|
||
<166;
|
||
<figureCitation box="[1258,1337,1812,1836]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="8" pageNumber="565">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[1325,1337,1812,1836]" italics="true" pageId="8" pageNumber="565">b</emphasis>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation box="[1357,1384,1812,1836]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="8" pageNumber="565">
|
||
2
|
||
<emphasis bold="true" box="[1371,1384,1812,1836]" italics="true" pageId="8" pageNumber="565">h</emphasis>
|
||
</figureCitation>
|
||
). Longer erect hairs present only on clypeus and ventral abdomen. Fine straight lineation on lateral sides of thorax limited to the basal region below metanotal groove (
|
||
<figureCitation box="[1299,1377,1908,1932]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="8" pageNumber="565">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[1364,1377,1908,1931]" italics="true" pageId="8" pageNumber="565">a</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.6987305" ID-Zenodo-Dep="6987305" httpUri="https://zenodo.org/record/6987305/files/figure.png" pageId="9" pageNumber="566" startId="9.[314,351,935,958]" targetBox="[314,1274,185,905]" targetPageId="9">
|
||
<paragraph blockId="9.[314,1274,935,1017]" pageId="9" pageNumber="566">
|
||
Fig. 4.
|
||
<taxonomicName authority="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví, 2022" authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[405,606,935,958]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="566" phylum="Arthropoda" rank="species" species="jambiensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[405,606,935,958]" italics="true" pageId="9" pageNumber="566">Overbeckia jambiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[613,689,935,958]" pageId="9" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype (worker) and lateral (
|
||
<emphasis bold="true" box="[1002,1013,935,958]" italics="true" pageId="9" pageNumber="566">a</emphasis>
|
||
), frontal (
|
||
<emphasis bold="true" box="[1113,1124,935,958]" italics="true" pageId="9" pageNumber="566">b</emphasis>
|
||
) and dorsal (
|
||
<emphasis bold="true" box="[1257,1266,935,958]" italics="true" pageId="9" pageNumber="566">c</emphasis>
|
||
) views. Detail of head sculpture and shape of frontal carinae (
|
||
<emphasis bold="true" box="[923,934,964,987]" italics="true" pageId="9" pageNumber="566">d</emphasis>
|
||
). Note that the right antenna has been glued into the antennal socket.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.6987307" ID-Zenodo-Dep="6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="9" pageNumber="566" startId="9.[314,351,1878,1901]" targetBox="[314,1274,1129,1848]" targetPageId="9">
|
||
<paragraph blockId="9.[314,1274,1878,1931]" pageId="9" pageNumber="566">
|
||
Fig. 5.
|
||
<taxonomicName authority="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví, 2022" authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[404,591,1878,1901]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="566" phylum="Arthropoda" rank="species" species="papuana" status="sp. nov.">
|
||
<emphasis bold="true" box="[404,591,1878,1901]" italics="true" pageId="9" pageNumber="566">Overbeckia papuana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[597,672,1878,1901]" pageId="9" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype (worker) and its lateral (
|
||
<emphasis bold="true" box="[1008,1019,1878,1901]" italics="true" pageId="9" pageNumber="566">a</emphasis>
|
||
), frontal (
|
||
<emphasis bold="true" box="[1117,1128,1878,1901]" italics="true" pageId="9" pageNumber="566">b</emphasis>
|
||
) and dorsal (
|
||
<emphasis bold="true" box="[1257,1266,1878,1901]" italics="true" pageId="9" pageNumber="566">c</emphasis>
|
||
) views. Detail of head sculpture and shape of frontal carinae (
|
||
<emphasis bold="true" box="[916,927,1908,1931]" italics="true" pageId="9" pageNumber="566">d</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.6987309" ID-Zenodo-Dep="6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="10" pageNumber="567" startId="10.[154,191,1653,1676]" targetBox="[154,1434,177,1622]" targetPageId="10">
|
||
<paragraph blockId="10.[154,1434,1652,1705]" pageId="10" pageNumber="567">
|
||
Fig. 6.
|
||
<taxonomicName authority="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví, 2022" authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[249,441,1652,1676]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="567" phylum="Arthropoda" rank="species" species="papuana" status="sp. nov.">
|
||
<emphasis bold="true" box="[249,441,1652,1676]" italics="true" pageId="10" pageNumber="567">Overbeckia papuana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[452,532,1653,1676]" pageId="10" pageNumber="567" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratypes (alates). Queen: lateral (
|
||
<emphasis bold="true" box="[896,907,1653,1676]" italics="true" pageId="10" pageNumber="567">a</emphasis>
|
||
), dorsal (
|
||
<emphasis bold="true" box="[1011,1020,1653,1676]" italics="true" pageId="10" pageNumber="567">c</emphasis>
|
||
) and frontal (
|
||
<emphasis bold="true" box="[1168,1179,1653,1676]" italics="true" pageId="10" pageNumber="567">e</emphasis>
|
||
) views; male: lateral (
|
||
<emphasis bold="true" box="[1409,1420,1653,1676]" italics="true" pageId="10" pageNumber="567">b</emphasis>
|
||
), dorsal (
|
||
<emphasis bold="true" box="[229,240,1682,1705]" italics="true" pageId="10" pageNumber="567">d</emphasis>
|
||
) and frontal (
|
||
<emphasis bold="true" box="[378,385,1682,1705]" italics="true" pageId="10" pageNumber="567">f</emphasis>
|
||
) views. Note that the petiole in queen (
|
||
<emphasis bold="true" box="[786,797,1682,1705]" italics="true" pageId="10" pageNumber="567">a</emphasis>
|
||
) looks wider and blunter due to being bent to the right site (
|
||
<emphasis bold="true" box="[1411,1420,1682,1705]" italics="true" pageId="10" pageNumber="567">c</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="12" lastPageNumber="569" pageId="10" pageNumber="567" type="description">
|
||
<paragraph blockId="10.[98,402,1758,1786]" box="[98,402,1758,1786]" pageId="10" pageNumber="567">
|
||
<heading bold="true" box="[98,402,1758,1786]" fontSize="11" level="3" pageId="10" pageNumber="567" reason="0">
|
||
<emphasis bold="true" box="[98,402,1758,1786]" pageId="10" pageNumber="567">Description of worker</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="10.[98,770,1809,1930]" lastBlockId="11.[98,770,180,1356]" lastPageId="11" lastPageNumber="568" pageId="10" pageNumber="567">
|
||
<emphasis bold="true" box="[98,262,1809,1833]" italics="true" pageId="10" pageNumber="567">Morphometrics</emphasis>
|
||
. Non type ‘HJ.3.1’ (
|
||
<specimenCount box="[502,638,1810,1834]" pageId="10" pageNumber="567" type="generic" typeStatus="syntypes">six syntypes</specimenCount>
|
||
and further five non-types workers, N.A. = measure not available hereafter; total measured
|
||
<emphasis bold="true" box="[411,429,1874,1897]" italics="true" pageId="10" pageNumber="567">N</emphasis>
|
||
= 11, min.– max): HL 1.08 (0.87–1.25), HW 1.14 (0.88–1.24), IOD 0.87 (0.67–0.92), CL 0.16 (0.15–0.19), CW 0.49 (0.34–0.54),
|
||
<collectionCode box="[1384,1420,1775,1798]" country="France" name="Musee de Lectoure" pageId="10" pageNumber="567">ML</collectionCode>
|
||
0.45 (0.33–0.47), EL 0.31 (0.25–0.36), EW 0.22 (0.16–0.26), FCDp 0.38 (0.29–0.44), FCDa 0.28 (0.21–0.29), SL 0.99 (0.70–1.08), SWmax 0.13 (0.11–0.17), SWmin 0.06 (0.05–0.09), WL 1.62 (1.24–1.76),
|
||
<collectionCode box="[1236,1274,1903,1926]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="567">PW</collectionCode>
|
||
0.79 (0.61–0.88), HFL 1.03 (0.92–1.08), HFT 0.24 (0.20–0.25), HTL N.A. (0.82–1.07), PetW 0.40 (0.32–0.47), PetL 0.22 (0.15–0.24),
|
||
<collectionCode box="[98,153,244,268]" pageId="11" pageNumber="568">FCDI</collectionCode>
|
||
134 (126–165), CI 105 (98–105), SI 87 (80–93), MI 41 (32–42).
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,180,1356]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,187,307,331]" italics="true" pageId="11" pageNumber="568">Head</emphasis>
|
||
. General morphology as for the genus (see above). Frontal carinae approaching one another less anteriorly (
|
||
<collectionCode box="[104,162,372,396]" pageId="11" pageNumber="568">FCDI</collectionCode>
|
||
= 134) and with relatively straight curvature, slightly concave near antennal sockets (
|
||
<figureCitation box="[459,536,403,428]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[523,536,403,427]" italics="true" pageId="11" pageNumber="568">d</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,180,1356]" box="[130,770,436,460]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,411,436,460]" italics="true" pageId="11" pageNumber="568">Mesosoma and metasoma</emphasis>
|
||
. As for the genus (see above).
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,180,1356]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,210,468,492]" italics="true" pageId="11" pageNumber="568">Pilosity</emphasis>
|
||
. Numerous tiny appressed hairs (approximately twice the length of 1 ommatidium or shorter) over scape, head and most of the thorax and abdomen; relatively long erect hairs (length ≥
|
||
<collectionCode box="[339,378,564,588]" pageId="11" pageNumber="568">SW</collectionCode>
|
||
) present only on clypeus, head and ventral abdomen and one pair of hairs present on frons in the middle above the eyes (
|
||
<figureCitation box="[442,525,628,652]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[512,525,628,651]" italics="true" pageId="11" pageNumber="568">a</emphasis>
|
||
</figureCitation>
|
||
); these longer hairs missing on genae and the occipital corners of the head including region above the eyes (in frontal view none of the longer hairs surpass the head margin); scape also without erect hairs except a tuft of ~8 setae growing from the apical tip (
|
||
<figureCitation box="[217,291,787,812]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[279,291,787,811]" italics="true" pageId="11" pageNumber="568">b</emphasis>
|
||
</figureCitation>
|
||
). No long erect hairs at the apex of petiole.
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,180,1356]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,232,820,844]" italics="true" pageId="11" pageNumber="568">Sculpture</emphasis>
|
||
. Fine punctation on dorsal surface of the head, in particular in the area between frontal carinae and above the clypeus (
|
||
<figureCitation box="[361,463,883,908]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[424,436,883,907]" italics="true" pageId="11" pageNumber="568">b</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[450,463,883,907]" italics="true" pageId="11" pageNumber="568">d</emphasis>
|
||
</figureCitation>
|
||
) and on mesosoma dorsum and lateral sites of thorax above metathoracic spiracle and dorsolaterally on propodeum (
|
||
<figureCitation box="[463,547,948,972]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[534,547,948,971]" italics="true" pageId="11" pageNumber="568">a</emphasis>
|
||
</figureCitation>
|
||
); this punctuation changes to the sculpturing rather forming a cell structure than individual pits posteriorly towards the occipital region; soft lineation on lateral sites of mesosoma, below the metathoracic spiracle and on propodeum between propodeal spiracle and hind coxae; same cuticular lineation also present on frontal coxae, petiole and anteriorly on the head near clypeus, but here much softer than on thorax (
|
||
<figureCitation box="[629,733,1171,1196]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[693,705,1171,1195]" italics="true" pageId="11" pageNumber="568">b</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[722,733,1172,1195]" italics="true" pageId="11" pageNumber="568">c</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,180,1356]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,260,1204,1228]" italics="true" pageId="11" pageNumber="568">Colouration</emphasis>
|
||
. Black head, thorax and petiole; dark brown scape except at the apex where light brown; brown abdomen and femurs; tarsi, funiculus and palps light brown to yellow; femurs brown in the middle but lighter near the joints (
|
||
<figureCitation box="[107,211,1332,1356]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[171,184,1332,1355]" italics="true" pageId="11" pageNumber="568">a</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[200,211,1332,1355]" italics="true" pageId="11" pageNumber="568">c</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,387,1409,1437]" box="[98,387,1409,1437]" pageId="11" pageNumber="568">
|
||
<heading bold="true" box="[98,387,1409,1437]" fontSize="11" level="3" pageId="11" pageNumber="568" reason="0">
|
||
<emphasis bold="true" box="[98,387,1409,1437]" pageId="11" pageNumber="568">Description of queen</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,1459,1932]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[98,262,1459,1483]" italics="true" pageId="11" pageNumber="568">Morphometrics</emphasis>
|
||
.
|
||
<typeStatus box="[294,384,1460,1484]" pageId="11" pageNumber="568">Syntype</typeStatus>
|
||
‘focol2566’: HL 1.43, HW 1.45, IOD 1.08, CL 0.30, CW 0.60,
|
||
<collectionCode box="[444,480,1492,1515]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="568">ML</collectionCode>
|
||
0.67, EL 0.53, EW 0.35, FCDp 0.47, FCDa 0.33, SL 1.17, SWmax 0.16, SWmin 0.08, WL 2.56,
|
||
<collectionCode box="[214,252,1556,1579]" country="Thailand" name="Paleontological Collections" pageId="11" pageNumber="568">PW</collectionCode>
|
||
1.14, HFL 1.08, HFT 0.36, HTL 1.34, PetW 0.58, PetL 0.28,
|
||
<collectionCode box="[286,342,1588,1612]" pageId="11" pageNumber="568">FCDI</collectionCode>
|
||
142, CI 101, SI 81, MI 47.
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,1459,1932]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,187,1620,1644]" italics="true" pageId="11" pageNumber="568">Head</emphasis>
|
||
(
|
||
<figureCitation box="[205,279,1620,1644]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[268,279,1620,1643]" italics="true" pageId="11" pageNumber="568">e</emphasis>
|
||
</figureCitation>
|
||
). Eyes almost twice as large as in worker. Frontal carinae approaching each other anteriorly above the posterior margin of clypeus relatively less when compared to queen of
|
||
<taxonomicName authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[298,424,1716,1740]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="568" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis bold="true" box="[298,424,1716,1740]" italics="true" pageId="11" pageNumber="568">O. papuana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode box="[438,496,1716,1740]" pageId="11" pageNumber="568">FCDI</collectionCode>
|
||
<160). Mandibles with four teeth; apical and subapical teeth approximately twice as long as and sharper than the other three teeth. Three ocelli well developed.
|
||
</paragraph>
|
||
<paragraph blockId="11.[98,770,1459,1932]" lastBlockId="11.[818,1490,180,876]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[130,244,1844,1867]" italics="true" pageId="11" pageNumber="568">Mesosoma</emphasis>
|
||
(
|
||
<figureCitation box="[264,373,1844,1868]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[331,344,1844,1867]" italics="true" pageId="11" pageNumber="568">a</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[362,373,1844,1867]" italics="true" pageId="11" pageNumber="568">c</emphasis>
|
||
</figureCitation>
|
||
). In lateral view, thorax dorsum flat at mesothorax, otherwise evenly rounded with convex prothoracic and propodeal angle. Overall morphology of thoracic segments and wings as in
|
||
<taxonomicName baseAuthorityName="Eisner and Wilson" baseAuthorityYear="1952" box="[1254,1389,180,204]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="568" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[1254,1389,180,204]" italics="true" pageId="11" pageNumber="568">Camponotus</emphasis>
|
||
</taxonomicName>
|
||
queens, except apical part of the forewing being relatively broader with evenly convex margins laterally towards the tip.
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,180,876]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[850,962,276,299]" italics="true" pageId="11" pageNumber="568">Metasoma</emphasis>
|
||
. Petiole similar to worker but relatively thicker and with less sharp apex. Abdomen as in workers but larger.
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,180,876]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[850,930,372,396]" italics="true" pageId="11" pageNumber="568">Pilosity</emphasis>
|
||
. Numerous tiny appressed (≥1/2 length of ocellus) hairs over scape; erect relatively long hairs (length>1 ocellus) present only on clypeus, ventral head, near ocelli, on mesothorax dorsum and on 3rd to 5th abdominal tergites (
|
||
<figureCitation box="[920,999,500,524]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[986,999,500,523]" italics="true" pageId="11" pageNumber="568">a</emphasis>
|
||
</figureCitation>
|
||
); these longer hairs missing on genae and the occipital corners of the head including region above the eyes (in frontal view no longer hairs surpass the head margins); scape also lacks erect hairs except five to six setae at the apical tip (
|
||
<figureCitation box="[1025,1103,628,652]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[1092,1103,628,651]" italics="true" pageId="11" pageNumber="568">e</emphasis>
|
||
</figureCitation>
|
||
). No long erect hairs on apex of petiole.
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,180,876]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[850,952,691,715]" italics="true" pageId="11" pageNumber="568">Sculpture</emphasis>
|
||
. Fine punctation on the dorsal surface of the head, and on mesosoma dorsum and propodeum; soft lineation on lateral sites of mesosoma, similar to workers.
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,180,876]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[850,980,787,811]" italics="true" pageId="11" pageNumber="568">Colouration</emphasis>
|
||
. Brown head, thorax, scape (except the tip that is lighter) and petiole; light brown abdomen; legs and funiculus yellow (
|
||
<figureCitation box="[1022,1126,852,876]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[1086,1099,852,875]" italics="true" pageId="11" pageNumber="568">a</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1115,1126,852,875]" italics="true" pageId="11" pageNumber="568">c</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,901,929,957]" box="[818,901,929,957]" pageId="11" pageNumber="568">
|
||
<heading bold="true" box="[818,901,929,957]" fontSize="11" level="3" pageId="11" pageNumber="568" reason="0">
|
||
<emphasis bold="true" box="[818,901,929,957]" pageId="11" pageNumber="568">Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,980,1069]" pageId="11" pageNumber="568">The generally lighter colouring may also reflect either the old age of the museum specimen (>100 years) or a relatively young queen collected from inside nest (alate).</paragraph>
|
||
<paragraph blockId="11.[818,1091,1121,1149]" box="[818,1091,1121,1149]" pageId="11" pageNumber="568">
|
||
<heading bold="true" box="[818,1091,1121,1149]" fontSize="11" level="3" pageId="11" pageNumber="568" reason="0">
|
||
<emphasis bold="true" box="[818,1091,1121,1149]" pageId="11" pageNumber="568">Description of male</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,1171,1932]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[818,982,1171,1195]" italics="true" pageId="11" pageNumber="568">Morphometrics</emphasis>
|
||
.
|
||
<typeStatus box="[1014,1104,1172,1196]" pageId="11" pageNumber="568">Syntype</typeStatus>
|
||
‘CASENT0905180’: HL 0.90, HW 1.08, IOD 0.69, CL 0.17, CW 0.40,
|
||
<collectionCode box="[1282,1318,1204,1227]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="568">ML</collectionCode>
|
||
0.34, EL 0.46, EW 0.32, FCDp 0.32, FCDa 0.14, SL 0.80, SWmax 0.09, SWmin 0.07, WL 1.95,
|
||
<collectionCode box="[1110,1148,1268,1291]" country="Thailand" name="Paleontological Collections" pageId="11" pageNumber="568">PW</collectionCode>
|
||
1.08, HFL 1.31, HFT 0.18, HTL 1.30, PetW N.A., PetL N.A.,
|
||
<collectionCode box="[1162,1216,1300,1324]" pageId="11" pageNumber="568">FCDI</collectionCode>
|
||
229, CI 120, SI 74, MI 38.
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,1171,1932]" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[850,907,1332,1356]" italics="true" pageId="11" pageNumber="568">Head</emphasis>
|
||
(
|
||
<figureCitation box="[924,995,1332,1356]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[987,995,1332,1356]" italics="true" pageId="11" pageNumber="568">f</emphasis>
|
||
</figureCitation>
|
||
). Eyes largely surpass the head margins in frontal view, with EL ~1/2 of HL, placed at ~1/2 of HL posteriorly. Frontal carinae approaching relatively less one another anteriorly above the posterior margin of clypeus compared to those in male of
|
||
<taxonomicName authorityName="Klimeš & Drescher & Buchori & Hidayat & Nazarreta & Potocký & Rimandai & Scheu & Matos-Maraví" authorityYear="2022" box="[1184,1316,1460,1484]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="568" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis bold="true" box="[1184,1316,1460,1484]" italics="true" pageId="11" pageNumber="568">O. papuana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode box="[1336,1394,1460,1484]" pageId="11" pageNumber="568">FCDI</collectionCode>
|
||
= 229), although this variance is not as distinct as among workers and among queens of the two species. Mandibles blunt without teeth. Three ocelli well developed and relatively larger than in queens (diameter as SWmax). Antennae with 13 segments not broadened towards the tip of funiculus like in the workers and queen (see frontal image of another male from the
|
||
<typeStatus box="[986,1035,1684,1708]" pageId="11" pageNumber="568">type</typeStatus>
|
||
series ‘focol2564’ with intact antennae) (AntWeb, see https://www.antweb.org). Scape long, at half of the length passing the head margin in frontal view.
|
||
</paragraph>
|
||
<paragraph blockId="11.[818,1490,1171,1932]" lastBlockId="12.[98,770,180,748]" lastPageId="12" lastPageNumber="569" pageId="11" pageNumber="568">
|
||
<emphasis bold="true" box="[850,964,1780,1803]" italics="true" pageId="11" pageNumber="568">Mesosoma</emphasis>
|
||
. Thorax dorsum flat at mesoscutum, otherwise evenly rounded in lateral view with convex prothoracic and propodeal angle, except the mesoscutellar disc that is placed higher and of more convex shape than the other thoracic parts in lateral view (
|
||
<figureCitation box="[1193,1276,1908,1932]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="11" pageNumber="568">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[1264,1276,1908,1932]" italics="true" pageId="11" pageNumber="568">b</emphasis>
|
||
</figureCitation>
|
||
). Suture between pronotum and mesoscutum not visible in dorsal view due to highly convex and enlarged mesoscutum compared to pronotum (
|
||
<figureCitation box="[188,266,244,268]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="12" pageNumber="569">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[253,266,244,268]" italics="true" pageId="12" pageNumber="569">d</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,180,748]" pageId="12" pageNumber="569">
|
||
<emphasis bold="true" box="[130,242,276,299]" italics="true" pageId="12" pageNumber="569">Metasoma</emphasis>
|
||
. Petiole triangular but relatively thicker and blunter at the apex.
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,180,748]" pageId="12" pageNumber="569">
|
||
<emphasis bold="true" box="[130,210,340,364]" italics="true" pageId="12" pageNumber="569">Pilosity</emphasis>
|
||
. Tiny appressed hairs present on scape and thorax but less numerous over the head compared to queen and workers; long erect hairs (length = SWmax) present only on clypeus, genae, ventral head, near ocelli (one long seta et each) and on 3rd to 5th abdominal tergites (
|
||
<figureCitation box="[608,685,468,492]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="12" pageNumber="569">
|
||
Fig. 2
|
||
<emphasis bold="true" box="[673,685,468,492]" italics="true" pageId="12" pageNumber="569">b</emphasis>
|
||
</figureCitation>
|
||
); these longer hairs missing completely on scape, petiole and at the occipital corners of the head including region above the eyes (in frontal view, no long hairs surpass the head margin).
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,180,748]" pageId="12" pageNumber="569">
|
||
<emphasis bold="true" box="[130,232,595,619]" italics="true" pageId="12" pageNumber="569">Sculpture</emphasis>
|
||
. Fine punctation on dorsal surface of the head, and on mesosoma dorsum and propodeum.
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,180,748]" pageId="12" pageNumber="569">
|
||
<emphasis bold="true" box="[130,260,660,684]" italics="true" pageId="12" pageNumber="569">Colouration</emphasis>
|
||
. Light brown to yellowish body, except brown head, thorax dorsum and antennae. Mandibles and appendages pale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="569" type="discussion">
|
||
<paragraph blockId="12.[98,220,801,829]" box="[98,220,801,829]" pageId="12" pageNumber="569">
|
||
<heading bold="true" box="[98,220,801,829]" fontSize="11" level="3" pageId="12" pageNumber="569" reason="0">
|
||
<emphasis bold="true" box="[98,220,801,829]" pageId="12" pageNumber="569">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,852,1932]" pageId="12" pageNumber="569">
|
||
The specimens of
|
||
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[310,463,852,876]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="569" phylum="Arthropoda" rank="species" species="subclavata">
|
||
<emphasis bold="true" box="[310,463,852,876]" italics="true" pageId="12" pageNumber="569">O. subclavata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[540,648,852,876]" name="Indonesia" pageId="12" pageNumber="569">Indonesia</collectingCountry>
|
||
that were examined have the posterior parts of the frontal carinae bent slightly inwards and therefore have slightly smaller
|
||
<collectionCode box="[714,770,916,940]" pageId="12" pageNumber="569">FCDI</collectionCode>
|
||
(mean 133) than the
|
||
<typeStatus box="[339,388,949,973]" pageId="12" pageNumber="569">type</typeStatus>
|
||
specimens from
|
||
<collectingCountry box="[579,691,948,972]" name="Singapore" pageId="12" pageNumber="569">Singapore</collectingCountry>
|
||
(mean 155) (
|
||
<figureCitation box="[167,243,979,1004]" captionStart="Fig" captionStartId="6.[314,351,934,957]" captionTargetBox="[314,1274,185,904]" captionTargetId="figure-471@6.[313,1274,184,904]" captionTargetPageId="6" captionText="Fig. 1. Overbeckia subclavata, non-type. Worker from Indonesia (specimen HJ.3.1) displaying the lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d). Note hind leg tibia and tarsi are missing (damaged)." figureDoi="http://doi.org/10.5281/zenodo.6987299" httpUri="https://zenodo.org/record/6987299/files/figure.png" pageId="12" pageNumber="569">
|
||
Fig. 1
|
||
<emphasis bold="true" box="[230,243,979,1003]" italics="true" pageId="12" pageNumber="569">d</emphasis>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation box="[259,286,979,1004]" captionStart="Fig" captionStartId="7.[208,245,1815,1838]" captionTargetBox="[209,1380,185,1785]" captionTargetId="figure-2@7.[208,1381,184,1785]" captionTargetPageId="7" captionText="Fig. 2. Overbeckia subclavata syntypes. Alate queen (FOCOL2566; photographer: Ch. Klingenberg): lateral (a), dorsal (c) and frontal (e) views; male (CASENT0905180; photographer: W. Ericson): lateral (b), dorsal (d) and frontal (f) views; worker (CASENT0101191; photographer A. Nobile): lateral (g) and frontal (h) views. Photos retrieved from AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.6987301" httpUri="https://zenodo.org/record/6987301/files/figure.png" pageId="12" pageNumber="569">
|
||
2
|
||
<emphasis bold="true" box="[273,286,979,1003]" italics="true" pageId="12" pageNumber="569">h</emphasis>
|
||
</figureCitation>
|
||
). However, the difference is small and this could possibly be due to measurement error from photographs. All other morphological parameters and the general appearance of the specimens are consistent between the two regions, therefore we regard this to be only a variation within the species.
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,852,1932]" pageId="12" pageNumber="569">
|
||
The specimen from the
|
||
<collectingCountry box="[411,536,1172,1196]" name="Philippines" pageId="12" pageNumber="569">Philippines</collectingCountry>
|
||
has a damaged left upper side of the head and a whitish dirt attached to the carinae. Therefore, we did not consider this for morphological measurements. However, absence of the longer hairs on the scape and the thorax, and the finely punctate head with numerous minute recumbent hairs, clearly fit the general appearance of
|
||
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[264,415,1363,1388]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="569" phylum="Arthropoda" rank="species" species="subclavata">
|
||
<emphasis bold="true" box="[264,415,1363,1388]" italics="true" pageId="12" pageNumber="569">O. subclavata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="12.[98,770,852,1932]" lastBlockId="12.[818,1490,180,364]" pageId="12" pageNumber="569">
|
||
<bibRefCitation author="Viehmeyer H" box="[130,333,1396,1420]" pageId="12" pageNumber="569" pagination="108 - 168" refId="ref20034" refString="Viehmeyer H (1916) Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Archiv fur Naturgeschichte - A 81 (8), 108 - 168." type="journal article" year="1916">Viehmeyer (1916)</bibRefCitation>
|
||
reported that
|
||
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[493,642,1396,1420]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="569" phylum="Arthropoda" rank="species" species="subclavata">
|
||
<emphasis bold="true" box="[493,642,1396,1420]" italics="true" pageId="12" pageNumber="569">O. subclavata</emphasis>
|
||
</taxonomicName>
|
||
nested in a hollow bamboo stem and in a hollow branch of a mango tree, therefore there is uncertainty whether the
|
||
<typeStatus box="[683,769,1460,1484]" pageId="12" pageNumber="569">syntype</typeStatus>
|
||
series of the species (
|
||
<specimenCount box="[363,502,1492,1516]" pageId="12" pageNumber="569" type="worker">six workers</specimenCount>
|
||
,
|
||
<specimenCount box="[521,658,1492,1516]" pageId="12" pageNumber="569" type="male">three males</specimenCount>
|
||
and
|
||
<specimenCount pageId="12" pageNumber="569" type="queen">one queen</specimenCount>
|
||
, available from AntWeb) belongs to a single nest. Five vials (samples?) collected in 1913 and two collected in 1914 were reported, therefore several colonies from one or more sites in
|
||
<collectingCountry box="[287,399,1620,1644]" name="Singapore" pageId="12" pageNumber="569">Singapore</collectingCountry>
|
||
might have been pooled for the
|
||
<typeStatus box="[98,184,1652,1676]" pageId="12" pageNumber="569">syntype</typeStatus>
|
||
series (most likely two colonies from two stems, a mango and a bamboo). There is also uncertainty as to whether all specimens collected in 1913–1914 are documented online at AntWeb, therefore more material might be scattered in museums worldwide. Nevertheless, all specimens collected by H. Overbeck illustrated in the AntWeb match
|
||
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[180,335,1843,1868]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="569" phylum="Arthropoda" rank="species" species="subclavata">
|
||
<emphasis bold="true" box="[180,335,1843,1868]" italics="true" pageId="12" pageNumber="569">O. subclavata</emphasis>
|
||
</taxonomicName>
|
||
. The arboricolous nesting in hollow plant structures of (probably) dead tissues suggests that the species nests in tree canopies, but is rather a generalist; that is, not using living tissues or myrmecophytic plants, as do more specialised arboreal species (
|
||
<bibRefCitation author="Klimes P & McArthur A" pageId="12" pageNumber="569" pagination="141 - 158" refId="ref18399" refString="Klimes P, McArthur A (2014) Diversity and ecology of arboricolous ant communities of Camponotus (Hymenoptera: Formicidae) in a New Guinea rainforest with descriptions of four new species. Myrmecological News 20, 141 - 158." type="journal article" year="2014">Klimes and McArthur 2014</bibRefCitation>
|
||
). Specimens of
|
||
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1057,1210,244,268]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="569" phylum="Arthropoda" rank="species" species="subclavata">
|
||
<emphasis bold="true" box="[1057,1210,244,268]" italics="true" pageId="12" pageNumber="569">O. subclavata</emphasis>
|
||
</taxonomicName>
|
||
were probably collected by H. Overbeck in a human-managed area but collections from Sumatra suggest that the species utilises a variety of habitats (see Distribution and Discussion for further information).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |