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<document id="A53FE9015E6009F9CF4770AE27A3E75C" ID="10.11646/zootaxa.4175.1.2" ID-DOI="10.11646/zootaxa.4175.1.2" ID-GBIF-Dataset="887d93ae-26dc-4192-87d4-5396e561df0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="160280" ID-ZooBank="011B74BE-40C0-4606-9354-C637F83C3E43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1476253530258" checkinUser="plazi" docAuthor="Shattuck, Steve &amp; Cover, Stefan" docDate="2016" docId="03E5E90BFF812331FF3C9FBAFC88777D" docLanguage="en" docName="zootaxa.4175.1.2.pdf" docOrigin="Zootaxa 4175 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nomamyrmex esenbeckii Westwood 1842" docType="treatment" docVersion="14" lastPageNumber="19" masterDocId="FFDC9173FF882338FFAB9D09FFC47079" masterDocTitle="Taxonomy of some little-understood North American ants (Hymenoptera: Formicidae)" masterLastPageNumber="22" masterPageNumber="10" pageNumber="19" updateTime="1698389545208" updateUser="plazi">
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<mods:title id="7F05FFFFE28EFD7A9D305B16C3CCFEF9">Taxonomy of some little-understood North American ants (Hymenoptera: Formicidae)</mods:title>
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<paragraph id="8BF3581DFF812331FF3C9FBAFD5772B5" blockId="9.[151,1437,691,1796]" box="[151,659,691,716]" pageId="9" pageNumber="19">
<heading id="D0BBEF71FF812331FF3C9FBAFD5772B5" bold="true" box="[151,659,691,716]" fontSize="10" level="2" pageId="9" pageNumber="19" reason="2">
<taxonomicName id="4C4C239EFF812331FF3C9FBAFD5772B5" ID-CoL="47NPQ" authority="Westwood, 1842" authorityName="Westwood" authorityYear="1842" box="[151,659,691,716]" class="Insecta" family="Formicidae" genus="Nomamyrmex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="esenbeckii">
<emphasis id="B938840FFF812331FF3C9FBAFD5772B5" bold="true" box="[151,659,691,716]" pageId="9" pageNumber="19">
<emphasis id="B938840FFF812331FF3C9FBAFE7D72B5" bold="true" box="[151,441,691,716]" italics="true" pageId="9" pageNumber="19">Nomamyrmex esenbeckii</emphasis>
(
<bibRefCitation id="EFDD25ECFF812331FE6C9FBAFD4F72B5" author="Westwood" box="[455,651,691,716]" pageId="9" pageNumber="19" refString="Westwood, J. O. (1842) Monograph of the hymenopterous group, Dorylides. In: Westwood, J. O. Arcana entomologica; or illustrations of new, rare, and interesting insects. Volume 1, No. 5. W. Smith, London, pp. 73 - 80." type="journal article" year="1842">Westwood, 1842</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3560B96FF812331FF6C9FDFFAC573B1" pageId="9" pageNumber="19" type="reference_group">
<paragraph id="8BF3581DFF812331FF6C9FDFFEB3734E" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
<materialsCitation id="3B245240FF812331FF6C9FDFFEB3734E" ID-GBIF-Occurrence="1316541008" collectingDate="1828-09-10" collectorName="W. Burchell &amp; Oxford University Museum &amp; Natural History" country="Brazil" latitude="-29.783333" location="Vendinha" longLatPrecision="1225" longitude="51.55" pageId="9" pageNumber="19" specimenCount="1" typeStatus="holotype">
<treatmentCitationGroup id="AB5C7F33FF812331FF6C9FDFFEB3734E" infered="true" pageId="9" pageNumber="19">
<taxonomicName id="4C4C239EFF812331FF6C9FDFFD507296" authority="Westwood, 1842: 75" authorityName="Westwood" authorityPageNumber="75" authorityYear="1842" box="[199,660,726,751]" class="Insecta" family="Formicidae" genus="Labidus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="esenbeckii">
<emphasis id="B938840FFF812331FF6C9FDFFE5B7296" box="[199,415,726,751]" italics="true" pageId="9" pageNumber="19">Labidus esenbeckii</emphasis>
<treatmentCitation id="0AED7E0CFF812331FE0C9FDFFD507296" author="Westwood" box="[423,660,726,751]" page="75" pageId="9" pageNumber="19" year="1842">
<bibRefCitation id="EFDD25ECFF812331FE0C9FDFFDAD7296" author="Westwood" box="[423,617,726,751]" pageId="9" pageNumber="19" refString="Westwood, J. O. (1842) Monograph of the hymenopterous group, Dorylides. In: Westwood, J. O. Arcana entomologica; or illustrations of new, rare, and interesting insects. Volume 1, No. 5. W. Smith, London, pp. 73 - 80." type="journal article" year="1842">Westwood, 1842</bibRefCitation>
: 75
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54F7E6BFFF812331FD0A9FDFFCCD7296" box="[673,777,726,751]" pageId="9" pageNumber="19" type="holotype">Holotype</typeStatus>
male,
<location id="8E930EC6FF812331FCF29FDFFC077296" LSID="urn:lsid:plazi:treatment:03E5E90BFF812331FF3C9FBAFC88777D:8E930EC6FF812331FCF29FDFFC077296" box="[857,963,726,751]" country="Brazil" latitude="-29.783333" longLatPrecision="1225" longitude="51.55" name="Vendinha" pageId="9" pageNumber="19">Vendinha</location>
[
<geoCoordinate id="EE783EDAFF812331FC789FDFFBEA7296" box="[979,1070,726,751]" direction="south" orientation="latitude" pageId="9" pageNumber="19" precision="925" value="-29.783333">29°47S</geoCoordinate>
<geoCoordinate id="EE783EDAFF812331FB9C9FDFFB527296" box="[1079,1174,726,751]" direction="east" orientation="longitude" pageId="9" pageNumber="19" precision="925" value="51.55">51°33E</geoCoordinate>
],
<collectingCountry id="F35B188DFF812331FB009FDFFB367296" box="[1195,1266,726,751]" name="Brazil" pageId="9" pageNumber="19">Brazil</collectingCountry>
,
<date id="FFF27EDDFF812331FB549FDEFF0B736A" pageId="9" pageNumber="19" value="1828-09-10">
<collectingDate id="EFB68735FF812331FB549FDEFF0B736A" pageId="9" pageNumber="19" value="1828-09-10">10 September 1828</collectingDate>
</date>
(
<collectorName id="26B93DCBFF812331FF419FF2FE44736D" box="[234,384,763,788]" pageId="9" pageNumber="19">W. Burchell</collectorName>
) (
<collectorName id="26B93DCBFF812331FE089FF2FD32736A" box="[419,758,763,788]" pageId="9" pageNumber="19">Oxford University Museum</collectorName>
of
<collectorName id="26B93DCBFF812331FC9F9FF2FC36736D" box="[820,1010,763,788]" pageId="9" pageNumber="19">Natural History</collectorName>
) [https://www.antweb.org/specimen/ CASENT0901959].
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="8BF3581DFF812331FF6C9E4DFA8573F9" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
<materialsCitation id="3B245240FF812331FF6C9E4DFA8573F9" ID-GBIF-Occurrence="1316541017" country="Mexico" latitude="18.916666" location="New" longLatPrecision="1273" longitude="-99.21667" pageId="9" pageNumber="19" specimenCount="1" typeStatus="holotype">
<treatmentCitationGroup id="AB5C7F33FF812331FF6C9E4DFA8573F9" infered="true" pageId="9" pageNumber="19">
<taxonomicName id="4C4C239EFF812331FF6C9E4DFD357325" authority="Santschi, 1929: 415" authorityName="Santschi" authorityPageNumber="415" authorityYear="1929" box="[199,753,835,860]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="crassicorne" subSpecies="mordax">
<emphasis id="B938840FFF812331FF6C9E4DFE3A7325" box="[199,510,835,860]" italics="true" pageId="9" pageNumber="19">Eciton crassicorne mordax</emphasis>
<treatmentCitation id="0AED7E0CFF812331FDA39E4AFD357325" author="Santschi" box="[520,753,835,860]" page="415" pageId="9" pageNumber="19" year="1929">
<bibRefCitation id="EFDD25ECFF812331FDA39E4AFD717322" author="Santschi" box="[520,693,835,860]" pageId="9" pageNumber="19" refString="Santschi, F. (1929) Sur quelques nouvelles fourmis du Bresil (Hym. Form.). Deutsche Entomologische Zeitschrift, 1928, 414 - 416. http: // dx. doi. org / 10.1002 / mmnd. 192919280507" type="journal article" year="1929">Santschi, 1929</bibRefCitation>
: 415
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54F7E6BFFF812331FCAB9E4AFCAC7325" box="[768,872,835,860]" pageId="9" pageNumber="19" type="holotype">Holotype</typeStatus>
worker, Cernavaca [
<geoCoordinate id="EE783EDAFF812331FBF79E4AFB797325" box="[1116,1213,835,860]" direction="north" orientation="latitude" pageId="9" pageNumber="19" precision="925" value="18.916666">18°55N</geoCoordinate>
<geoCoordinate id="EE783EDAFF812331FB6C9E4AFAF47325" box="[1223,1328,835,860]" direction="west" orientation="longitude" pageId="9" pageNumber="19" precision="925" value="-99.21667">99°13W</geoCoordinate>
],
<collectingCountry id="F35B188DFF812331FAEC9E4AFA587325" box="[1351,1436,835,860]" name="Mexico" pageId="9" pageNumber="19">Mexico</collectingCountry>
(Naturhistorisches Museum Basel) [https://www.antweb.org/specimen/CASENT0911413].
<location id="8E930EC6FF812331FB3C9E61FB087306" LSID="urn:lsid:plazi:treatment:03E5E90BFF812331FF3C9FBAFC88777D:8E930EC6FF812331FB3C9E61FB087306" box="[1175,1228,872,895]" country="Mexico" latitude="18.916666" longLatPrecision="1273" longitude="-99.21667" name="New" pageId="9" pageNumber="19">New</location>
synonym.
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="8BF3581DFF812331FF6C9E85FAC573B1" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
<materialsCitation id="3B245240FF812331FF6C9E85FAC573B1" ID-GBIF-Occurrence="1316540997" collectorName="C. Schaeffer &amp; Brooklyn Museum" latitude="25.9" location="New" longLatPrecision="1244" longitude="-97.5" municipality="June" pageId="9" pageNumber="19" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<treatmentCitationGroup id="AB5C7F33FF812331FF6C9E85FAC573B1" infered="true" pageId="9" pageNumber="19">
<taxonomicName id="4C4C239EFF812331FF6C9E85FD2773DD" authority="Santschi, 1920: 366" authorityName="Santschi" authorityPageNumber="366" authorityYear="1920" box="[199,739,907,932]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="esenbecki" subSpecies="wilsoni">
<emphasis id="B938840FFF812331FF6C9E85FE2C73DD" box="[199,488,907,932]" italics="true" pageId="9" pageNumber="19">Eciton esenbecki wilsoni</emphasis>
<treatmentCitation id="0AED7E0CFF812331FE5E9E82FD2773DD" author="Santschi" box="[501,739,907,932]" page="366" pageId="9" pageNumber="19" year="1920">
<bibRefCitation id="EFDD25ECFF812331FE5E9E82FD6073DA" author="Santschi" box="[501,676,907,932]" pageId="9" pageNumber="19" refString="Santschi, F. (1920) Formicides africains et americains nouveaux. Annales de la Societe Entomologique de France, 88, 361 - 390." type="journal article" year="1920">Santschi, 1920</bibRefCitation>
: 366
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54F7E6BFFF812331FD589E82FC9F73DD" box="[755,859,907,932]" pageId="9" pageNumber="19" type="holotype">Holotype</typeStatus>
male, Las Borragas, near Brownsville [
<geoCoordinate id="EE783EDAFF812331FA979E82FA5873DD" box="[1340,1436,907,932]" direction="north" orientation="latitude" pageId="9" pageNumber="19" precision="925" value="25.9">25°54N</geoCoordinate>
<geoCoordinate id="EE783EDAFF812331FF3C9EA7FEC473BE" box="[151,256,942,967]" direction="west" orientation="longitude" pageId="9" pageNumber="19" precision="925" value="-97.5">97°30W</geoCoordinate>
],
<collectingRegion id="498896FFFF812331FEB99EB9FE9273BE" box="[274,342,944,967]" country="United States of America" name="Texas" pageId="9" pageNumber="19">Texas</collectingRegion>
,
<collectingMunicipality id="6B97C267FF812331FECA9EB9FE5173BE" box="[353,405,944,967]" pageId="9" pageNumber="19">June</collectingMunicipality>
[year uncertain] (
<collectorName id="26B93DCBFF812331FDF69EA6FD2F73BE" box="[605,747,942,967]" pageId="9" pageNumber="19">C. Schaeffer</collectorName>
) (
<collectorName id="26B93DCBFF812331FCAA9EA7FC1C73BE" box="[769,984,942,967]" pageId="9" pageNumber="19">Brooklyn Museum</collectorName>
, not seen).
<location id="8E930EC6FF812331FBFC9EB9FB4873BE" LSID="urn:lsid:plazi:treatment:03E5E90BFF812331FF3C9FBAFC88777D:8E930EC6FF812331FBFC9EB9FB4873BE" box="[1111,1164,944,967]" latitude="25.9" longLatPrecision="1244" longitude="-97.5" municipality="June" name="New" pageId="9" pageNumber="19" stateProvince="Texas">New</location>
synonym.
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3560B96FF812331FF6C9EDAFCAD74BA" pageId="9" pageNumber="19" type="discussion">
<paragraph id="8BF3581DFF812331FF6C9EDAFCAD74BA" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
As noted by
<bibRefCitation id="EFDD25ECFF812331FEFF9EDAFDC67392" author="Watkins" box="[340,514,979,1004]" pageId="9" pageNumber="19" refString="Watkins, J. F. II. (1977) The species and subspecies of Nomamyrmex (Dorylinae: Formicidae). Journal of the Kansas Entomological Society, 50, 203 - 214." type="journal article" year="1977">Watkins (1977)</bibRefCitation>
,
<taxonomicName id="4C4C239EFF812331FDA69EDDFCE77395" box="[525,803,979,1004]" class="Insecta" family="Formicidae" genus="Nomamymrex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="esenbeckii">
<emphasis id="B938840FFF812331FDA69EDDFCE77395" box="[525,803,979,1004]" italics="true" pageId="9" pageNumber="19">Nomamymrex esenbeckii</emphasis>
</taxonomicName>
has accumulated a large number of names to describe its various forms. The number of these names considered as valid has gradually decreased as additional material has revealed intermediate morphologies, diverse combinations of character states and distinct geographic patterns. As a result we are currently down to just a single species with two recognized subspecies,
<taxonomicName id="4C4C239EFF812331FBDF9949FB29742E" box="[1140,1261,1086,1111]" class="Insecta" family="Formicidae" genus="Nomamymrex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="mordax">
<emphasis id="B938840FFF812331FBDF9949FB29742E" box="[1140,1261,1086,1111]" italics="true" pageId="9" pageNumber="19">N. mordax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C239EFF812331FA839949FA58742E" box="[1320,1436,1086,1111]" class="Insecta" family="Formicidae" genus="Nomamymrex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B938840FFF812331FA839949FA58742E" box="[1320,1436,1086,1111]" italics="true" pageId="9" pageNumber="19">N. wilsoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDD25ECFF812331FF35996AFE807402" author="Watkins" box="[158,324,1123,1148]" pageId="9" pageNumber="19" refString="Watkins, J. F. II. (1977) The species and subspecies of Nomamyrmex (Dorylinae: Formicidae). Journal of the Kansas Entomological Society, 50, 203 - 214." type="journal article" year="1977">Watkins, 1977</bibRefCitation>
;
<bibRefCitation id="EFDD25ECFF812331FEE4996AFE147402" author="Wild" box="[335,464,1123,1148]" pageId="9" pageNumber="19" refString="Wild, A. L. (2007) A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa, 1622, 1 - 55." type="journal article" year="2007">Wild, 2007</bibRefCitation>
). However, even this relatively simple system has proven problematic. In a detailed but unpublished analysis, Roy and Gordon Snelling (http://www.armyants.org/armyants/genusaccounts/
<taxonomicName id="4C4C239EFF812331FB5F998FFA5074E6" box="[1268,1428,1158,1183]" class="Insecta" family="Formicidae" genus="Nomamyrmex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="genus">nomamyrmex</taxonomicName>
/ noma.html, accessed
<date id="FFF27EDDFF812331FE2C99A2FDDA74BA" box="[391,542,1195,1219]" pageId="9" pageNumber="19" value="2014-06-19">19 June 2014</date>
) provide the following notes:
</paragraph>
</subSubSection>
<subSubSection id="C3560B96FF812331FF6C99C6FC8276C5" pageId="9" pageNumber="19" type="description">
<paragraph id="8BF3581DFF812331FF6C99C6FE99760A" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
“It is true that the morphological features on which the several subspecies of
<taxonomicName id="4C4C239EFF812331FBF699D9FB33749E" box="[1117,1271,1230,1255]" class="Insecta" family="Formicidae" genus="Nomamymrex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="esenbeckii">
<emphasis id="B938840FFF812331FBF699D9FB33749E" box="[1117,1271,1230,1255]" italics="true" pageId="9" pageNumber="19">N. esenbeckii</emphasis>
</taxonomicName>
are based are generally representative of their respective populations. Equally true, however, is that many specimens within these populations do not possess the required characteristics of that named population. Samples from areas of sympatry are, as may be expected, intermediate between the respective phenotypes. In the worker caste, there is a north/south trend that is continuous. In northern specimens, the posterior margin of the mesonotum, in dorsal view, is definitely concave, and the longitudinal rugules of the propodeal dorsum are short and weak. Proceeding into
<collectingRegion id="498896FFFF812331FAE1988AFA5875E5" box="[1354,1436,1411,1436]" country="Paraguay" name="Central" pageId="9" pageNumber="19">Central</collectingRegion>
America and through South America, the margin of the mesonotum becomes straight or even convex; the propodeal rugulae strengthen and run the entire length of the dorsal face. Similar trends are evident in the male structures employed by
<bibRefCitation id="EFDD25ECFF812331FE0098E7FD9E767E" author="Watkins" box="[427,602,1518,1543]" pageId="9" pageNumber="19" refString="Watkins, J. F. II. (1977) The species and subspecies of Nomamyrmex (Dorylinae: Formicidae). Journal of the Kansas Entomological Society, 50, 203 - 214." type="journal article" year="1977">Watkins (1977)</bibRefCitation>
in segregating
<taxonomicName id="4C4C239EFF812331FCA598F9FC61767E" box="[782,933,1518,1543]" class="Insecta" family="Formicidae" genus="Nomamymrex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="esenbeckii">
<emphasis id="B938840FFF812331FCA598F9FC61767E" box="[782,933,1518,1543]" italics="true" pageId="9" pageNumber="19">N. esenbeckii</emphasis>
</taxonomicName>
into four subspecies. In fact, differences in genitalic structures may be greater within any given population than those that separate purported subspecies. Under the circumstances, there would appear to be little justification for these subspecies and we propose to reduce all to synonymy.”
</paragraph>
<paragraph id="8BF3581DFF812331FF6C9B77FC8276C5" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
<bibRefCitation id="EFDD25ECFF812331FF6C9B77FE9676EE" author="Wild" box="[199,338,1662,1687]" pageId="9" pageNumber="19" refString="Wild, A. L. (2007) A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa, 1622, 1 - 55." type="journal article" year="2007">Wild (2007)</bibRefCitation>
accepted these recommendations when he synonymized
<taxonomicName id="4C4C239EFF812331FC4E9B89FB4876EE" box="[997,1164,1664,1687]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="crassicorne">
<emphasis id="B938840FFF812331FC4E9B89FB4876EE" box="[997,1164,1664,1687]" italics="true" pageId="9" pageNumber="19">N. crassicorne</emphasis>
</taxonomicName>
and we now apply this careful analysis by the Snellings to the remaining subspecies.
</paragraph>
</subSubSection>
<subSubSection id="C3560B96FF812331FF6C9BC1FC88777D" pageId="9" pageNumber="19" type="distribution">
<paragraph id="8BF3581DFF812331FF6C9BC1FC88777D" blockId="9.[151,1437,691,1796]" pageId="9" pageNumber="19">
<materialsCitation id="3B245240FF812331FF6C9BC1FC8C777D" ID-GBIF-Occurrence="1316541018" country="Argentina and Paraguay" location="southern Texas" pageId="9" pageNumber="19" specimenCount="1" stateProvince="Central">
<taxonomicName id="4C4C239EFF812331FF6C9BC1FE2576A6" box="[199,481,1734,1759]" class="Insecta" family="Formicidae" genus="Nomamymrex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="esenbeckii">
<emphasis id="B938840FFF812331FF6C9BC1FE2576A6" box="[199,481,1734,1759]" italics="true" pageId="9" pageNumber="19">Nomamymrex esenbeckii</emphasis>
</taxonomicName>
is exceptionally widespread, occurring from
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, United States, south through
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America to northern
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</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>