2217 lines
329 KiB
XML
2217 lines
329 KiB
XML
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<mods:title id="3675002F44E52D0D4E23082CFD2130A7">Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)</mods:title>
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<mods:namePart id="FA89F7D44B512D50C2ED603EBA1A890F">Kerr, Isaac A. R.</mods:namePart>
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<mods:namePart id="539DD813739CC83B956393D1E5AD7238">Van Zoelen, Jacob D.</mods:namePart>
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<mods:namePart id="8ADBD08836CBAC0D9AAE13C1DAA8DB69">Worthy, Trevor H.</mods:namePart>
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<mods:namePart id="8D8AAF36C2E3227AB41DE87501AA0127">Prideaux, Gavin J.</mods:namePart>
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<mods:title id="4F5714D025199E1B4B66176C4822DD5A">Megataxa</mods:title>
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<treatment id="03E587FDFF44D58FFF00772CFBB5F7E8" ID-DOI="http://doi.org/10.5281/zenodo.10993789" ID-Zenodo-Dep="10993789" LSID="urn:lsid:plazi:treatment:03E587FDFF44D58FFF00772CFBB5F7E8" httpUri="http://treatment.plazi.org/id/03E587FDFF44D58FFF00772CFBB5F7E8" lastPageId="186" lastPageNumber="187" pageId="168" pageNumber="169">
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<taxonomicName id="4C4C4D68FF44D59DFF00772CFD92FC69" authority="Plane, 1967" authorityName="Plane" authorityYear="1967" box="[136,578,874,901]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="otibandus">
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<emphasis id="B938EAF9FF44D59DFF00772CFD92FC69" bold="true" box="[136,578,874,901]" pageId="168" pageNumber="169">
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<emphasis id="B938EAF9FF44D59DFF00772CFE7BFC69" bold="true" box="[136,427,874,901]" italics="true" pageId="168" pageNumber="169">Protemnodon otibandus</emphasis>
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<bibRefCitation id="EFDD4B1AFF44D59DFE3A772DFD92FC69" author="Plane, M. D." box="[434,578,875,901]" pageId="168" pageNumber="169" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane, 1967</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3566560FF44D59DFF0077E8FCD5FAE8" pageId="168" pageNumber="169" type="reference_group">
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<paragraph id="8BF336EBFF44D59DFF0077E8FEA3FBE8" blockId="168.[136,779,942,1285]" pageId="168" pageNumber="169">
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<taxonomicName id="4C4C4D68FF44D59DFF0077E8FE3CFC28" authority="Plane, 1967" authorityName="Plane" authorityYear="1967" box="[136,492,942,964]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF44D59DFF0077E8FEA0FC2F" box="[136,368,942,963]" italics="true" pageId="168" pageNumber="169">Protemnodon otibandus</emphasis>
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<bibRefCitation id="EFDD4B1AFF44D59DFEFE77E8FE3CFC28" author="Plane, M. D." box="[374,492,942,964]" pageId="168" pageNumber="169" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane, 1967</bibRefCitation>
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</taxonomicName>
|
||
:
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||
<emphasis id="B938EAF9FF44D59DFE7E77E8FE9CFC08" italics="true" pageId="168" pageNumber="169">Bull. Bur. Min. Res., Geo. & Geophys., Aus.</emphasis>
|
||
, 86, pp. 26–44, figs 5–10. See also:
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFD147788FEE3FBE8" author="Mahoney, J. A. & Ride, W. D. L." pageId="168" pageNumber="169" pagination="1 - 250" refId="ref151155" refString="Mahoney, J. A. & Ride, W. D. L. (1975) Index to the genera and species of fossil Mammalia described from Australia and New Guinea between 1838 and 1968 (including citations of type species and primary type specimens). Special Publication of the Western Australian Museum, 6, 1 - 250." type="journal article" year="1975">Mahoney& Ride (1975)</bibRefCitation>
|
||
, p. 23.
|
||
</paragraph>
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<paragraph id="8BF336EBFF44D59DFF007048FE2BFB88" blockId="168.[136,779,942,1285]" pageId="168" pageNumber="169">
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<taxonomicName id="4C4C4D68FF44D59DFF007048FD8FFBC8" authority="Bartholomai, 1973" authorityName="Bartholomai" authorityYear="1973" box="[136,607,1038,1060]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF44D59DFF007048FE70FBCF" box="[136,416,1038,1059]" italics="true" pageId="168" pageNumber="169">Protemnodon chinchillaensis</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFE2E7048FD8FFBC8" author="Bartholomai, A." box="[422,607,1038,1060]" pageId="168" pageNumber="169" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai, 1973</bibRefCitation>
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</taxonomicName>
|
||
:
|
||
<emphasis id="B938EAF9FF44D59DFDE07049FCD5FBC8" box="[616,773,1038,1060]" italics="true" pageId="168" pageNumber="169">Mem. Qld. Mus.</emphasis>
|
||
, 16, pp. 347–354, pl. 20, 21, 23. See also
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFDDA7068FF2FFB89" author="Flannery, T. F. & Archer, M." pageId="168" pageNumber="169" pagination="357 - 383" refId="ref146359" refString="Flannery, T. F. & Archer, M. (1984) The macropodoids (Marsupialia) of the early Pliocene Bow Local Fauna, central eastern New South Wales. The Australian Zoologist, 21 (4), 357 - 383." type="journal article" year="1984">Flannery & Archer (1984)</bibRefCitation>
|
||
, pp. 375–376, figs A & B.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFF007028FE66FB48" blockId="168.[136,779,942,1285]" pageId="168" pageNumber="169">
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<taxonomicName id="4C4C4D68FF44D59DFF007028FE56FB68" authority="Owen" authorityName=", Owen" authorityYear="1874" box="[136,390,1134,1156]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF44D59DFF007028FE92FB6F" box="[136,322,1134,1155]" italics="true" pageId="168" pageNumber="169">Protemnodon anak</emphasis>
|
||
Owen
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFE1B7029FDF1FB68" author="De Vis, C. W." box="[403,545,1134,1156]" pageId="168" pageNumber="169" pagination="75 - 133" refId="ref145589" refString="De Vis, C. W. (1895) A review of the fossil jaws of the Macropodidae in the Queensland Museum. Proceedings of the Linnean Society of New South Wales, 10, 75 - 133." type="journal article" year="1895">De Vis (1895)</bibRefCitation>
|
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(
|
||
<emphasis id="B938EAF9FF44D59DFDBA7029FDA2FB68" box="[562,626,1135,1156]" italics="true" pageId="168" pageNumber="169">partim</emphasis>
|
||
), pp. 104–109. Not
|
||
<taxonomicName id="4C4C4D68FF44D59DFF6F70C9FE62FB48" authority=", Owen, 1874" authorityName=", Owen" authorityYear="1874" box="[231,434,1166,1188]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF44D59DFF6F70C9FEFEFB4F" box="[231,302,1166,1188]" italics="true" pageId="168" pageNumber="169">P. anak</emphasis>
|
||
,
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFEB270C9FE62FB48" author="Owen, R." box="[314,434,1167,1188]" pageId="168" pageNumber="169" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
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<paragraph id="8BF336EBFF44D59DFF0070E8FE04FB08" blockId="168.[136,779,942,1285]" pageId="168" pageNumber="169">
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<treatmentCitationGroup id="AB5C11C5FF44D59DFF0070E8FD31FB28" box="[136,737,1198,1221]" pageId="168" pageNumber="169">
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<taxonomicName id="4C4C4D68FF44D59DFF0070E8FE18FB28" authority="Bartholomai" authorityName=", Bartholomai" authorityYear="1973" box="[136,456,1198,1220]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF44D59DFF0070E8FE97FB2F" box="[136,327,1198,1219]" italics="true" pageId="168" pageNumber="169">Protemnodon devisi</emphasis>
|
||
Bartholomai
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AED10FAFF44D59DFE4770E8FD0EFB28" author="Tedford, R. H. & Wells, R. T. & Barghoorn, S. F." box="[463,734,1198,1221]" page="187" pageId="168" pageNumber="169" year="1992">
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFE4770E8FD46FB29" author="Tedford, R. H. & Wells, R. T. & Barghoorn, S. F." box="[463,662,1198,1221]" pageId="168" pageNumber="169" pagination="173 - 193" refId="ref155532" refString="Tedford, R. H., Wells, R. T. & Barghoorn, S. F. (1992) Tirari Formation and contained faunas, Pliocene of the Lake Eyre Basin, South Australia. The Beagle: Records of the Museums and Art Galleries of the Northern Territory, 9 (1), 173 - 193. https: // doi. org / 10.5962 / p. 263124" type="journal article" year="1992">
|
||
Tedford
|
||
<emphasis id="B938EAF9FF44D59DFDA870E9FD80FB2F" box="[544,592,1198,1220]" italics="true" pageId="168" pageNumber="169">et al.</emphasis>
|
||
(1992)
|
||
</bibRefCitation>
|
||
, p. 187
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
Not
|
||
<taxonomicName id="4C4C4D68FF44D59DFF357089FE00FB08" authority=", Bartholomai, 1973" authorityName=", Bartholomai" authorityYear="1973" box="[189,464,1230,1252]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF44D59DFF357089FEDDFB0F" box="[189,269,1230,1252]" italics="true" pageId="168" pageNumber="169">P. devisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFE917088FE00FB08" author="Bartholomai, A." box="[281,464,1230,1252]" pageId="168" pageNumber="169" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFF0070A8FCD5FAE8" blockId="168.[136,779,942,1285]" box="[136,773,1262,1285]" pageId="168" pageNumber="169">
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||
<taxonomicName id="4C4C4D68FF44D59DFF0070A8FEBBFAE8" box="[136,363,1262,1284]" class="Mammalia" family="Macropodidae" genus="Protemnodon" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B938EAF9FF44D59DFF0070A8FEDAFAEF" box="[136,266,1262,1283]" italics="true" pageId="168" pageNumber="169">Protemnodon</emphasis>
|
||
Owen sp.
|
||
</taxonomicName>
|
||
indet.;
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFE3B70A8FDACFAE9" author="Tedford, R. H. & Wells, R. T. & Barghoorn, S. F." box="[435,636,1262,1285]" pageId="168" pageNumber="169" pagination="173 - 193" refId="ref155532" refString="Tedford, R. H., Wells, R. T. & Barghoorn, S. F. (1992) Tirari Formation and contained faunas, Pliocene of the Lake Eyre Basin, South Australia. The Beagle: Records of the Museums and Art Galleries of the Northern Territory, 9 (1), 173 - 193. https: // doi. org / 10.5962 / p. 263124" type="journal article" year="1992">
|
||
Tedford
|
||
<emphasis id="B938EAF9FF44D59DFD8C70A9FDE5FAEF" box="[516,565,1262,1284]" italics="true" pageId="168" pageNumber="169">et al.</emphasis>
|
||
(1992)
|
||
</bibRefCitation>
|
||
, pp. 186–187.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF44D59CFF00716DFEC9FD29" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169" type="materials_examined">
|
||
<paragraph id="8BF336EBFF44D59DFF00716DFD69F949" blockId="168.[136,778,1323,1701]" pageId="168" pageNumber="169">
|
||
<materialsCitation id="3B243CB6FF44D59DFF00716DFD69F949" collectionCode="CPC, UCMP" country="Papua New Guinea" elevation="250" latitude="-7.2166667" location="west bank of Upper Watut River" longLatPrecision="1303" longitude="146.53334" municipality="Otibanda Fm." pageId="168" pageNumber="169" specimenCode="CPC 6771, UCMP 69895" specimenCount="1" stateProvince="Morobe Province" typeStatus="holotype">
|
||
<emphasis id="B938EAF9FF44D59DFF00716DFED4FAA9" bold="true" box="[136,260,1323,1349]" pageId="168" pageNumber="169">
|
||
<typeStatus id="54F78849FF44D59DFF00716DFF2FFAA9" box="[136,255,1323,1349]" pageId="168" pageNumber="169" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DBEA9E90FF44D59DFF00710DFED7FA89" box="[136,263,1355,1381]" pageId="168" pageNumber="169">CPC 6771</specimenCode>
|
||
: near-complete R dentary preserving
|
||
<date id="FFF2102BFF44D59DFD58710DFD36FA89" box="[720,742,1355,1381]" pageId="168" pageNumber="169">i1</date>
|
||
& p3–m4; posteromedial fracture along margin of pterygoid fossa with pterygoid process not preserved. Figured
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFD4371CDFF05FA2A" author="Plane, M. D." pageId="168" pageNumber="169" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane (1967)</bibRefCitation>
|
||
, figs 5 & 6. Reassociated here are the holotype, CPC 6771, and specimen
|
||
<specimenCode id="DBEA9E90FF44D59DFE8B71ADFE7CF9EA" box="[259,428,1515,1542]" pageId="168" pageNumber="169">UCMP 69895</specimenCode>
|
||
, a semi-complete L dentary preserving the base of the
|
||
<date id="FFF2102BFF44D59DFE53724DFE21F9C9" box="[475,497,1547,1573]" pageId="168" pageNumber="169">i1</date>
|
||
and p3–m4, based on extreme similarity in dimensions, degree of molar wear, morphology, shape of mandibular symphyseal plates and preservation. UCMP 69895 was collected by the same expedition from the same locality as the holotype.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFF00728DFEE2F908" blockId="168.[136,778,1738,2054]" box="[136,306,1738,1765]" pageId="168" pageNumber="169">
|
||
<emphasis id="B938EAF9FF44D59DFF00728DFEE2F908" bold="true" box="[136,306,1738,1765]" pageId="168" pageNumber="169">Type locality:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFF0072ADFD58F7E9" blockId="168.[136,778,1738,2054]" pageId="168" pageNumber="169">
|
||
‘Watut 1’, UCMP V6234,
|
||
<location id="8E936030FF44D59DFE4272ADFF16F8C9" LSID="urn:lsid:plazi:treatment:03E587FDFF44D58FFF00772CFBB5F7E8:8E936030FF44D59DFE4272ADFF16F8C9" country="Papua New Guinea" latitude="-7.2166667" longLatPrecision="1303" longitude="146.53334" municipality="Otibanda Fm." name="west bank of Upper Watut River" pageId="168" pageNumber="169" stateProvince="Morobe Province">west bank of Upper Watut River</location>
|
||
(
|
||
<geoCoordinate id="EE78502CFF44D59DFF69734DFE90F8C9" box="[225,320,1803,1829]" degrees="7" direction="south" minutes="13" orientation="latitude" pageId="168" pageNumber="169" precision="925" value="-7.2166667">7°13’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE78502CFF44D59DFEDF734DFE05F8C9" box="[343,469,1803,1829]" degrees="146" direction="east" minutes="32" orientation="longitude" pageId="168" pageNumber="169" precision="925" value="146.53334">146°32’ E</geoCoordinate>
|
||
),
|
||
<collectingMunicipality id="6B97AC91FF44D59DFE7C734DFD4BF8C9" box="[500,667,1803,1829]" pageId="168" pageNumber="169">Otibanda Fm.</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="4988F809FF44D59DFD39734DFF3EF8A9" country="Papua New Guinea" name="Morobe" pageId="168" pageNumber="169">Morobe Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35B767BFF44D59DFF73736DFE0BF8A9" box="[251,475,1835,1861]" name="Papua New Guinea" pageId="168" pageNumber="169">Papua New Guinea</collectingCountry>
|
||
. The type section contains fossiliferous sandstone and mudstone with conglomerate and intercalated pyroclastic rocks. Potassium–Argon dating of a pyroclastic rock sample, taken from an outcrop of the Otibanda Fm. approximately
|
||
<quantity id="4CB49B0EFF44D59DFDA473EDFDA6F829" box="[556,630,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="168" pageNumber="169" unit="m" value="250.0">
|
||
<elevation id="0061D1D8FF44D59DFDA473EDFDA6F829" box="[556,630,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="168" pageNumber="169" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
downstream from the type locality, gave an age of 2.9 ± 0.4 Ma (
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFD43738DFEF8F7EA" author="Hoch, E. & Holm, P. M." pageId="168" pageNumber="169" pagination="181 - 195" refId="ref148234" refString="Hoch, E. & Holm, P. M. (1986) New K / Ar age determinations of the Awe Fauna Gangue, Papua New Guinea: consequences for Papuaustralian Late Cenozoic biostratigraphy. Modern Geology, 10, 181 - 195." type="journal article" year="1986">Hoch & Holm 1986</bibRefCitation>
|
||
), placing it in the late Pliocene.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA274ECFC14FF28" blockId="168.[809,1453,170,677]" box="[810,964,170,196]" pageId="168" pageNumber="169">
|
||
<emphasis id="B938EAF9FF44D59DFCA274ECFC14FF28" bold="true" box="[810,964,170,196]" pageId="168" pageNumber="169">
|
||
<typeStatus id="54F78849FF44D59DFCA274ECFC74FF28" box="[810,932,170,196]" pageId="168" pageNumber="169" type="paratype">Paratype</typeStatus>
|
||
(s):
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2748DFC01FD49" blockId="168.[809,1453,170,677]" pageId="168" pageNumber="169">
|
||
Premaxilla fragments:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBB3748DFB5DFF09" box="[1083,1165,203,229]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69832. Partial maxilla:
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA274ADFCACFEE9" box="[810,892,235,261]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69851–69854,
|
||
<collectionCode id="ED5DAE2EFF44D59DFBA474ADFBAEFEE9" box="[1068,1150,235,261]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69857. I1:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB7674ADFA80FEE9" box="[1278,1360,235,261]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69790– 69800,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC08754DFC02FEC9" box="[896,978,267,293]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69877,
|
||
<collectionCode id="ED5DAE2EFF44D59DFBA7754DFB51FEC9" box="[1071,1153,267,293]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69859. I2:
|
||
<collectionCode id="ED5DAE2EFF44D59DFA8C754DFA86FEC9" box="[1284,1366,267,293]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
63631,
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA2756DFCACFEA9" box="[810,892,299,325]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69803,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC6D756DFBE7FEA9" box="[997,1079,299,325]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69804,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB28756DFB22FEA9" box="[1184,1266,299,325]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69806,
|
||
<collectionCode id="ED5DAE2EFF44D59DFAD3756DFA7DFEA9" box="[1371,1453,299,325]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69825,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC0C750DFC06FE89" box="[900,982,331,357]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69826. I3:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBEC750DFB66FE89" box="[1124,1206,331,357]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69827–69831. DP2:
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA2752DFCACFE69" box="[810,892,363,389]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69833. P3:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB8A752DFB84FE69" box="[1026,1108,363,389]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69834–69839,
|
||
<collectionCode id="ED5DAE2EFF44D59DFA8D752DFA87FE69" box="[1285,1367,363,389]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69863,
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA275CDFCACFE49" box="[810,892,395,421]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69864. M1:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB9F75CDFBB9FE49" box="[1047,1129,395,421]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69840–69842. M3:
|
||
<collectionCode id="ED5DAE2EFF44D59DFAD275CDFA7CFE49" box="[1370,1452,395,421]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69843,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC0675EDFC30FE29" box="[910,992,427,453]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69844. M4:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB1775EDFB21FE29" box="[1183,1265,427,453]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69847–69849. Dentary fragments:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBA4758DFBAEFE09" box="[1068,1150,459,485]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69896,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB7E758DFA98FE09" box="[1270,1352,459,485]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69897,
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA175ADFCABFDE9" box="[809,891,491,517]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69981,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC6C75ADFBE6FDE9" box="[996,1078,491,517]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69986,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB1775ADFB21FDE9" box="[1183,1265,491,517]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69987,
|
||
<collectionCode id="ED5DAE2EFF44D59DFAD275ADFA7CFDE9" box="[1370,1452,491,517]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69991,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC09764DFC03FDC9" box="[897,979,523,549]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69899,
|
||
<collectionCode id="ED5DAE2EFF44D59DFBBB764DFB55FDC9" box="[1075,1157,523,549]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69900. Juvenile dentary:
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA1766DFCB0FDA9" box="[809,864,555,581]" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="168" pageNumber="169">CPC</collectionCode>
|
||
6772. i1:
|
||
<collectionCode id="ED5DAE2EFF44D59DFC5C766DFBF6FDA9" box="[980,1062,555,581]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69860,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB0A766DFB04FDA9" box="[1154,1236,555,581]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69871–69878. p3:
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA1760DFCABFD89" box="[809,891,587,613]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69861,
|
||
<collectionCode id="ED5DAE2EFF44D59DFC6C760DFBE6FD89" box="[996,1078,587,613]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69863,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB17760DFB21FD89" box="[1183,1265,587,613]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69864,
|
||
<collectionCode id="ED5DAE2EFF44D59DFAD2760DFA7CFD89" box="[1370,1452,587,613]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69879. m1:
|
||
<collectionCode id="ED5DAE2EFF44D59DFC4C762DFBC6FD69" box="[964,1046,619,645]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69865. m3:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB37762DFAC1FD69" box="[1215,1297,619,645]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69870. m4:
|
||
<collectionCode id="ED5DAE2EFF44D59DFCA176CDFCABFD49" box="[809,891,651,677]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
69898.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2768CFBFAFD08" blockId="168.[810,1066,714,740]" box="[810,1066,714,740]" pageId="168" pageNumber="169">
|
||
<emphasis id="B938EAF9FF44D59DFCA2768CFBFAFD08" bold="true" box="[810,1066,714,740]" pageId="168" pageNumber="169">Referred specimens:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2774AFBDFFCC9" blockId="168.[809,1452,779,1829]" box="[810,1039,779,805]" pageId="168" pageNumber="169">
|
||
<emphasis id="B938EAF9FF44D59DFCA2774AFBDFFCC9" box="[810,1039,779,805]" italics="true" pageId="168" pageNumber="169">
|
||
<collectingCountry id="F35B767BFF44D59DFCA2774AFBDFFCC9" box="[810,1039,779,805]" name="Papua New Guinea" pageId="168" pageNumber="169">Papua New Guinea</collectingCountry>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2776DFC3EFC49" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Site 2, Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBAB776DFBA2FCA9" box="[1059,1138,811,837]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45345 vertebral fragment,
|
||
<collectionCode id="ED5DAE2EFF44D59DFCD7770DFCA2FC89" box="[863,882,843,869]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
calcaneus, talar fragment, metatarsals
|
||
<collectionCode id="ED5DAE2EFF44D59DFAB1770DFA87FC89" box="[1337,1367,843,869]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
and
|
||
<collectionCode id="ED5DAE2EFF44D59DFA1C770DFA78FC89" box="[1428,1448,843,869]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
, distal pedal phalanx
|
||
<collectionCode id="ED5DAE2EFF44D59DFBC9772DFB8EFC69" box="[1089,1118,875,901]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
, and proximal & middle pedal phalanges
|
||
<collectionCode id="ED5DAE2EFF44D59DFC5D77CDFC39FC49" box="[981,1001,907,933]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA277EDFA5BFBE9" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Sunshine 3, Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBCB77EDFB42FC29" box="[1091,1170,939,965]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
71414 partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFAB877EDFA93FC29" box="[1328,1347,939,965]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
talus and metatarsal
|
||
<collectionCode id="ED5DAE2EFF44D59DFC51778DFC26FC09" box="[985,1014,971,997]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
, partial LR metatarsals
|
||
<collectionCode id="ED5DAE2EFF44D59DFB75778DFAC1FC09" box="[1277,1297,971,997]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
and proximal pedal phalanges
|
||
<collectionCode id="ED5DAE2EFF44D59DFB9277ADFBE7FBE9" box="[1050,1079,1003,1029]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
, and middle pedal phalanx
|
||
<collectionCode id="ED5DAE2EFF44D59DFAE177ADFA57FBE9" box="[1385,1415,1003,1029]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2704DFC07FBA9" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Sunshine General, Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB04704DFB0BFBC9" box="[1164,1243,1035,1061]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45344 LR femoral fragments.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2700DFA79FA69" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Watut 1, Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBA1700DFBA8FB89" box="[1065,1144,1099,1125]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70036
|
||
<collectionCode id="ED5DAE2EFF44D59DFB46700DFB31FB89" box="[1230,1249,1099,1125]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
i1;
|
||
<collectionCode id="ED5DAE2EFF44D59DFA87700DFA8EFB89" box="[1295,1374,1099,1125]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70045 clavicle;
|
||
<collectionCode id="ED5DAE2EFF44D59DFC46702DFBCDFB69" box="[974,1053,1131,1157]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70059 partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFB52702DFB3DFB69" box="[1242,1261,1131,1157]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
humerus, ulna, radius and articulated manus;
|
||
<collectionCode id="ED5DAE2EFF44D59DFB3370CDFADAFB49" box="[1211,1290,1163,1189]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70054 partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFCD770EDFCA2FB29" box="[863,882,1195,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
radius;
|
||
<collectionCode id="ED5DAE2EFF44D59DFC5470EDFBFBFB29" box="[988,1067,1195,1221]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70065 partial L femur;
|
||
<collectionCode id="ED5DAE2EFF44D59DFAD570EDFA7CFB29" box="[1373,1452,1195,1221]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70066 partial L femur;
|
||
<collectionCode id="ED5DAE2EFF44D59DFBEE708DFB65FB09" box="[1126,1205,1227,1253]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70038, 70039, 70584 and 70585 femoral fragments, partial LR tibiae, L and partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFC6E714DFC29FAC9" box="[998,1017,1291,1317]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
calcaneus, L cuboid, LR metatarsals
|
||
<collectionCode id="ED5DAE2EFF44D59DFCD7716DFCADFAA9" box="[863,893,1323,1349]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
and
|
||
<collectionCode id="ED5DAE2EFF44D59DFC30716DFC1CFAA9" box="[952,972,1323,1349]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
, and proximal and distal pedal phalanges
|
||
<collectionCode id="ED5DAE2EFF44D59DFCD7710DFCAEFA89" box="[863,894,1355,1381]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
;
|
||
<collectionCode id="ED5DAE2EFF44D59DFC02710DFC09FA89" box="[906,985,1355,1381]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70078 L cuboid;
|
||
<collectionCode id="ED5DAE2EFF44D59DFB2C710DFB23FA89" box="[1188,1267,1355,1381]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70006 partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFA10710DFA7BFA89" box="[1432,1451,1355,1381]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
metatarsal
|
||
<collectionCode id="ED5DAE2EFF44D59DFC51712DFC28FA69" box="[985,1016,1387,1413]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
;
|
||
<collectionCode id="ED5DAE2EFF44D59DFB8B712DFB82FA69" box="[1027,1106,1387,1413]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
70007 partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFB7C712DFAD7FA69" box="[1268,1287,1387,1413]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
metatarsal
|
||
<collectionCode id="ED5DAE2EFF44D59DFA00712DFA76FA69" box="[1416,1446,1387,1413]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA171CDFABCF9E9" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Woodard 2, Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBCF71CDFB46FA49" box="[1095,1174,1419,1445]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45246 partial juvenile
|
||
<collectionCode id="ED5DAE2EFF44D59DFA1071CDFA7BFA49" box="[1432,1451,1419,1445]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
premaxilla, partial L maxilla, partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFB7D71EDFAD8FA29" box="[1269,1288,1451,1477]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
maxilla,
|
||
<collectionCode id="ED5DAE2EFF44D59DFAE671EDFA51FA29" box="[1390,1409,1451,1477]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
P3, and partial mandible;
|
||
<collectionCode id="ED5DAE2EFF44D59DFBC5718DFB4CFA09" box="[1101,1180,1483,1509]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45248 partial
|
||
<collectionCode id="ED5DAE2EFF44D59DFAB0718DFA9BFA09" box="[1336,1355,1483,1509]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
maxilla;
|
||
<collectionCode id="ED5DAE2EFF44D59DFCD771ADFC7EF9E9" box="[863,942,1515,1541]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45250
|
||
<collectionCode id="ED5DAE2EFF44D59DFB8971ADFBC4F9E9" box="[1025,1044,1515,1541]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
talus;
|
||
<collectionCode id="ED5DAE2EFF44D59DFBD671ADFB7DF9E9" box="[1118,1197,1515,1541]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45253 L cuboid.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA1724DFBA5F949" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Woodard 3/Site 3,Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB0D724DFB04F9C9" box="[1157,1236,1547,1573]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45247 L calcaneus,
|
||
<collectionCode id="ED5DAE2EFF44D59DFCD7726DFCA2F9A9" box="[863,882,1579,1605]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
talus, articulated
|
||
<collectionCode id="ED5DAE2EFF44D59DFBC7726DFBB2F9A9" box="[1103,1122,1579,1605]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
cuboid, ectocuneiform and metatarsals
|
||
<collectionCode id="ED5DAE2EFF44D59DFC6D720DFC2AF989" box="[997,1018,1611,1637]" pageId="168" pageNumber="169">II</collectionCode>
|
||
,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB80720DFBF4F989" box="[1032,1060,1611,1637]" pageId="168" pageNumber="169">III</collectionCode>
|
||
and
|
||
<collectionCode id="ED5DAE2EFF44D59DFBD6720DFBABF989" box="[1118,1147,1611,1637]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
,
|
||
<collectionCode id="ED5DAE2EFF44D59DFB00720DFB4BF989" box="[1160,1179,1611,1637]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
metatarsal
|
||
<collectionCode id="ED5DAE2EFF44D59DFA97720DFAE3F989" box="[1311,1331,1611,1637]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
, proximal, middle and distal pedal phalanges
|
||
<collectionCode id="ED5DAE2EFF44D59DFB52722DFB27F969" box="[1242,1271,1643,1669]" pageId="168" pageNumber="169">IV</collectionCode>
|
||
, and middle and distal pedal phalanges
|
||
<collectionCode id="ED5DAE2EFF44D59DFBD472CDFBA0F949" box="[1116,1136,1675,1701]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA172EDFA84F909" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Woodard 4, Watut River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFBC972EDFB40F929" box="[1089,1168,1707,1733]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
45244 partial premaxilla, maxillae and L dentary, and tibial fragments.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA172ADFA92F8C9" blockId="168.[809,1452,779,1829]" pageId="168" pageNumber="169">
|
||
Northwest bank of Kikori River,
|
||
<collectingRegion id="4988F809FF44D59DFB4B72ADFABDF8E9" box="[1219,1389,1771,1797]" country="Papua New Guinea" name="Gulf" pageId="168" pageNumber="169">Gulf Province</collectingRegion>
|
||
:
|
||
<collectionCode id="ED5DAE2EFF44D59DFAF672ADFA7BF8E9" box="[1406,1451,1771,1797]" country="Australia" name="Australian Museum" pageId="168" pageNumber="169" type="Museum">AM</collectionCode>
|
||
F134486 pelvic fragment with acetabulum.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2730DFC30F889" blockId="168.[810,1452,1867,2053]" box="[810,992,1867,1893]" pageId="168" pageNumber="169">
|
||
<emphasis id="B938EAF9FF44D59DFCA2730DFC30F889" box="[810,992,1867,1893]" italics="true" pageId="168" pageNumber="169">
|
||
<collectingRegion id="4988F809FF44D59DFCA2730DFC30F889" box="[810,992,1867,1893]" country="Australia" name="South Australia" pageId="168" pageNumber="169">South Australia</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA2732DFAD0F849" blockId="168.[810,1452,1867,2053]" pageId="168" pageNumber="169">
|
||
Stirton Quarry (
|
||
<collectionCode id="ED5DAE2EFF44D59DFC55732DFC22F86A" box="[989,1010,1899,1926]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="168" pageNumber="169" type="Museum">V</collectionCode>
|
||
5772), Lake Kanunka:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB7F732DFA96F869" box="[1271,1350,1899,1925]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
56894 L dentary;
|
||
<collectionCode id="ED5DAE2EFF44D59DFC4A73CDFBC1F849" box="[962,1041,1931,1957]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="168" pageNumber="169" type="University or college">UCMP</collectionCode>
|
||
156893
|
||
<collectionCode id="ED5DAE2EFF44D59DFBFA73CDFB55F849" box="[1138,1157,1931,1957]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
calcaneus.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFCA273EDFC69F7E9" blockId="168.[810,1452,1867,2053]" pageId="168" pageNumber="169">
|
||
Site 6 (
|
||
<geoCoordinate id="EE78502CFF44D59DFC0573EDFBEBF829" box="[909,1083,1963,1990]" degrees="27" direction="south" minutes="51.327" orientation="latitude" pageId="168" pageNumber="169" precision="1" value="-27.85545">27° 51.327’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE78502CFF44D59DFBDA73EDFAC0F829" box="[1106,1296,1963,1990]" degrees="137" direction="east" minutes="53.385" orientation="longitude" pageId="168" pageNumber="169" precision="1" value="137.88976">137° 53.385’ E</geoCoordinate>
|
||
), Toolapinna Waterhole, Warburton River:
|
||
<collectionCode id="ED5DAE2EFF44D59DFB46738DFACFF809" box="[1230,1311,1995,2021]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="168" pageNumber="169" type="Museum">SAMA</collectionCode>
|
||
P50566
|
||
<collectionCode id="ED5DAE2EFF44D59DFA11738DFA7CF809" box="[1433,1452,1995,2021]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="168" pageNumber="169">R</collectionCode>
|
||
dentary.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF0074EDFEAFFF09" blockId="169.[136,778,171,293]" pageId="169" pageNumber="170">
|
||
Toolapinna Waterhole, Warburton River:
|
||
<collectionCode id="ED5DAE2EFF45D59CFDD474EDFD7DFF29" box="[604,685,171,197]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="169" pageNumber="170" type="Museum">SAMA</collectionCode>
|
||
P25504 partial L maxilla.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF0074ADFEDEFEC9" blockId="169.[136,778,171,293]" pageId="169" pageNumber="170">
|
||
Lawson–Daily Quarry, Lake Palankarinna:
|
||
<collectionCode id="ED5DAE2EFF45D59CFDE774ADFD6EFEE9" box="[623,702,235,261]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="169" pageNumber="170" type="University or college">UCMP</collectionCode>
|
||
57195 L tibia.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF00750DFEC4FE89" blockId="169.[136,779,331,454]" box="[136,276,331,357]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFF00750DFEC4FE89" box="[136,276,331,357]" italics="true" pageId="169" pageNumber="170">
|
||
<collectingRegion id="4988F809FF45D59CFF00750DFEC4FE89" box="[136,276,331,357]" country="Australia" name="Queensland" pageId="169" pageNumber="170">Queensland</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF00752DFE1EFE29" blockId="169.[136,779,331,454]" pageId="169" pageNumber="170">
|
||
Chinchilla, Darling Downs (site unknown):
|
||
<collectionCode id="ED5DAE2EFF45D59CFDFE752DFD73FE69" box="[630,675,363,389]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="169" pageNumber="170" type="Museum">QM</collectionCode>
|
||
F4719 L maxilla;
|
||
<collectionCode id="ED5DAE2EFF45D59CFEAA75CDFE9FFE49" box="[290,335,395,421]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="169" pageNumber="170" type="Museum">QM</collectionCode>
|
||
F5239 partial juvenile L maxilla;
|
||
<collectionCode id="ED5DAE2EFF45D59CFD5575CDFCDAFE49" box="[733,778,395,421]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="169" pageNumber="170" type="Museum">QM</collectionCode>
|
||
F5246 partial
|
||
<collectionCode id="ED5DAE2EFF45D59CFED375EDFEBEFE29" box="[347,366,427,453]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="169" pageNumber="170">R</collectionCode>
|
||
dentary.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF0075AAFE83FDE9" blockId="169.[136,777,491,582]" box="[136,339,491,517]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFF0075AAFE83FDE9" box="[136,339,491,517]" italics="true" pageId="169" pageNumber="170">
|
||
<collectingRegion id="4988F809FF45D59CFF0075AAFE83FDE9" box="[136,339,491,517]" country="Australia" name="New South Wales" pageId="169" pageNumber="170">New South Wales</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF00764DFDDFFDA9" blockId="169.[136,777,491,582]" pageId="169" pageNumber="170">
|
||
Bow LF, Hunter Valley:
|
||
<collectionCode id="ED5DAE2EFF45D59CFE21764DFE06FDC9" box="[425,470,523,549]" country="Australia" name="Australian Museum" pageId="169" pageNumber="170" type="Museum">AM</collectionCode>
|
||
F59530 partial L maxilla;
|
||
<collectionCode id="ED5DAE2EFF45D59CFF35766DFF3AFDA9" box="[189,234,555,581]" country="Australia" name="Australian Museum" pageId="169" pageNumber="170" type="Museum">AM</collectionCode>
|
||
F59533 partial L maxilla.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF00762AFF36FD69" blockId="169.[136,778,620,709]" box="[136,230,620,645]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFF00762AFF36FD69" box="[136,230,620,645]" italics="true" pageId="169" pageNumber="170">
|
||
<collectingRegion id="4988F809FF45D59CFF00762AFF36FD69" box="[136,230,620,645]" country="Australia" name="Victoria" pageId="169" pageNumber="170">Victoria</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF0076CDFEC9FD29" blockId="169.[136,778,620,709]" pageId="169" pageNumber="170">
|
||
Nowa Nowa Arm, Lake Tyers:
|
||
<collectionCode id="ED5DAE2EFF45D59CFE7176CDFDEAFD49" box="[505,570,651,677]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="169" pageNumber="170" type="Museum">NMV</collectionCode>
|
||
P26893 partial
|
||
<collectionCode id="ED5DAE2EFF45D59CFD7F76CDFCDAFD49" box="[759,778,651,677]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="169" pageNumber="170">R</collectionCode>
|
||
maxilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF45D59CFF0076ACFC62FDE9" pageId="169" pageNumber="170" type="diagnosis">
|
||
<paragraph id="8BF336EBFF45D59CFF0076ACFE03FCE8" blockId="169.[136,779,746,2085]" box="[136,467,746,772]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFF0076ACFE03FCE8" bold="true" box="[136,467,746,772]" pageId="169" pageNumber="170">Revised specific diagnosis:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF00774DFD94FBA9" blockId="169.[136,779,746,2085]" pageId="169" pageNumber="170">
|
||
<taxonomicName id="4C4C4D68FF45D59CFF00774DFE77FCC9" authorityName="Plane" authorityYear="1967" box="[136,423,779,805]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF45D59CFF00774DFE77FCC9" box="[136,423,779,805]" italics="true" pageId="169" pageNumber="170">Protemnodon otibandus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all other species of
|
||
<taxonomicName id="4C4C4D68FF45D59CFEDC776DFE3BFCA9" authorityName="Owen" authorityYear="1874" box="[340,491,811,837]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF45D59CFEDC776DFE3BFCA9" box="[340,491,811,837]" italics="true" pageId="169" pageNumber="170">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
by one unique skeletal characteristic and by a combination of other dental and skeletal characteristics. The pes of
|
||
<taxonomicName id="4C4C4D68FF45D59CFDAF772AFD65FC69" authorityName="Plane" authorityYear="1967" box="[551,693,875,901]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF45D59CFDAF772AFD65FC69" box="[551,693,875,901]" italics="true" pageId="169" pageNumber="170">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from all other species of
|
||
<taxonomicName id="4C4C4D68FF45D59CFE5977CDFDB8FC49" authorityName="Owen" authorityYear="1874" box="[465,616,907,933]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF45D59CFE5977CDFDB8FC49" box="[465,616,907,933]" italics="true" pageId="169" pageNumber="170">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
in having a calcaneus with a slightly medially offset head relative to the calcaneal tuberosity. In
|
||
<taxonomicName id="4C4C4D68FF45D59CFE6C778AFDB4FC09" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[484,612,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFE6C778AFDB4FC09" box="[484,612,971,997]" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
this condition is more extreme, with the head more strongly medially displaced; in all other species of
|
||
<taxonomicName id="4C4C4D68FF45D59CFE7A704DFD59FBC9" authorityName="Owen" authorityYear="1874" box="[498,649,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF45D59CFE7A704DFD59FBC9" box="[498,649,1035,1061]" italics="true" pageId="169" pageNumber="170">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
the head of the calcaneus is not medially displaced.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF30700DFD3DFAC9" blockId="169.[136,779,746,2085]" pageId="169" pageNumber="170">
|
||
<taxonomicName id="4C4C4D68FF45D59CFF30700DFE1AFB89" authorityName="Plane" authorityYear="1967" box="[184,458,1099,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF45D59CFF30700DFE1AFB89" box="[184,458,1099,1125]" italics="true" pageId="169" pageNumber="170">Protemnodon otibandus</emphasis>
|
||
</taxonomicName>
|
||
is most similar in dentary morphology to
|
||
<taxonomicName id="4C4C4D68FF45D59CFECB702AFE4BFB69" authorityName=", Owen" authorityYear="1874" box="[323,411,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF45D59CFECB702AFE4BFB69" box="[323,411,1131,1157]" italics="true" pageId="169" pageNumber="170">P. anak</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C4D68FF45D59CFE38702AFD93FB69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[432,579,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF45D59CFE38702AFD93FB69" box="[432,579,1131,1157]" italics="true" pageId="169" pageNumber="170">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF45D59CFDD9702DFD61FB69" bold="true" box="[593,689,1131,1157]" pageId="169" pageNumber="170">
|
||
<taxonomicNameLabel id="A20B5782FF45D59CFDD9702DFD61FB69" box="[593,689,1131,1157]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF45D59CFD7D702AFF3AFB49" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFD7D702AFF3AFB49" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
. It differs from that of
|
||
<taxonomicName id="4C4C4D68FF45D59CFE7A70CAFD95FB49" authorityName=", Owen" authorityYear="1874" box="[498,581,1163,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF45D59CFE7A70CAFD95FB49" box="[498,581,1163,1189]" italics="true" pageId="169" pageNumber="170">P. anak</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF45D59CFDF570CAFCDAFB49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[637,778,1163,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF45D59CFDF570CAFCDAFB49" box="[637,778,1163,1189]" italics="true" pageId="169" pageNumber="170">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF45D59CFF0070EDFF36FB29" bold="true" box="[136,230,1195,1221]" pageId="169" pageNumber="170">
|
||
<taxonomicNameLabel id="A20B5782FF45D59CFF0070EDFF36FB29" box="[136,230,1195,1221]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in its less robust anterior dentary, particularly around the i1. It differs further from
|
||
<taxonomicName id="4C4C4D68FF45D59CFDB9708AFD54FB09" authorityName=", Owen" authorityYear="1874" box="[561,644,1227,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF45D59CFDB9708AFD54FB09" box="[561,644,1227,1253]" italics="true" pageId="169" pageNumber="170">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in its more dorsally deflected diastema; and from
|
||
<taxonomicName id="4C4C4D68FF45D59CFDD770AAFD35FAE9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[607,741,1259,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFDD770AAFD35FAE9" box="[607,741,1259,1285]" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having a more anterodorsally situated mental foramen.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF30716DFD98F969" blockId="169.[136,779,746,2085]" pageId="169" pageNumber="170">
|
||
<taxonomicName id="4C4C4D68FF45D59CFF30716DFE1DFAA9" authorityName="Plane" authorityYear="1967" box="[184,461,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF45D59CFF30716DFE1DFAA9" box="[184,461,1323,1349]" italics="true" pageId="169" pageNumber="170">Protemnodon otibandus</emphasis>
|
||
</taxonomicName>
|
||
is most similar in dental morphology to
|
||
<taxonomicName id="4C4C4D68FF45D59CFEB0710AFE7BFA89" authorityName="Bartholomai" authorityYear="1978" box="[312,427,1356,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF45D59CFEB0710AFE7BFA89" box="[312,427,1356,1381]" italics="true" pageId="169" pageNumber="170">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF45D59CFE6C710AFDB3FA89" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[484,611,1355,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFE6C710AFDB3FA89" box="[484,611,1355,1381]" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
. The dentition differs from
|
||
<taxonomicName id="4C4C4D68FF45D59CFEAB712AFE4BFA69" authorityName="Bartholomai" authorityYear="1978" box="[291,411,1388,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF45D59CFEAB712AFE4BFA69" box="[291,411,1388,1413]" italics="true" pageId="169" pageNumber="170">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having: P3 with a broader posterior relative to the anterior cusp and a higher lingual crest; relatively broader upper molars with a less anteriorly prominent precingulum, a more raised, more distinct postparacrista, and a urocrista; and more robust, less lanceolate i1. The dentition differs from that of
|
||
<taxonomicName id="4C4C4D68FF45D59CFD7D724AFF3AF9A9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFD7D724AFF3AF9A9" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having: P3 with more raised, more distinct buccal ridgelets; less rounded molars; and i1 with a thinner, more raised ventrolingual crest.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFF3072CDFC62FDE9" blockId="169.[136,779,746,2085]" lastBlockId="169.[810,1452,171,517]" pageId="169" pageNumber="170">
|
||
<taxonomicName id="4C4C4D68FF45D59CFF3072CDFE15F949" authorityName="Plane" authorityYear="1967" box="[184,453,1675,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF45D59CFF3072CDFE15F949" box="[184,453,1675,1701]" italics="true" pageId="169" pageNumber="170">Protemnodon otibandus</emphasis>
|
||
</taxonomicName>
|
||
is most similar in aspects of skeletal morphology to
|
||
<taxonomicName id="4C4C4D68FF45D59CFE1272EAFDCEF929" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[410,542,1707,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFE1272EAFE7BF929" box="[410,427,1708,1733]" italics="true" pageId="169" pageNumber="170">P</emphasis>
|
||
.
|
||
<emphasis id="B938EAF9FF45D59CFE3472EDFDCEF929" box="[444,542,1707,1733]" italics="true" pageId="169" pageNumber="170">tumbuna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF45D59CFDD272EAFD1EF929" authorityName="Bartholomai" authorityYear="1978" box="[602,718,1708,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF45D59CFDD272EAFD1EF929" box="[602,718,1708,1733]" italics="true" pageId="169" pageNumber="170">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
. The hindlimb differs from
|
||
<taxonomicName id="4C4C4D68FF45D59CFE08728AFE2DF909" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[384,509,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFE08728AFE2DF909" box="[384,509,1739,1765]" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having: a femur with a less elongate and more proximally situated quadratus tubercle; and a narrower tibia with a thinner, more raised proximolateral crest and a less distally extensive cnemial crest with a more defined peak. The pes further differs from that of
|
||
<taxonomicName id="4C4C4D68FF45D59CFEAA732AFE74F869" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[290,420,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF45D59CFEAA732AFE74F869" box="[290,420,1899,1925]" italics="true" pageId="169" pageNumber="170">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having: a calcaneus with a more planar plantar surface with less flaring of the medial margin and a smaller, shallower fossa cranial to the lateral talar facet; and more transversely compressed metatarsal V with dorsoplantarly shorter proximolateral process, longer metatarsal IV facet and larger medial plantar tubercle. The hindlimb differs from that of
|
||
<taxonomicName id="4C4C4D68FF45D59CFA9274EAFA5DFF29" authorityName="Bartholomai" authorityYear="1978" box="[1306,1421,172,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF45D59CFA9274EAFA5DFF29" box="[1306,1421,172,197]" italics="true" pageId="169" pageNumber="170">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having a tibia with a larger, broader proximal fibular facet. The pes differs from that of
|
||
<taxonomicName id="4C4C4D68FF45D59CFBF174AAFB3EFEE9" authorityName="Bartholomai" authorityYear="1978" box="[1145,1262,236,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF45D59CFBF174AAFB3EFEE9" box="[1145,1262,236,261]" italics="true" pageId="169" pageNumber="170">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having: talus with a deeper trochlea and a deeper concavity between the posterior plantar tubercle and the talar head; cuboid with a less dorsomedially flared dorsomedial section, a much smaller talar facet, and a larger, more plantomedially projected medial plantar tubercle; metatarsal IV with a dorsoplantarly shorter, more plantarly situated proximal cuboid fossa; and distal phalanx IV with a more rounded dorsal peak.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF45D59CFCA2766DFB4DFD69" pageId="169" pageNumber="170" type="etymology">
|
||
<paragraph id="8BF336EBFF45D59CFCA2766DFC6BFDA9" blockId="169.[810,1451,555,645]" box="[810,955,555,581]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFCA2766DFC6BFDA9" bold="true" box="[810,955,555,581]" pageId="169" pageNumber="170">Etymology:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFCA2760DFB4DFD69" blockId="169.[810,1451,555,645]" pageId="169" pageNumber="170">
|
||
In reference to the geological formation of the
|
||
<typeStatus id="54F78849FF45D59CFAF3760DFA7BFD89" box="[1403,1451,587,613]" pageId="169" pageNumber="170">type</typeStatus>
|
||
locality, the Otibanda Formation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF45D58CFCA276ECFCA7F7C9" lastPageId="185" lastPageNumber="186" pageId="169" pageNumber="170" type="description">
|
||
<paragraph id="8BF336EBFF45D59CFCA276ECFB70FD28" blockId="169.[810,1451,682,837]" box="[810,1184,682,709]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFCA276ECFB70FD28" bold="true" box="[810,1184,682,709]" pageId="169" pageNumber="170">Description and comparisons:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFCA2768DFC07FCA9" blockId="169.[810,1451,682,837]" pageId="169" pageNumber="170">
|
||
For craniodental description, see the original description by
|
||
<bibRefCitation id="EFDD4B1AFF45D59CFCD876ADFC0DFCE9" author="Plane, M. D." box="[848,989,747,773]" pageId="169" pageNumber="170" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane (1967</bibRefCitation>
|
||
, pp. 28–39). For comparative purposes, the cranium, dentary and dentition are figured below (
|
||
<figureCitation id="13772A6EFF45D59CFCBA776DFC56FCA9" box="[818,902,811,837]" captionStart="FIGURE 98" captionStartId="170.[136,235,1740,1764]" captionTargetBox="[162,1425,178,1676]" captionTargetId="figure-27@170.[151,1436,167,1687]" captionTargetPageId="170" captionText="FIGURE 98. upper dentition of P. otibandus: (a–b) surface scan images of left I1 of UCMP 45246 in (a) lateral/buccal, and (b) anterior views; (c–d) surface scan images of left premaxilla fragment with partial I1 and I2–3 of paratype UCMP 69832 in (c) lateral/buccal, and (d) occlusal views; (e–g) surface scan images of partial left maxilla with P3–M3 of paratype UCMP 69857 in (e) buccal, (f) lingual, and (g) occlusal views; and (h) partial right maxilla of NMV P22650 from Jemmy’s Point, Lake Tyers, in occlusal view." figureDoi="http://doi.org/10.5281/zenodo.10993992" httpUri="https://zenodo.org/record/10993992/files/figure.png" pageId="169" pageNumber="170">Figs 98</figureCitation>
|
||
&
|
||
<figureCitation id="13772A6EFF45D59CFC22776DFC1AFCA9" box="[938,970,811,837]" captionStart="FIGURE 99" captionStartId="171.[136,235,1836,1860]" captionTargetBox="[162,1425,209,1809]" captionTargetId="figure-17@171.[151,1436,198,1817]" captionTargetPageId="171" captionText="FIGURE 99. dentaries and lower dentition of P. otibandus: (a–b) surface scan images of re-articulated right dentary of holotype CPC 6771 and left dentary of UCMP 69895 in (a) occlusal, and (b) ventral views; (c–d) surface scan images of right dentary of CPC 6771 in (c) buccal, and (d) lingual views; (e) partial right dentary of SAMA P50566 from Toolapinna Waterhole, Lake Eyre Basin, in buccal view; and (f–h) surface scan images of right p3–m4 of CPC 6771 in (f) occlusal, (g) buccal, and (h) lingual views." figureDoi="http://doi.org/10.5281/zenodo.10993994" httpUri="https://zenodo.org/record/10993994/files/figure.png" pageId="169" pageNumber="170">99</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFCA2772DFB5CFC68" blockId="169.[810,1452,874,2085]" box="[810,1164,874,901]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFCA2772DFB5CFC68" bold="true" box="[810,1164,874,901]" pageId="169" pageNumber="170">Pectoral girdle and forelimb</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFCA277CDFC64FB09" blockId="169.[810,1452,874,2085]" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFCA277CDFBF7FC49" box="[810,1063,907,933]" italics="true" pageId="169" pageNumber="170">
|
||
Clavicle (
|
||
<figureCitation id="13772A6EFF45D59CFC1F77CDFBF0FC49" box="[919,1056,907,933]" captionStart="FIGURE 100" captionStartId="172.[136,235,1900,1924]" captionTargetBox="[162,1425,212,1869]" captionTargetId="figure-27@172.[151,1436,201,1880]" captionTargetPageId="172" captionText="FIGURE 100. surface scan images of right upper forelimb elements of P. otibandus: (a–c) clavicle of UCMP 70045 in (a) anterior, (b) dorsal, and (c) ventral views; (d–e) distal humeral fragment of UCMP 70059 in (d) caudal, and (e) cranial views; and (f–g) radius of UCMP 70059 in (f) cranial, and (g) caudal views." figureDoi="http://doi.org/10.5281/zenodo.10993996" httpUri="https://zenodo.org/record/10993996/files/figure.png" pageId="169" pageNumber="170">Fig. 100a–c</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: craniocaudally compressed in the middle section and at the acromial articular end, becomes rounded in cross-section toward the sternal articular end; curves gently posteriorly towards both ends in caudal view, to be gently convex anteriorly; steadily increases in depth medially to the slightly thickened, smoothly ventrally curved acromial articular end, and forms a blunted point. Sternal articular end thickened, rounded in medial view; epiphysis not preserved, but the medial surface has a convex posterior part and a small, deep fossa in its centre.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D59CFCD270ADFAD9F969" blockId="169.[810,1452,874,2085]" pageId="169" pageNumber="170">
|
||
The clavicle of
|
||
<taxonomicName id="4C4C4D68FF45D59CFB9770AAFB60FAE9" authorityName="Plane" authorityYear="1967" box="[1055,1200,1259,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF45D59CFB9770AAFB60FAE9" box="[1055,1200,1259,1285]" italics="true" pageId="169" pageNumber="170">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of all compared species in being more craniocaudally compressed. It further differs from that of
|
||
<taxonomicName id="4C4C4D68FF45D59CFA90716AFA7CFAA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1304,1452,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF45D59CFA90716AFA7CFAA9" box="[1304,1452,1323,1349]" italics="true" pageId="169" pageNumber="170">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF45D59CFCA2710DFC58FA89" bold="true" box="[810,904,1355,1381]" pageId="169" pageNumber="170">
|
||
<taxonomicNameLabel id="A20B5782FF45D59CFCA2710DFC58FA89" box="[810,904,1355,1381]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being slightly less robust, with a relatively slightly smaller acromial articular end; from
|
||
<taxonomicName id="4C4C4D68FF45D59CFA93712DFA7BFA69" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1307,1451,1387,1413]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF45D59CFA93712DFA7BFA69" box="[1307,1451,1387,1413]" italics="true" pageId="169" pageNumber="170">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being more curved, with a relatively larger acromial articular end; from
|
||
<taxonomicName id="4C4C4D68FF45D59CFB8D71EDFBB2FA29" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1029,1122,1451,1477]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF45D59CFB8D71EDFBB2FA29" box="[1029,1122,1451,1477]" italics="true" pageId="169" pageNumber="170">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF45D59CFB1171EAFAEAFA29" authorityName="Desmarest" authorityYear="1817" box="[1177,1338,1451,1477]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF45D59CFB1171EAFAEAFA29" box="[1177,1338,1451,1477]" italics="true" pageId="169" pageNumber="170">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in having a slightly more caudally curved and less posteriorly curved acromial end, and a much less anteroposteriorly concave sternal articular surface; and from
|
||
<taxonomicName id="4C4C4D68FF45D59CFABA724AFA7BF9C9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1330,1451,1547,1573]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="169" pageNumber="170" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF45D59CFABA724AFA7BF9C9" box="[1330,1451,1547,1573]" italics="true" pageId="169" pageNumber="170">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and deeper, with a more ventrally curved acromial end with a more pointed tip, and the sternal end not strongly anteroposteriorly compressed.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF45D598FCD272CAFE7DFEC9" blockId="169.[810,1452,874,2085]" lastBlockId="173.[136,779,171,2085]" lastPageId="173" lastPageNumber="174" pageId="169" pageNumber="170">
|
||
<emphasis id="B938EAF9FF45D59CFCD272CAFBBFF949" box="[858,1135,1675,1701]" italics="true" pageId="169" pageNumber="170">
|
||
Humerus (
|
||
<figureCitation id="13772A6EFF45D59CFC5072CDFBB7F949" box="[984,1127,1675,1701]" captionStart="FIGURE 100" captionStartId="172.[136,235,1900,1924]" captionTargetBox="[162,1425,212,1869]" captionTargetId="figure-27@172.[151,1436,201,1880]" captionTargetPageId="172" captionText="FIGURE 100. surface scan images of right upper forelimb elements of P. otibandus: (a–c) clavicle of UCMP 70045 in (a) anterior, (b) dorsal, and (c) ventral views; (d–e) distal humeral fragment of UCMP 70059 in (d) caudal, and (e) cranial views; and (f–g) radius of UCMP 70059 in (f) cranial, and (g) caudal views." figureDoi="http://doi.org/10.5281/zenodo.10993996" httpUri="https://zenodo.org/record/10993996/files/figure.png" pageId="169" pageNumber="170">Fig. 100d–e</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: only one distal fragment known; distal end broad. Lateral supracondylar ridge quite thick and convex cranially at distal end; lateral epicondyle not preserved. Capitulum and ulnar facet broad, weakly projected and laterally situated, close to the lateral epicondyle; combined width is roughly three-quarters of the epicondylar width; capitulum smoothly convex; ulnar facet with the medial margin relatively straight and unbevelled; trochlea wide and quite shallow. Olecranon fossa broad and deep; radial fossa quite large and deep compared to the coronoid fossa; coronoid fossa partially preserved, appears shallow, broad and proximodistally short. Medial supracondylar bridge partially preserved, appears broad, and thin craniocaudally; supracondylar foramen appears large and quite flattened craniocaudally. Medial epicondyle large, rounded, rugose and very slightly distally projected.
|
||
</paragraph>
|
||
<caption id="DF336663FF46D59FFF00728AFEC8F889" ID-DOI="http://doi.org/10.5281/zenodo.10993992" ID-Zenodo-Dep="10993992" httpUri="https://zenodo.org/record/10993992/files/figure.png" pageId="170" pageNumber="171" startId="170.[136,235,1740,1764]" targetBox="[162,1425,178,1676]" targetPageId="170" targetType="figure">
|
||
<paragraph id="8BF336EBFF46D59FFF00728AFEC8F889" blockId="170.[136,1452,1740,1893]" pageId="170" pageNumber="171">
|
||
<emphasis id="B938EAF9FF46D59FFF00728AFEC2F909" bold="true" box="[136,274,1740,1765]" pageId="170" pageNumber="171">FIGURE 98.</emphasis>
|
||
upper dentition of
|
||
<taxonomicName id="4C4C4D68FF46D59FFE51728BFD87F909" authorityName="Plane" authorityYear="1967" box="[473,599,1741,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="170" pageNumber="171" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF46D59FFE51728BFD87F909" box="[473,599,1741,1765]" italics="true" pageId="170" pageNumber="171">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
: (a–b) surface scan images of left I1 of UCMP 45246 in (a) lateral/buccal, and (b) anterior views; (c–d) surface scan images of left premaxilla fragment with partial I1 and I2–3 of paratype UCMP 69832 in (c) lateral/buccal, and (d) occlusal views; (e–g) surface scan images of partial left maxilla with P3–M3 of paratype UCMP 69857 in (e) buccal, (f) lingual, and (g) occlusal views; and (h) partial right maxilla of NMV P22650 from Jemmy’s Point, Lake Tyers, in occlusal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF336663FF47D59EFF00736AFAA1F849" ID-DOI="http://doi.org/10.5281/zenodo.10993994" ID-Zenodo-Dep="10993994" httpUri="https://zenodo.org/record/10993994/files/figure.png" pageId="171" pageNumber="172" startId="171.[136,235,1836,1860]" targetBox="[162,1425,209,1809]" targetPageId="171" targetType="figure">
|
||
<paragraph id="8BF336EBFF47D59EFF00736AFAA1F849" blockId="171.[136,1452,1836,1957]" pageId="171" pageNumber="172">
|
||
<emphasis id="B938EAF9FF47D59EFF00736AFEC1F8A9" bold="true" box="[136,273,1836,1861]" pageId="171" pageNumber="172">FIGURE 99.</emphasis>
|
||
dentaries and lower dentition of
|
||
<taxonomicName id="4C4C4D68FF47D59EFDEE736BFD33F8A9" authorityName="Plane" authorityYear="1967" box="[614,739,1837,1861]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="171" pageNumber="172" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF47D59EFDEE736BFD33F8A9" box="[614,739,1837,1861]" italics="true" pageId="171" pageNumber="172">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
: (a–b) surface scan images of re-articulated right dentary of holotype CPC 6771 and left dentary of UCMP 69895 in (a) occlusal, and (b) ventral views; (c–d) surface scan images of right dentary of CPC 6771 in (c) buccal, and (d) lingual views; (e) partial right dentary of SAMA P50566 from Toolapinna Waterhole, Lake Eyre Basin, in buccal view; and (f–h) surface scan images of right p3–m4 of CPC 6771 in (f) occlusal, (g) buccal, and (h) lingual views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF336663FF40D599FF00732AFD30F829" ID-DOI="http://doi.org/10.5281/zenodo.10993996" ID-Zenodo-Dep="10993996" httpUri="https://zenodo.org/record/10993996/files/figure.png" pageId="172" pageNumber="173" startId="172.[136,235,1900,1924]" targetBox="[162,1425,212,1869]" targetPageId="172" targetType="figure">
|
||
<paragraph id="8BF336EBFF40D599FF00732AFD30F829" blockId="172.[136,1452,1900,1989]" pageId="172" pageNumber="173">
|
||
<emphasis id="B938EAF9FF40D599FF00732AFECDF868" bold="true" box="[136,285,1900,1924]" pageId="172" pageNumber="173">FIGURE 100.</emphasis>
|
||
surface scan images of right upper forelimb elements of
|
||
<taxonomicName id="4C4C4D68FF40D599FCD1732BFC05F869" authorityName="Plane" authorityYear="1967" box="[857,981,1901,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="172" pageNumber="173" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF40D599FCD1732BFC05F869" box="[857,981,1901,1925]" italics="true" pageId="172" pageNumber="173">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
: (a–c) clavicle of UCMP 70045 in (a) anterior, (b) dorsal, and (c) ventral views; (d–e) distal humeral fragment of UCMP 70059 in (d) caudal, and (e) cranial views; and (f–g) radius of UCMP 70059 in (f) cranial, and (g) caudal views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF41D598FF30756DFD15FCC9" blockId="173.[136,779,171,2085]" pageId="173" pageNumber="174">
|
||
The humerus of
|
||
<taxonomicName id="4C4C4D68FF41D598FEE5756AFE26FEA9" authorityName="Plane" authorityYear="1967" box="[365,502,299,325]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF41D598FEE5756AFE26FEA9" box="[365,502,299,325]" italics="true" pageId="173" pageNumber="174">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
does not differ from that of
|
||
<taxonomicName id="4C4C4D68FF41D598FF22750AFEFAFE89" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[170,298,331,357]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF41D598FF22750AFEFAFE89" box="[170,298,331,357]" italics="true" pageId="173" pageNumber="174">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
. It differs from
|
||
<taxonomicName id="4C4C4D68FF41D598FE62750AFDEFFE89" authorityName=", Owen" authorityYear="1874" box="[490,575,331,357]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF41D598FE62750AFDEFFE89" box="[490,575,331,357]" italics="true" pageId="173" pageNumber="174">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with a smaller capitulum and ulnar facet, a narrower medial supracondylar bridge, shallower trochlea, and a more curved, less distinct and less distally projected medial margin of the ulnar facet; from
|
||
<taxonomicName id="4C4C4D68FF41D598FDC4758AFD0FFE09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[588,735,459,485]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF41D598FDC4758AFD0FFE09" box="[588,735,459,485]" italics="true" pageId="173" pageNumber="174">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF41D598FD60758DFF69FDE9" bold="true" pageId="173" pageNumber="174">
|
||
<taxonomicNameLabel id="A20B5782FF41D598FD60758DFF69FDE9" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF41D598FF7D75AAFE86FDE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[245,342,492,517]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF41D598FF7D75AAFE86FDE9" box="[245,342,492,517]" italics="true" pageId="173" pageNumber="174">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF41D598FED775ADFE6CFDE9" bold="true" box="[351,444,491,517]" pageId="173" pageNumber="174">
|
||
<taxonomicNameLabel id="A20B5782FF41D598FED775ADFE6CFDE9" box="[351,444,491,517]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a less projected capitulum and ulnar facet, and less bevelled medial margin of the ulnar facet; from
|
||
<taxonomicName id="4C4C4D68FF41D598FDDD766DFD39FDA9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[597,745,555,581]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF41D598FDDD766DFD39FDA9" box="[597,745,555,581]" italics="true" pageId="173" pageNumber="174">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having a broader medial supracondylar bridge; from
|
||
<taxonomicName id="4C4C4D68FF41D598FF00762DFF39FD69" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[136,233,619,645]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF41D598FF00762DFF39FD69" box="[136,233,619,645]" italics="true" pageId="173" pageNumber="174">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in having a broader distal end, broader distal part of the lateral supracondylar ridge, and a relatively larger ulnar facet; from
|
||
<taxonomicName id="4C4C4D68FF41D598FE2176EAFD9FFD29" authorityName="Desmarest" authorityYear="1817" box="[425,591,683,709]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF41D598FE2176EAFD9FFD29" box="[425,591,683,709]" italics="true" pageId="173" pageNumber="174">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF41D598FD1976EAFCDAFD29" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[657,778,683,709]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF41D598FD1976EAFCDAFD29" box="[657,778,683,709]" italics="true" pageId="173" pageNumber="174">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in having a broader distal end, broader distal part of the lateral supracondylar ridge, and a shallower trochlea; and additionally differs from
|
||
<taxonomicName id="4C4C4D68FF41D598FE2A774AFDC6FCC9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[418,534,779,805]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF41D598FE2A774AFDC6FCC9" box="[418,534,779,805]" italics="true" pageId="173" pageNumber="174">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF41D598FF30776DFEE7F929" blockId="173.[136,779,171,2085]" pageId="173" pageNumber="174">
|
||
<emphasis id="B938EAF9FF41D598FF30776DFE66FCA9" box="[184,438,811,837]" italics="true" pageId="173" pageNumber="174">
|
||
Ulna (
|
||
<figureCitation id="13772A6EFF41D598FE9D776DFE7EFCA9" box="[277,430,811,837]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="173" pageNumber="174">Fig. 101a–i</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: moderately large, deep, transversely compressed, curved cranially in lateral view and gently medially curved in cranial view.Olecranon quite elongate, transversely compressed and tapers in height to a blunt, slightly caudally deflected in lateral view, with a squared distal end. Proximal section of the lateral surface of the ulna is flat; proximomedial flexor fossa smoothly, deeply concave. Facet for the humeral articulation is quite broad, with no visible trochlear notch; medial part deeply concave, more elongate and situated more cranially than the lateral part; lateral part relatively shallow, smoothly concave and tilted and projected laterally, with the lateral margin abraded in the sole known specimen; anconeal process low relative to the coronoid process, with an obtuse, slightly rounded mesial angle; coronoid process tall, quite narrow, and smoothly rounded in posterior view; merges gently into the shaft distally (
|
||
<figureCitation id="13772A6EFF41D598FDB1716DFD7AFAA9" box="[569,682,1323,1349]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="173" pageNumber="174">Fig. 101a</figureCitation>
|
||
). Radial facet mostly abraded, but appears roughly semicircular and abuts the lateral three-quarters of the anterior margin of the lateral part of anconeal facet. Ulnar tuberosity low, narrow, quite elongate, slightly thickened and rugose. Proximal part of the shaft is slightly crushed; interpreted as having been transversely compressed and deep; caudal margin straight beneath the olecranon and the humeral facet, curves gently cranially past the humeral facet, and straightens distally; midpoint of the cranial surface of the shaft has thin ridge tilted laterally. Distal end has a cone-shaped base, smoothly narrows to a projected, globular styloid process.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF41D598FF30728DFAB2FD29" blockId="173.[136,779,171,2085]" lastBlockId="173.[810,1452,171,1765]" pageId="173" pageNumber="174">
|
||
The ulna of
|
||
<taxonomicName id="4C4C4D68FF41D598FECB728AFE1DF909" authorityName="Plane" authorityYear="1967" box="[323,461,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF41D598FECB728AFE1DF909" box="[323,461,1739,1765]" italics="true" pageId="173" pageNumber="174">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
does not differ from that of
|
||
<taxonomicName id="4C4C4D68FF41D598FF0072AAFED8F8E9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,264,1771,1797]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF41D598FF0072AAFED8F8E9" box="[136,264,1771,1797]" italics="true" pageId="173" pageNumber="174">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
. It differs from
|
||
<taxonomicName id="4C4C4D68FF41D598FE4B72AAFDC6F8E9" authorityName=", Owen" authorityYear="1874" box="[451,534,1771,1797]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF41D598FE4B72AAFDC6F8E9" box="[451,534,1771,1797]" italics="true" pageId="173" pageNumber="174">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller and lacking a large caudomedial eminence on the olecranon, with a less deep proximal shaft and a more concave medial surface beneath the humeral facet; from
|
||
<taxonomicName id="4C4C4D68FF41D598FDFB730AFCDAF889" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[627,778,1867,1893]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF41D598FDFB730AFCDAF889" box="[627,778,1867,1893]" italics="true" pageId="173" pageNumber="174">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF41D598FF00732DFF38F869" bold="true" box="[136,232,1899,1925]" pageId="173" pageNumber="174">
|
||
<taxonomicNameLabel id="A20B5782FF41D598FF00732DFF38F869" box="[136,232,1899,1925]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being more gracile and more transversely compressed, with a relatively longer olecranon; from
|
||
<taxonomicName id="4C4C4D68FF41D598FD7D73CAFF1BF829" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF41D598FD7D73CAFF1BF829" italics="true" pageId="173" pageNumber="174">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF41D598FF5073EDFEE7F829" bold="true" box="[216,311,1963,1989]" pageId="173" pageNumber="174">
|
||
<taxonomicNameLabel id="A20B5782FF41D598FF5073EDFEE7F829" box="[216,311,1963,1989]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being generally smaller, deeper and more dorsoventrally compressed, with relatively longer olecranon, more deeply concave humeral facet, narrower coronoid process with a gentler slope anteriorly into the shaft (rather than meeting the shaft at almost a right angle), and a shallower proximomedial flexor fossa; from
|
||
<taxonomicName id="4C4C4D68FF41D598FA1A748DFC4AFEE9" baseAuthorityName="Flannery" baseAuthorityYear="1989" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF41D598FA1A748DFC4AFEE9" italics="true" pageId="173" pageNumber="174">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being generally larger, and deeper and more transversely compressed proximally, with a less cranially deflected olecranon, taller, less medially flared coronoid process, and a lower, more proximally situated cranial ridge on the shaft; from
|
||
<taxonomicName id="4C4C4D68FF41D598FBBA752DFB5DFE69" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1074,1165,363,389]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF41D598FBBA752DFB5DFE69" box="[1074,1165,363,389]" italics="true" pageId="173" pageNumber="174">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being more robust, and deeper and more transversely compressed proximally, with a less cranially deflected olecranon, less medially flared anconeal process, and a taller coronoid process; from
|
||
<taxonomicName id="4C4C4D68FF41D598FA05758AFC75FDE9" authorityName="Desmarest" authorityYear="1817" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF41D598FA05758AFC75FDE9" italics="true" pageId="173" pageNumber="174">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more robust, and deeper and more transversely compressed proximally, with a less tall olecranon, less medially flared anconeal process, and a taller coronoid process; and from
|
||
<taxonomicName id="4C4C4D68FF41D598FB44760AFA93FD89" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1228,1347,587,613]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF41D598FB44760AFA93FD89" box="[1228,1347,587,613]" italics="true" pageId="173" pageNumber="174">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a relatively longer, cranially undeflected and unprojected olecranon, lower, narrower anconeal process, and a lower, less medially flared coronoid process.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF41D598FCD2768DFAD2FAC9" blockId="173.[810,1452,171,1765]" pageId="173" pageNumber="174">
|
||
<emphasis id="B938EAF9FF41D598FCD2768DFB96FD09" box="[858,1094,715,741]" italics="true" pageId="173" pageNumber="174">
|
||
Radius (
|
||
<figureCitation id="13772A6EFF41D598FC31768DFBEEFD09" box="[953,1086,715,741]" captionStart="FIGURE 100" captionStartId="172.[136,235,1900,1924]" captionTargetBox="[162,1425,212,1869]" captionTargetId="figure-27@172.[151,1436,201,1880]" captionTargetPageId="172" captionText="FIGURE 100. surface scan images of right upper forelimb elements of P. otibandus: (a–c) clavicle of UCMP 70045 in (a) anterior, (b) dorsal, and (c) ventral views; (d–e) distal humeral fragment of UCMP 70059 in (d) caudal, and (e) cranial views; and (f–g) radius of UCMP 70059 in (f) cranial, and (g) caudal views." figureDoi="http://doi.org/10.5281/zenodo.10993996" httpUri="https://zenodo.org/record/10993996/files/figure.png" pageId="173" pageNumber="174">Fig. 100f–g</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: elongate, with a gently arched shaft curving mediocaudally past the radial tubercle. Radial head broad, oval and smoothly concave. Radial neck narrows to the radial tubercule. Radial tubercule rugose, oval and smoothly projected. Shaft cross-section oval immediately distal to the radial tubercle, quickly transitions to strongly craniomedially-caudolaterally compressed by the midpoint; gently broadens distally. Cranial ridge short, quite thin and located immediately proximal to the midpoint of the shaft; caudal ridge relatively thinner and more raised on the caudomedial to caudal surface. Distal end large and craniocaudally compressed with a small tubercle on the cranial surface; scaphoidal facet broad, slightly caudally tilted and gently concave, located caudal to a low, shallow fossa across the cranial section; styloid process broad, craniocaudally compressed and tapering to a rounded tip. Ulnar notch broad, very shallowly concave and elongate, slowly tapers proximally along the caudolateral surface.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF41D598FCD2716DFA4CF909" blockId="173.[810,1452,171,1765]" pageId="173" pageNumber="174">
|
||
The radius of
|
||
<taxonomicName id="4C4C4D68FF41D598FC78716AFBA8FAA9" authorityName="Plane" authorityYear="1967" box="[1008,1144,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF41D598FC78716AFBA8FAA9" box="[1008,1144,1323,1349]" italics="true" pageId="173" pageNumber="174">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
cannot be not differentiated from that of
|
||
<taxonomicName id="4C4C4D68FF41D598FC33710AFBEAFA89" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[955,1082,1355,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF41D598FC33710AFBEAFA89" box="[955,1082,1355,1381]" italics="true" pageId="173" pageNumber="174">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
. It differs from
|
||
<taxonomicName id="4C4C4D68FF41D598FB7B710AFA96FA89" authorityName=", Owen" authorityYear="1874" box="[1267,1350,1355,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF41D598FB7B710AFA96FA89" box="[1267,1350,1355,1381]" italics="true" pageId="173" pageNumber="174">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being generally smaller, with the distal end more craniocaudally compressed; from
|
||
<taxonomicName id="4C4C4D68FF41D598FB8171CAFB72FA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1033,1186,1419,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF41D598FB8171CAFB72FA49" box="[1033,1186,1419,1445]" italics="true" pageId="173" pageNumber="174">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF41D598FB3871CDFAC1FA49" bold="true" box="[1200,1297,1419,1445]" pageId="173" pageNumber="174">
|
||
<taxonomicNameLabel id="A20B5782FF41D598FB3871CDFAC1FA49" box="[1200,1297,1419,1445]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a less raised cranial ridge and a shallower proximal shaft; from
|
||
<taxonomicName id="4C4C4D68FF41D598FCEF718AFC16FA09" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[871,966,1484,1509]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="viator">
|
||
<emphasis id="B938EAF9FF41D598FCEF718AFC16FA09" box="[871,966,1484,1509]" italics="true" pageId="173" pageNumber="174">P. viator</emphasis>
|
||
</taxonomicName>
|
||
in being shorter, with a more raised caudal ridge and a less proximally situated cranial ridge; from
|
||
<taxonomicName id="4C4C4D68FF41D598FCA2724DFC13F9C9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[810,963,1547,1573]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF41D598FCA2724DFC13F9C9" box="[810,963,1547,1573]" italics="true" pageId="173" pageNumber="174">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having a more raised cranial ridge, less elongate caudal ridge, and a caudally (rather than cranially) tilted scaphoidal facet; from
|
||
<taxonomicName id="4C4C4D68FF41D598FB60720DFA96F989" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1256,1350,1611,1637]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF41D598FB60720DFA96F989" box="[1256,1350,1611,1637]" italics="true" pageId="173" pageNumber="174">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being more robust, with a larger radial tubercle, and more raised caudal ridge; from
|
||
<taxonomicName id="4C4C4D68FF41D598FB8F72CAFB7AF949" authorityName="Desmarest" authorityYear="1817" box="[1031,1194,1675,1701]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF41D598FB8F72CAFB7AF949" box="[1031,1194,1675,1701]" italics="true" pageId="173" pageNumber="174">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being more robust, with a relatively larger proximal epiphysis and a more raised caudal ridge; and from
|
||
<taxonomicName id="4C4C4D68FF41D598FBF1728AFB3DF909" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1145,1261,1739,1765]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="173" pageNumber="174" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF41D598FBF1728AFB3DF909" box="[1145,1261,1739,1765]" italics="true" pageId="173" pageNumber="174">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF41D598FCA2734DFCAFF8C9" blockId="173.[810,1452,1803,2085]" box="[810,895,1803,1829]" pageId="173" pageNumber="174">
|
||
<emphasis id="B938EAF9FF41D598FCA2734DFCAFF8C9" bold="true" box="[810,895,1803,1829]" italics="true" pageId="173" pageNumber="174">
|
||
<collectingRegion id="4988F809FF41D598FCA2734DFCAFF8C9" box="[810,895,1803,1829]" country="Papua New Guinea" name="Manus" pageId="173" pageNumber="174">Manus</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF41D59BFCA2736DFDA1FDA9" blockId="173.[810,1452,1803,2085]" lastBlockId="174.[136,779,171,2085]" lastPageId="174" lastPageNumber="175" pageId="173" pageNumber="174">
|
||
<emphasis id="B938EAF9FF41D598FCA2736DFB88F8A9" box="[810,1112,1835,1861]" italics="true" pageId="173" pageNumber="174">
|
||
Scaphoid (
|
||
<figureCitation id="13772A6EFF41D598FC2F736DFB9EF8A9" box="[935,1102,1835,1861]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="173" pageNumber="174">Fig. 101e–g, i</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: broad, roughly semicircular in dorsal view with a broadly convex proximal surface; dorsopalmarly shortens medial to the midpoint. Radial facet broad and strongly, smoothly convex, extends from posteriorly adjacent to the triquetral facet, across the posterior surface, to the posterior margin of the palmar process; becomes gently concave medially due to a low, rounded eminence on the medial margin and a broad, raised, anteroposteriorly short crest located slightly medial of the centre of the dorsal surface, for articulation with the styloid process of radius. Palmar process broad, tapers in dorsal view to a rounded tip, slightly anteroposteriorly compressed and curves distinctly palmarly toward the tip. Facet for the hamatum large, covers anterolateral surface. Facet for the capitatum slightly broader than the hamatal facet; covers the centre of the anterior surface. Facet for the trapezoid palmarly convex and dorsally concave beneath anterior of dorsal crest, abuts capitatum facet on anterodorsal surface of palmar process. Facet for the trapezium tall, rounded and convex, wraps around the anteromedial surface of the palmar process.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFF30760DFEE6FB89" blockId="174.[136,779,171,2085]" pageId="174" pageNumber="175">
|
||
The scaphoid of
|
||
<taxonomicName id="4C4C4D68FF42D59BFEF2760AFDD6FD89" authorityName="Plane" authorityYear="1967" box="[378,518,587,613]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF42D59BFEF2760AFDD6FD89" box="[378,518,587,613]" italics="true" pageId="174" pageNumber="175">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF42D59BFD7D760AFF2DFD69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF42D59BFD7D760AFF2DFD69" italics="true" pageId="174" pageNumber="175">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF42D59BFE80762DFEB6FD69" bold="true" box="[264,358,619,645]" pageId="174" pageNumber="175">
|
||
<taxonomicNameLabel id="A20B5782FF42D59BFE80762DFEB6FD69" box="[264,358,619,645]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a slightly more palmarly curved and anteriorly deflected palmar process, with a shallower fossa on the medial part of the palmar surface; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFF42768AFEFDFD09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[202,301,716,741]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF42D59BFF42768AFEFDFD09" box="[202,301,716,741]" italics="true" pageId="174" pageNumber="175">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF42D59BFEB1768DFE47FD09" bold="true" box="[313,407,715,741]" pageId="174" pageNumber="175">
|
||
<taxonomicNameLabel id="A20B5782FF42D59BFEB1768DFE47FD09" box="[313,407,715,741]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a raised dorsal ridge, a taller, more convex radial facet, larger medial eminence, and a less dorsopalmarly compressed palmar process; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFEF5776DFDDDFCA9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[381,525,811,837]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF42D59BFEF5776DFDDDFCA9" box="[381,525,811,837]" italics="true" pageId="174" pageNumber="175">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being broader, with a raised dorsal ridge and a broader, more anteropalmarly deflected palmar process; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFE62772DFD96FC69" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[490,582,875,901]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF42D59BFE62772DFD96FC69" box="[490,582,875,901]" italics="true" pageId="174" pageNumber="175">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being broader, with a larger palmomedial fossa and a more dorsopalmarly compressed palmar process; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFD8077EAFD78FC29" authorityName="Desmarest" authorityYear="1817" box="[520,680,939,965]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF42D59BFD8077EAFD78FC29" box="[520,680,939,965]" italics="true" pageId="174" pageNumber="175">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being relatively broader, with a broader, more smoothly convex radial facet and a less deep, more palmarly curved palmar process; and from
|
||
<taxonomicName id="4C4C4D68FF42D59BFED5704AFE03FBC9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[349,467,1035,1061]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF42D59BFED5704AFE03FBC9" box="[349,467,1035,1061]" italics="true" pageId="174" pageNumber="175">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, with a low, rounded eminence present on the medial margin of the radial facet.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFF30702DFE00FAC9" blockId="174.[136,779,171,2085]" pageId="174" pageNumber="175">
|
||
<emphasis id="B938EAF9FF42D59BFF30702DFE67FB69" box="[184,439,1131,1157]" italics="true" pageId="174" pageNumber="175">
|
||
Pisiform (
|
||
<figureCitation id="13772A6EFF42D59BFEA3702DFE7FFB69" box="[299,431,1131,1157]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="174" pageNumber="175">Fig. 101e–i</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: quite elongate and distinctly dorsopalmarly flattened; large, squared proximal end constricts to a narrow waist and flares anteroposteriorly to the thickened, rounded distal end. The posteromedial facet for the styloid process of the ulna is small, located on the posteromedial margin.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFF30716DFE9DF969" blockId="174.[136,779,171,2085]" pageId="174" pageNumber="175">
|
||
The pisiform of
|
||
<taxonomicName id="4C4C4D68FF42D59BFE0B716AFDC3FAA9" authorityName="Plane" authorityYear="1967" box="[387,531,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF42D59BFE0B716AFDC3FAA9" box="[387,531,1323,1349]" italics="true" pageId="174" pageNumber="175">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF42D59BFF00710AFF3CFA89" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,236,1356,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="viator">
|
||
<emphasis id="B938EAF9FF42D59BFF00710AFF3CFA89" box="[136,236,1356,1381]" italics="true" pageId="174" pageNumber="175">P. viator</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with a more dorsopalmarly compressed proximal end and a much smaller proximal ulnar facet; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFEC771CDFE31FA49" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[335,481,1419,1445]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF42D59BFEC771CDFE31FA49" box="[335,481,1419,1445]" italics="true" pageId="174" pageNumber="175">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger relative to the other carpals and more elongate, with a narrower waist and a more dorsopalmarly compressed distal end; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFF0071ADFF35F9E9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[136,229,1515,1541]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF42D59BFF0071ADFF35F9E9" box="[136,229,1515,1541]" italics="true" pageId="174" pageNumber="175">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF42D59BFE9571AAFE6EF9E9" authorityName="Desmarest" authorityYear="1817" box="[285,446,1515,1541]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF42D59BFE9571AAFE6EF9E9" box="[285,446,1515,1541]" italics="true" pageId="174" pageNumber="175">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being more elongate, with a narrower waist and a more dorsopalmarly compressed base lacking an anterodorsal tubercle; and from
|
||
<taxonomicName id="4C4C4D68FF42D59BFD10726AFCDAF9A9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[664,778,1579,1605]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF42D59BFD10726AFCDAF9A9" box="[664,778,1579,1605]" italics="true" pageId="174" pageNumber="175">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more robust, with a relatively larger, deeper distal end.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFF3072CAFD55F889" blockId="174.[136,779,171,2085]" pageId="174" pageNumber="175">
|
||
<emphasis id="B938EAF9FF42D59BFF3072CAFE08F949" box="[184,472,1675,1701]" italics="true" pageId="174" pageNumber="175">
|
||
Triquetrum (
|
||
<figureCitation id="13772A6EFF42D59BFEC272CDFE1FF949" box="[330,463,1675,1701]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="174" pageNumber="175">Fig. 101e–i</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: broad and blocky; dorsal surface gently convex with small ridges along the anterior and posterior margins.The palmar surface forms a rounded point at the posterior margin. Facet for the styloid process of the ulna rounded and concave. Facet for the pisiform long and dorsopalmarly short. Facet for the hamatum large, occupying most of the anterior surface.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFF30732DFABFFEA9" blockId="174.[136,779,171,2085]" lastBlockId="174.[810,1452,171,2053]" pageId="174" pageNumber="175">
|
||
The triquetrum of
|
||
<taxonomicName id="4C4C4D68FF42D59BFE19732AFDCEF869" authorityName="Plane" authorityYear="1967" box="[401,542,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF42D59BFE19732AFDCEF869" box="[401,542,1899,1925]" italics="true" pageId="174" pageNumber="175">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF42D59BFF0073CAFECCF849" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,284,1931,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF42D59BFF0073CAFECCF849" box="[136,284,1931,1957]" italics="true" pageId="174" pageNumber="175">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF42D59BFEAE73CDFE54F849" bold="true" box="[294,388,1931,1957]" pageId="174" pageNumber="175">
|
||
<taxonomicNameLabel id="A20B5782FF42D59BFEAE73CDFE54F849" box="[294,388,1931,1957]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a flatter dorsal surface and a slightly smaller pisiform facet; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFDFF73EDFCDAF829" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[631,778,1963,1989]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF42D59BFDFF73EDFCDAF829" box="[631,778,1963,1989]" italics="true" pageId="174" pageNumber="175">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger relative to the other carpals, with a broader and flatter dorsal surface and a slightly more laterally and less palmarly situated facet for the pisiform; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFD667C4DFCB4FF29" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[750,868,171,2085]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF42D59BFD667C4DFCB4FF29" box="[750,868,171,2085]" italics="true" pageId="174" pageNumber="175">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in having flatter dorsal surface, with a smaller and more palmarly situated facet for the pisiform; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFA05748AFC75FEE9" authorityName="Desmarest" authorityYear="1817" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF42D59BFA05748AFC75FEE9" italics="true" pageId="174" pageNumber="175">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being deeper, with a flatter dorsal surface; and from
|
||
<taxonomicName id="4C4C4D68FF42D59BFC2A754AFBCCFEC9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[930,1052,267,293]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF42D59BFC2A754AFBCCFEC9" box="[930,1052,267,293]" italics="true" pageId="174" pageNumber="175">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being absolutely larger, larger relative to the size of the pisiform, and more robust.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFCD2750AFBD6FC49" blockId="174.[810,1452,171,2053]" pageId="174" pageNumber="175">
|
||
<emphasis id="B938EAF9FF42D59BFCD2750AFBABFE89" box="[858,1147,331,357]" italics="true" pageId="174" pageNumber="175">
|
||
Hamatum (
|
||
<figureCitation id="13772A6EFF42D59BFC6A750DFBA3FE89" box="[994,1139,331,357]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="174" pageNumber="175">Fig. 101e–h</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: quite deep and blocky, with a narrow, blunted palmar process projecting palmomedially from the posteromedial edge; dorsal surface is gently convex; a small, rounded eminence projects posteromedially. Facet for the triquetrum quite broad and mostly posterior facing, occupies the lateral part of the posterior surface except for on the palmar process; a small posterior part of the facet extends onto the posteromedial process of the hamatum, with the medial margin distinct and separate from the facet for the scaphoid. Facet for the scaphoid small and concave, on posteromedial surface. Facet for the capitatum small, narrow and facing anteromedially, with slight dorsal lip. Facet for metacarpal III narrow and faces anteromedially. Facet for metacarpal IV broad, narrows palmarly onto the medial base of the palmar process. Facet for metacarpal V quite large and anterolaterally facing, covers the majority of the anterior surface of the palmar process, with a rounded palmar lip.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFCD277EDFC0AF9E9" blockId="174.[810,1452,171,2053]" pageId="174" pageNumber="175">
|
||
The hamatum of
|
||
<taxonomicName id="4C4C4D68FF42D59BFB9777EAFB7AFC29" authorityName="Plane" authorityYear="1967" box="[1055,1194,939,965]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF42D59BFB9777EAFB7AFC29" box="[1055,1194,939,965]" italics="true" pageId="174" pageNumber="175">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF42D59BFA1F77EAFC4FFC09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF42D59BFA1F77EAFC4FFC09" italics="true" pageId="174" pageNumber="175">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF42D59BFC2E778DFBD1FC09" bold="true" box="[934,1025,971,997]" pageId="174" pageNumber="175">
|
||
<taxonomicNameLabel id="A20B5782FF42D59BFC2E778DFBD1FC09" box="[934,1025,971,997]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and deeper, with the triquetral facet distinctly separate from the scaphoidal facet, and a smaller, posteropalmarly projected palmar process; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFC5B706AFBE6FBA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[979,1078,1068,1093]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF42D59BFC5B706AFBE6FBA9" box="[979,1078,1068,1093]" italics="true" pageId="174" pageNumber="175">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF42D59BFBC9706DFB4FFBA9" bold="true" box="[1089,1183,1067,1093]" pageId="174" pageNumber="175">
|
||
<taxonomicNameLabel id="A20B5782FF42D59BFBC9706DFB4FFBA9" box="[1089,1183,1067,1093]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a smaller triquetral facet that is less deeply inset, less extensive medially, and distinctly separate from the scaphoidal facet; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFBB970CDFB14FB49" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1073,1220,1163,1189]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF42D59BFBB970CDFB14FB49" box="[1073,1220,1163,1189]" italics="true" pageId="174" pageNumber="175">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having a broader triquetral facet and larger facets for the capitatum and metacarpal III; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFB95708DFBADFB09" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1053,1149,1227,1253]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF42D59BFB95708DFBADFB09" box="[1053,1149,1227,1253]" italics="true" pageId="174" pageNumber="175">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in having a slightly more palmarly situated triquetral facet and a less anteriorly deflected and extensive palmar process; from
|
||
<taxonomicName id="4C4C4D68FF42D59BFA05714AFC75FAA9" authorityName="Desmarest" authorityYear="1817" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF42D59BFA05714AFC75FAA9" italics="true" pageId="174" pageNumber="175">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly deeper, with a more laterally situated triquetral facet relative to the scaphoidal facet, and a less palmarly projected, much less anteriorly curved palmar process; and from
|
||
<taxonomicName id="4C4C4D68FF42D59BFBDD71CAFB1BFA49" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1109,1227,1419,1445]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="174" pageNumber="175" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF42D59BFBDD71CAFB1BFA49" box="[1109,1227,1419,1445]" italics="true" pageId="174" pageNumber="175">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and deeper, with the scaphoidal and triquetral facets distinct rather than abutting, and a broader, less palmarly projected palmar process.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF42D59BFCD2724DFC37F7E9" blockId="174.[810,1452,171,2053]" pageId="174" pageNumber="175">
|
||
<emphasis id="B938EAF9FF42D59BFCD2724DFBA2F9C9" box="[858,1138,1547,1573]" italics="true" pageId="174" pageNumber="175">
|
||
Capitatum (
|
||
<figureCitation id="13772A6EFF42D59BFC69724DFBBAF9C9" box="[993,1130,1547,1573]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="174" pageNumber="175">Fig. 101e–g</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: tall and blocky, with a flat dorsal surface and a rugose, slightly concave, roughly triangular palmar surface. Posteropalmar process very slight to absent. Facet for the scaphoid located on the medial component of the posterior surface. Facet for the triquetrum slightly larger than the scaphoidal facet, situated on the posterolateral surface. Facet for metacarpal III broad, with semi-distinct anterior and anterolateral components; a small, dorsally opening mesial fossa on a slight corner separates the slightly broader anterolateral facet from the smaller anterior facet. Facet for the posterolateral articulation of metacarpal II small, quite tall and narrow, medially abuts the anterior section of the facet for metacarpal III. Facet for the trapezoid tall, narrow and gently concave, anteromedially abuts the scaphoidal facet.
|
||
</paragraph>
|
||
<caption id="DF336663FF43D59AFF0073CAFE59F7E8" ID-DOI="http://doi.org/10.5281/zenodo.10993998" ID-Zenodo-Dep="10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="175" pageNumber="176" startId="175.[136,235,1932,1956]" targetBox="[195,1391,177,1902]" targetPageId="175" targetType="figure">
|
||
<paragraph id="8BF336EBFF43D59AFF0073CAFE59F7E8" blockId="175.[136,1452,1932,2053]" pageId="175" pageNumber="176">
|
||
<emphasis id="B938EAF9FF43D59AFF0073CAFECDF849" bold="true" box="[136,285,1932,1957]" pageId="175" pageNumber="176">FIGURE 101.</emphasis>
|
||
surface scan images of right lower forelimb elements of
|
||
<taxonomicName id="4C4C4D68FF43D59AFCD373CBFBF5F849" authority="UCMP" authorityName="UCMP" box="[859,1061,1933,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="175" pageNumber="176" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF43D59AFCD373CBFC08F849" box="[859,984,1933,1957]" italics="true" pageId="175" pageNumber="176">P. otibandus</emphasis>
|
||
UCMP
|
||
</taxonomicName>
|
||
70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF5CD585FF3074EDFDC1FDC9" blockId="176.[136,779,171,2085]" pageId="176" pageNumber="177">
|
||
The capitatum of
|
||
<taxonomicName id="4C4C4D68FF5CD585FE0474EAFDCAFF29" box="[396,538,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5CD585FE0474EAFDCAFF29" box="[396,538,171,197]" italics="true" pageId="176" pageNumber="177">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF5CD585FF00748AFECEFF09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,286,203,229]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5CD585FF00748AFECEFF09" box="[136,286,203,229]" italics="true" pageId="176" pageNumber="177">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5CD585FEA3748DFE5AFF09" bold="true" box="[299,394,203,229]" pageId="176" pageNumber="177">
|
||
<taxonomicNameLabel id="A20B5782FF5CD585FEA3748DFE5AFF09" box="[299,394,203,229]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF5CD585FE44748AFDE0FF09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[460,560,204,229]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5CD585FE44748AFDE0FF09" box="[460,560,204,229]" italics="true" pageId="176" pageNumber="177">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5CD585FDB5748DFD4CFF09" bold="true" box="[573,668,203,229]" pageId="176" pageNumber="177">
|
||
<taxonomicNameLabel id="A20B5782FF5CD585FDB5748DFD4CFF09" box="[573,668,203,229]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and lacking a blunt posteropalmar process; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FF00754DFEC8FEC9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[136,280,267,293]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF5CD585FF00754DFEC8FEC9" box="[136,280,267,293]" italics="true" pageId="176" pageNumber="177">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being generally squarer and less rounded, lacking a posteropalmar process and in having the dorsal surface flat instead of convex; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FDB4750DFD4EFE89" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[572,670,331,357]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5CD585FDB4750DFD4EFE89" box="[572,670,331,357]" italics="true" pageId="176" pageNumber="177">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly anteroposteriorly deeper; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FDC1752AFD3BFE69" authorityName="Desmarest" authorityYear="1817" box="[585,747,363,389]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5CD585FDC1752AFD3BFE69" box="[585,747,363,389]" italics="true" pageId="176" pageNumber="177">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being squarer in dorsal view, with a more medially and less anteriorly facing facet for metacarpal II and a less medially projected facet for the trapezoid; and from
|
||
<taxonomicName id="4C4C4D68FF5CD585FD66758AFF09FDE9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5CD585FD66758AFF09FDE9" italics="true" pageId="176" pageNumber="177">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, deeper, and squarer in dorsal view, with a larger, deeper hamatal facet.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FF30766DFE07FAE9" blockId="176.[136,779,171,2085]" pageId="176" pageNumber="177">
|
||
<emphasis id="B938EAF9FF5CD585FF30766DFE3AFDA9" box="[184,490,555,581]" italics="true" pageId="176" pageNumber="177">
|
||
Metacarpals (
|
||
<figureCitation id="13772A6EFF5CD585FED2766DFE32FDA9" box="[346,482,555,581]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="176" pageNumber="177">Fig. 101e–i</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: short and quite robust, with metacarpals II–IV relatively more elongate; each shaft narrows to a distinct waist; tall proximally, becoming dorsopalmarly compressed distally, particularly II–IV; all with large, broad and rounded distal keels. Metacarpal I: shortest, with a tall, narrow facet for the trapezium laterally situated on the proximal surface; a large, blunt tubercle on the palmomedial surface of the proximal end projects medially; metacarpals I and V have distinctly asymmetrical distal ends, skewed mesially. Metacarpal II: substantially shorter than III and IV, with the lateral surface of the proximal end articulating against the anterolateral surface of the capitatum and the lateral surface of the proximal end of metacarpal III; distal end has a distinctly enlarged lateral section. Metacarpal III: longest, with the proximal end separated into a longer, medially deflected medial process and a short, laterally deflected lateral process. Metacarpal IV: has a broad proximal facet for articulation with the capitatum, tilted dorsally and slightly medially, with the dorsal margin distinctly concave in dorsal view. Metacarpal V: proximal end has a large, blunt tubercle on the palmolateral surface that is strongly projected laterally; shaft curves slightly palmarly.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FF30714DFEC4F989" blockId="176.[136,779,171,2085]" pageId="176" pageNumber="177">
|
||
The metacarpals of
|
||
<taxonomicName id="4C4C4D68FF5CD585FE10714AFDF3FAC9" box="[408,547,1291,1317]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5CD585FE10714AFDF3FAC9" box="[408,547,1291,1317]" italics="true" pageId="176" pageNumber="177">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differ from those of
|
||
<taxonomicName id="4C4C4D68FF5CD585FF00716AFF0CFAA9" authorityName=", Owen" authorityYear="1874" box="[136,220,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF5CD585FF00716AFF0CFAA9" box="[136,220,1323,1349]" italics="true" pageId="176" pageNumber="177">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being slightly smaller; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FDC5716AFD0FFAA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[589,735,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5CD585FDC5716AFD0FFAA9" box="[589,735,1323,1349]" italics="true" pageId="176" pageNumber="177">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5CD585FD60716DFF69FA89" bold="true" pageId="176" pageNumber="177">
|
||
<taxonomicNameLabel id="A20B5782FF5CD585FD60716DFF69FA89" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and slightly more gracile; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FD7D710AFF1BFA69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5CD585FD7D710AFF1BFA69" italics="true" pageId="176" pageNumber="177">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5CD585FF5A712DFEFBFA69" bold="true" box="[210,299,1387,1413]" pageId="176" pageNumber="177">
|
||
<taxonomicNameLabel id="A20B5782FF5CD585FF5A712DFEFBFA69" box="[210,299,1387,1413]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and more robust; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FD78712DFF28FA49" baseAuthorityName="Flannery" baseAuthorityYear="1989" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF5CD585FD78712DFF28FA49" italics="true" pageId="176" pageNumber="177">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being more elongate, with shafts with more pronounced waists, generally broader proximal articular surfaces, flatter and less convex distal articular surfaces and generally squarer distal ends; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FDC171ADFD76F9E9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[585,678,1515,1541]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5CD585FDC171ADFD76F9E9" box="[585,678,1515,1541]" italics="true" pageId="176" pageNumber="177">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being relatively slightly broader; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FD88724AFD73F9C9" authorityName="Desmarest" authorityYear="1817" box="[512,675,1547,1573]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5CD585FD88724AFD73F9C9" box="[512,675,1547,1573]" italics="true" pageId="176" pageNumber="177">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly more elongate; and from
|
||
<taxonomicName id="4C4C4D68FF5CD585FDA8726AFD4AF9A9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[544,666,1579,1605]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5CD585FDA8726AFD4AF9A9" box="[544,666,1579,1605]" italics="true" pageId="176" pageNumber="177">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FF30722DFD28F7E9" blockId="176.[136,779,171,2085]" pageId="176" pageNumber="177">
|
||
<emphasis id="B938EAF9FF5CD585FF30722DFD9FF969" box="[184,591,1643,1669]" italics="true" pageId="176" pageNumber="177">
|
||
Manual phalanges (
|
||
<figureCitation id="13772A6EFF5CD585FE3F722DFD98F969" box="[439,584,1643,1669]" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="176" pageNumber="177">Fig. 101e–i</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: short, broad, dorsopalmarly compressed and quite symmetrical, particularly proximal and middle phalanges; proximal phalanx II more gracile than other proximal phalanges; proximal phalanges slightly shorter than the corresponding distal phalanges. Proximal phalanges: palmar tubercles raised, rounded and proximally projected; trochlea moderately shallow and quite narrow. Middle phalanges: very short, broad and dorsopalmarly compressed; shafts have very slight waists. Distal phalanges: quite long, with the palmar tubercle large and palmodistally projected (
|
||
<figureCitation id="13772A6EFF5CD585FD5573EDFF12F80A" captionStart="FIGURE 101" captionStartId="175.[136,235,1932,1956]" captionTargetBox="[195,1391,177,1902]" captionTargetId="figure-17@175.[185,1401,172,1910]" captionTargetPageId="175" captionText="FIGURE 101. surface scan images of right lower forelimb elements of P. otibandus UCMP 70059: (a–d) partial ulna in (a) lateral, (b) medial, (c) caudal, and (d) cranial views; and (e–i) partial articulated manus, with ulnar styloid process attached, in (e) dorsal, (f) dorsolateral, (g) palmar, (h) lateral, and (i) medial views. Elements not preserved in the manus are the trapezoid, trapezium, and distal phalanges I and IV." figureDoi="http://doi.org/10.5281/zenodo.10993998" httpUri="https://zenodo.org/record/10993998/files/figure.png" pageId="176" pageNumber="177">Fig. 101h</figureCitation>
|
||
),and the shaft palmarly curved,slightly dorsopalmarly compressed and smoothly rounded on the dorsal surface.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FF307C4DFA88FC89" blockId="176.[136,779,171,2085]" lastBlockId="176.[810,1452,171,869]" pageId="176" pageNumber="177">
|
||
The manual phalanges of
|
||
<taxonomicName id="4C4C4D68FF5CD585FE437C4AFD80F7C9" box="[459,592,2059,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5CD585FE437C4AFD80F7C9" box="[459,592,2059,2085]" italics="true" pageId="176" pageNumber="177">P.otibandus</emphasis>
|
||
</taxonomicName>
|
||
differ from those of
|
||
<taxonomicName id="4C4C4D68FF5CD585FCC374EAFC70FF29" authorityName=", Owen" authorityYear="1874" box="[843,928,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF5CD585FCC374EAFC70FF29" box="[843,928,171,197]" italics="true" pageId="176" pageNumber="177">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with more palmarly curved distal phalanges; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FBAC748AFB61FF09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1060,1201,203,229]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5CD585FBAC748AFB61FF09" box="[1060,1201,203,229]" italics="true" pageId="176" pageNumber="177">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5CD585FB3F748DFADFFF09" bold="true" box="[1207,1295,203,229]" pageId="176" pageNumber="177">
|
||
<taxonomicNameLabel id="A20B5782FF5CD585FB3F748DFADFFF09" box="[1207,1295,203,229]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having less dorsopalmar compression, longer, narrower proximal and middle shafts, and more palmarly curved distal phalangeal shafts; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FC40756AFBFEFEA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[968,1070,300,325]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5CD585FC40756AFBFEFEA9" box="[968,1070,300,325]" italics="true" pageId="176" pageNumber="177">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5CD585FBB6756DFB4FFEA9" bold="true" box="[1086,1183,299,325]" pageId="176" pageNumber="177">
|
||
<taxonomicNameLabel id="A20B5782FF5CD585FBB6756DFB4FFEA9" box="[1086,1183,299,325]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and more gracile, with less dorsopalmarly compressed distal phalanges with more projected palmar tubercles; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FCA275CDFC6EFE49" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[810,958,395,421]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF5CD585FCA275CDFC6EFE49" box="[810,958,395,421]" italics="true" pageId="176" pageNumber="177">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being broader, far more dorsopalmarly compressed, particularly the middle and distal phalanges, and significantly shorter relative to the metacarpals, with broader, less V-shaped trochleae, more dorsally tilted proximal surfaces of proximal phalanges, and dorsally rounded distal phalanges with less palmar curvature of the shaft; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FC69760DFB90FD89" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[993,1088,587,613]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5CD585FC69760DFB90FD89" box="[993,1088,587,613]" italics="true" pageId="176" pageNumber="177">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly broader, with proximal phalanges having a broader distal end relative to the proximal end, and distal phalanges having longer, more dorsopalmarly compressed and palmarly curved shafts; from
|
||
<taxonomicName id="4C4C4D68FF5CD585FC37768AFBB3FD09" authorityName="Desmarest" authorityYear="1817" box="[959,1123,715,741]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5CD585FC37768AFBB3FD09" box="[959,1123,715,741]" italics="true" pageId="176" pageNumber="177">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in having the proximal and middle phalanges with slightly more dorsally tilted proximal surfaces and less V-shaped trochleae, and more dorsopalmarly compressed distal phalanges with palmarly curved shafts; and from
|
||
<taxonomicName id="4C4C4D68FF5CD585FBBF770AFB7AFC89" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1079,1194,843,869]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5CD585FBBF770AFB7AFC89" box="[1079,1194,843,869]" italics="true" pageId="176" pageNumber="177">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FCA277CCFC4EFC49" blockId="176.[810,1452,906,2085]" box="[810,926,906,933]" pageId="176" pageNumber="177">
|
||
<emphasis id="B938EAF9FF5CD585FCA277CCFC4EFC49" bold="true" box="[810,926,906,933]" italics="true" pageId="176" pageNumber="177">Hindlimb</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FCA277EAFA5AFA09" blockId="176.[810,1452,906,2085]" pageId="176" pageNumber="177">
|
||
<emphasis id="B938EAF9FF5CD585FCA277EAFBC7FC29" box="[810,1047,939,965]" italics="true" pageId="176" pageNumber="177">
|
||
Femur (
|
||
<figureCitation id="13772A6EFF5CD585FC0E77EDFBC0FC29" box="[902,1040,939,965]" captionStart="FIGURE 102" captionStartId="177.[136,235,1932,1956]" captionTargetBox="[207,1381,182,1900]" captionTargetId="figure-17@177.[197,1391,172,1910]" captionTargetPageId="177" captionText="FIGURE 102. surface scan images of partial hindlimb elements of P. otibandus: (a) proximal right femoral fragment of UCMP 70665 in dorsal view; (b–c) partial proximal right femur of UCMP 45344 in (b) dorsal, and (c) ventral views; (d–e) left distal femoral epiphysis of UCMP 45344 in (d) distal, and (e) lateral views; (f–i) partial proximal left tibia of UCMP 70584 in (f) lateral, (g) medial, (h) cranial, and (i) proximal views; and (j–k) partial right tibial shaft in (j) lateral, and (k) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994000" httpUri="https://zenodo.org/record/10994000/files/figure.png" pageId="176" pageNumber="177">Fig. 102a–e</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: mid-sized and gracile. Head large, very broad, rounded, hemispherical, dorsomedially projected and situated quite close to the medial base of the greater trochanter with the thick epiphyseal surface continuous between the two; rotated distally such that the articular surface faces slightly medially; the base of the trochanter is deep. Proximal end fairly broad, with the dorsal surface convex in the mesial third. Greater trochanter large, blunt, transversely compressed and slightly medially deflected; thickened on the dorsal surface of the base. Greater trochanteric ridge and trochanteric fossa poorly preserved in available specimens; appear elongate in UCMP 45344, with a deep fossa in UCMP 70065. Intertrochanteric crest quite thick, moderately raised and curves distally with slight medial deflection; ventromedially orientated in cross-section; abuts the small, rugose lesser trochanter. Lesser trochanteric ridge thick, raised, distally elongate and ventromedially projected.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5CD585FCD271ADFAACF7C9" blockId="176.[810,1452,906,2085]" pageId="176" pageNumber="177">
|
||
Shaft straight, round in cross-section, with the caudal surface flattened and broadened distally for the insertion of the ischiatic portion of the mm. adductores. The quadratus tubercule is situated around the midpoint of the ventral surface of the shaft; raised and oval. Distolateral fossa for the m. flexor digitorum superficialis elongate, shallow and lateral-facing. Distal end tall and robust. Lateral trochlear crest broad and rounded relative to the medial crest, but subequal in height. Trochlea deep and approaches Vshaped (
|
||
<figureCitation id="13772A6EFF5CD585FC02734DFC2EF8CA" box="[906,1022,1803,1830]" captionStart="FIGURE 102" captionStartId="177.[136,235,1932,1956]" captionTargetBox="[207,1381,182,1900]" captionTargetId="figure-17@177.[197,1391,172,1910]" captionTargetPageId="177" captionText="FIGURE 102. surface scan images of partial hindlimb elements of P. otibandus: (a) proximal right femoral fragment of UCMP 70665 in dorsal view; (b–c) partial proximal right femur of UCMP 45344 in (b) dorsal, and (c) ventral views; (d–e) left distal femoral epiphysis of UCMP 45344 in (d) distal, and (e) lateral views; (f–i) partial proximal left tibia of UCMP 70584 in (f) lateral, (g) medial, (h) cranial, and (i) proximal views; and (j–k) partial right tibial shaft in (j) lateral, and (k) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994000" httpUri="https://zenodo.org/record/10994000/files/figure.png" pageId="176" pageNumber="177">Fig. 102d</figureCitation>
|
||
). Intercondylar fossa large and deep, extends dorsodistally to just past the midpoint of the distal surface. Condyles broad, subequal in width and gently convex ventrally. Lateral epicondyle tall, rounded and laterally very projected; lateral gastrocnemial fossa deep and rounded. Fibular facet quite broad, laterally projected and slightly caudally deflected. Medial epicondyle has a slight lateral projection of the ventral component; medial gastrocnemial fossa broad and very slightly concave.
|
||
</paragraph>
|
||
<caption id="DF336663FF5DD584FF0073CAFAC9F7E9" ID-DOI="http://doi.org/10.5281/zenodo.10994000" ID-Zenodo-Dep="10994000" httpUri="https://zenodo.org/record/10994000/files/figure.png" pageId="177" pageNumber="178" startId="177.[136,235,1932,1956]" targetBox="[207,1381,182,1900]" targetPageId="177" targetType="figure">
|
||
<paragraph id="8BF336EBFF5DD584FF0073CAFAC9F7E9" blockId="177.[136,1452,1932,2053]" pageId="177" pageNumber="178">
|
||
<emphasis id="B938EAF9FF5DD584FF0073CAFECEF849" bold="true" box="[136,286,1932,1957]" pageId="177" pageNumber="178">FIGURE 102.</emphasis>
|
||
surface scan images of partial hindlimb elements of
|
||
<taxonomicName id="4C4C4D68FF5DD584FCCA73CBFC11F849" box="[834,961,1933,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="177" pageNumber="178" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5DD584FCCA73CBFC11F849" box="[834,961,1933,1957]" italics="true" pageId="177" pageNumber="178">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
: (a) proximal right femoral fragment of UCMP 70665 in dorsal view; (b–c) partial proximal right femur of UCMP 45344 in (b) dorsal, and (c) ventral views; (d–e) left distal femoral epiphysis of UCMP 45344 in (d) distal, and (e) lateral views; (f–i) partial proximal left tibia of UCMP 70584 in (f) lateral, (g) medial, (h) cranial, and (i) proximal views; and (j–k) partial right tibial shaft in (j) lateral, and (k) medial views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF5ED587FF3074EDFD76FB49" blockId="178.[136,779,171,2085]" pageId="178" pageNumber="179">
|
||
The femur of
|
||
<taxonomicName id="4C4C4D68FF5ED587FED974EAFE08FF29" box="[337,472,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5ED587FED974EAFE08FF29" box="[337,472,171,197]" italics="true" pageId="178" pageNumber="179">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF5ED587FD3174EAFCD9FF29" authorityName=", Owen" authorityYear="1874" box="[697,777,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF5ED587FD3174EAFCD9FF29" box="[697,777,171,197]" italics="true" pageId="178" pageNumber="179">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with a deeper, narrower trochlea and a narrower, more raised medial trochlear crest; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FD7E74AAFF2AFEC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5ED587FD7E74AAFF2AFEC9" italics="true" pageId="178" pageNumber="179">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5ED587FF76754DFE85FEC9" bold="true" box="[254,341,267,293]" pageId="178" pageNumber="179">
|
||
<taxonomicNameLabel id="A20B5782FF5ED587FF76754DFE85FEC9" box="[254,341,267,293]" pageId="178" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a narrower, deeper trochlea, and a narrower medial trochlear crest and medial condyle; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FEAE750AFE51FE89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[294,385,332,357]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5ED587FEAE750AFE51FE89" box="[294,385,332,357]" italics="true" pageId="178" pageNumber="179">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5ED587FE0D750DFE0DFE89" bold="true" box="[389,477,331,357]" pageId="178" pageNumber="179">
|
||
<taxonomicNameLabel id="A20B5782FF5ED587FE0D750DFE0DFE89" box="[389,477,331,357]" pageId="178" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a less medially projected lesser trochanter, and a narrower medial trochlear crest and medial condyle; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FDE175CAFD39FE49" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[617,745,395,421]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF5ED587FDE175CAFD39FE49" box="[617,745,395,421]" italics="true" pageId="178" pageNumber="179">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having a less elongate, more proximally situated quadratus tubercle; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FEBD758DFE17FE09" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[309,455,459,485]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF5ED587FEBD758DFE17FE09" box="[309,455,459,485]" italics="true" pageId="178" pageNumber="179">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having a more elongate, medially deflected and transversely compressed greater trochanter, deeper trochlea, trochlear crests subequal in height, and a less elongate and more proximally situated lateral gastrocnemial fossa; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FE7C760DFD9DFD89" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[500,589,587,613]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5ED587FE7C760DFD9DFD89" box="[500,589,587,613]" italics="true" pageId="178" pageNumber="179">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in having a more medially deflected greater trochanter, head positioned closer to the base of the greater trochanter, more distally extensive lesser trochanteric ridge, relatively broader distal condyles, trochlear crests subequal in height, more laterally projected lateral epicondyle, and a more proximally situated lateral gastrocnemial fossa; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FD8C774AFD73FCC9" authorityName="Desmarest" authorityYear="1817" box="[516,675,779,805]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5ED587FD8C774AFD73FCC9" box="[516,675,779,805]" italics="true" pageId="178" pageNumber="179">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being generally larger, with a more elongate, more transversely compressed greater trochanter, more distally extensive lesser trochanteric ridge, relatively broader distal condyles, deeper trochlea, more laterally projected lateral epicondyle, and a more proximally situated lateral gastrocnemial fossa; and from
|
||
<taxonomicName id="4C4C4D68FF5ED587FF78778AFEB0FC09" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[240,352,971,997]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5ED587FF78778AFEB0FC09" box="[240,352,971,997]" italics="true" pageId="178" pageNumber="179">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a relatively larger, more dorsally projected and more medially tilted head, more rounded greater trochanter, more distally extensive lesser trochanteric crest, more distally situated quadratus tubercle, broader, less raised trochlear crests, slightly deeper trochlea, larger fibular facet on the lateral condyle, and a more laterally projected lateral epicondyle.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5ED587FF3070EDFEE3F889" blockId="178.[136,779,171,2085]" pageId="178" pageNumber="179">
|
||
<emphasis id="B938EAF9FF5ED587FF3070EDFE48FB29" box="[184,408,1195,1221]" italics="true" pageId="178" pageNumber="179">
|
||
Tibia (
|
||
<figureCitation id="13772A6EFF5ED587FE8070EDFE41FB29" box="[264,401,1195,1221]" captionStart="FIGURE 102" captionStartId="177.[136,235,1932,1956]" captionTargetBox="[207,1381,182,1900]" captionTargetId="figure-17@177.[197,1391,172,1910]" captionTargetPageId="177" captionText="FIGURE 102. surface scan images of partial hindlimb elements of P. otibandus: (a) proximal right femoral fragment of UCMP 70665 in dorsal view; (b–c) partial proximal right femur of UCMP 45344 in (b) dorsal, and (c) ventral views; (d–e) left distal femoral epiphysis of UCMP 45344 in (d) distal, and (e) lateral views; (f–i) partial proximal left tibia of UCMP 70584 in (f) lateral, (g) medial, (h) cranial, and (i) proximal views; and (j–k) partial right tibial shaft in (j) lateral, and (k) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994000" httpUri="https://zenodo.org/record/10994000/files/figure.png" pageId="178" pageNumber="179">Fig. 102f–k</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: quite small, short and robust. Proximal epiphysis partially preserved; cranial component narrow and moderately cranially tilted; lateral condyle caudolaterally rounded and projected; intercondylar eminence tall, thick and elongate; medial condyle not known. Proximal fibular facet large, deep, broad and caudolaterally projected, with a slight distal deflection. Cnemial crest thick and shallow (
|
||
<figureCitation id="13772A6EFF5ED587FD8571CDFDACFA4A" box="[525,636,1419,1446]" captionStart="FIGURE 102" captionStartId="177.[136,235,1932,1956]" captionTargetBox="[207,1381,182,1900]" captionTargetId="figure-17@177.[197,1391,172,1910]" captionTargetPageId="177" captionText="FIGURE 102. surface scan images of partial hindlimb elements of P. otibandus: (a) proximal right femoral fragment of UCMP 70665 in dorsal view; (b–c) partial proximal right femur of UCMP 45344 in (b) dorsal, and (c) ventral views; (d–e) left distal femoral epiphysis of UCMP 45344 in (d) distal, and (e) lateral views; (f–i) partial proximal left tibia of UCMP 70584 in (f) lateral, (g) medial, (h) cranial, and (i) proximal views; and (j–k) partial right tibial shaft in (j) lateral, and (k) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994000" httpUri="https://zenodo.org/record/10994000/files/figure.png" pageId="178" pageNumber="179">Fig. 102f</figureCitation>
|
||
); peak quite distinct and rugose, crest broadens distally, gently lowers, becomes rounder and merges gently into the shaft. Proximolateral crest raised and rounded proximally, rapidly narrowing and steadily becoming very raised before merging into the distal fibular facet at the midpoint of the shaft. Distal fibular facet extends from the midpoint of the shaft to the distal epiphysis; deep, distinct, slightly convex and slightly craniolaterally projected, particularly proximally. Distal shaft broad, particularly distally. Distal end very broad, with a gently convex trochlear articular surface, roughly rectangular and slightly caudally tilted; medial tuberosity large and elongate, and medial malleolus elongate and ventrally projected with caudal part curving caudomedially.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5ED587FF30732DFB3AFCE9" blockId="178.[136,779,171,2085]" lastBlockId="178.[810,1452,171,965]" pageId="178" pageNumber="179">
|
||
The tibia of
|
||
<taxonomicName id="4C4C4D68FF5ED587FED1732AFE3AF869" box="[345,490,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5ED587FED1732AFE3AF869" box="[345,490,1899,1925]" italics="true" pageId="178" pageNumber="179">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF5ED587FD7D732AFF6EF849" authorityName=", Owen" authorityYear="1874" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF5ED587FD7D732AFF6EF849" italics="true" pageId="178" pageNumber="179">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being shorter, with a relatively slightly broader distal epiphysis; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FE1E73EAFDFDF829" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[406,557,1963,1989]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5ED587FE1E73EAFDFDF829" box="[406,557,1963,1989]" italics="true" pageId="178" pageNumber="179">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5ED587FDB273EDFD4AF829" bold="true" box="[570,666,1963,1989]" pageId="178" pageNumber="179">
|
||
<taxonomicNameLabel id="A20B5782FF5ED587FDB273EDFD4AF829" box="[570,666,1963,1989]" pageId="178" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with larger, broader proximal fibular facet and a shallower, relatively less distally extensive cnemial crest; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FF4C7C4AFEF2F7C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[196,290,2060,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5ED587FF4C7C4AFEF2F7C9" box="[196,290,2060,2085]" italics="true" pageId="178" pageNumber="179">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5ED587FEAF7C4DFE50F7C9" bold="true" box="[295,384,2059,2085]" pageId="178" pageNumber="179">
|
||
<taxonomicNameLabel id="A20B5782FF5ED587FEAF7C4DFE50F7C9" box="[295,384,2059,2085]" pageId="178" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF5ED587FE3C7C4AFDF4F7C9" authorityName="Bartholomai" authorityYear="1978" box="[436,548,2060,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF5ED587FE3C7C4AFDF4F7C9" box="[436,548,2060,2085]" italics="true" pageId="178" pageNumber="179">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being smaller and more robust, with a larger, broader proximal fibular facet, gently cranially tilted cranial section of the proximal epiphysis, and a slightly more curved cnemial crest in cranial view; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FC75754AFBAAFEC9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1021,1146,267,293]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF5ED587FC75754AFBAAFEC9" box="[1021,1146,267,293]" italics="true" pageId="178" pageNumber="179">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in being more gracile, with a thinner, more raised proximolateral crest, and a less distally extensive cnemial crest with a more defined peak; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FCE5752DFBD4FE69" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[877,1028,363,389]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF5ED587FCE5752DFBD4FE69" box="[877,1028,363,389]" italics="true" pageId="178" pageNumber="179">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having broader proximal fibular facet, thicker proximal part of the proximolateral crest, and a deeper, more distinct distal fibular facet; from
|
||
<taxonomicName id="4C4C4D68FF5ED587FA1875EDFCB4FE09" baseAuthorityName="Desmarest" baseAuthorityYear="1822" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5ED587FA1875EDFCB4FE09" italics="true" pageId="178" pageNumber="179">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF5ED587FC2C758AFB9AFE09" authorityName="Desmarest" authorityYear="1817" box="[932,1098,459,485]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5ED587FC2C758AFB9AFE09" box="[932,1098,459,485]" italics="true" pageId="178" pageNumber="179">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being more robust, with a cranially tilted cranial section of the proximal epiphysis, broader proximal fibular facet, relatively longer cnemial crest with less distinct distal peak, thicker proximolateral crest, deeper, more distinct distal fibular facet and more craniocaudally compressed distal epiphysis; and from
|
||
<taxonomicName id="4C4C4D68FF5ED587FA18762AFCABFD49" baseAuthorityName="Desmarest" baseAuthorityYear="1804" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="178" pageNumber="179" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5ED587FA18762AFCABFD49" italics="true" pageId="178" pageNumber="179">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a deeper proximal fibular facet, thicker proximolateral crest, relatively more elongate cnemial crest with a less angular, less distinct distal peak, and a broader talar trochlea.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5ED587FCD2774DFB56FC29" blockId="178.[810,1452,171,965]" pageId="178" pageNumber="179">
|
||
<emphasis id="B938EAF9FF5ED587FCD2774DFC75FCC9" box="[858,933,779,805]" italics="true" pageId="178" pageNumber="179">Fibula</emphasis>
|
||
: only known specimen is a fragmentary distal shaft; articulates with the lateral surface of the distal half of the tibia. Proximal end interpreted as being large and broad based on the size of the proximal fibular facet on the tibia. Distal shaft quite deep and gently crescentic in cross-section, thickens distally.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5ED587FCA277ADFC84FBE9" blockId="178.[810,1452,1003,2085]" box="[810,852,1003,1029]" pageId="178" pageNumber="179">
|
||
<emphasis id="B938EAF9FF5ED587FCA277ADFC84FBE9" bold="true" box="[810,852,1003,1029]" italics="true" pageId="178" pageNumber="179">Pes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5ED587FCA2704DFCB8FA49" blockId="178.[810,1452,1003,2085]" pageId="178" pageNumber="179">
|
||
<emphasis id="B938EAF9FF5ED587FCA2704DFBE9FBC9" box="[810,1081,1035,1061]" italics="true" pageId="178" pageNumber="179">
|
||
Calcaneus (
|
||
<figureCitation id="13772A6EFF5ED587FC27704DFBE2FBC9" box="[943,1074,1035,1061]" captionStart="FIGURE 103" captionStartId="179.[136,235,1932,1956]" captionTargetBox="[198,1389,177,1895]" captionTargetId="figure-18@179.[188,1399,167,1905]" captionTargetPageId="179" captionText="FIGURE 103. left tarsal elements of P. otibandus: (a–d, f) surface scan images of partial calcaneus of UCMP 70584 in (a) dorsal, (b) plantar, (c) medial, (d) lateral, and (f) caudodorsal views; (e) calcaneus of UCMP 45247 in cranial view; (g–i) surface scan images of talus of UCMP 70584 in (g) dorsal, (h) cranial, and (i) medial views; and (j–l) surface scan images of cuboid of UCMP 70078 in (j) caudal, (k) cranial, and (l) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994002" httpUri="https://zenodo.org/record/10994002/files/figure.png" pageId="178" pageNumber="179">Fig. 103a–f</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: short, low and robust. Calcaneal tuberosity broad, low and domed in cross-section, almost squared immediately caudal to the calcaneal head. Caudal epiphysis low and rounded in caudal view with a thin, shallow, transverse channel on the caudal surface. Plantar surface thick, rugose, quite broad and gently narrowing cranially, and curving slightly laterally to a rounded point lateral to the cranial plantar tubercle; a thicker and more plantarly projected caudal section is formed from the caudal epiphysis. Cranial plantar tubercule small, rounded, plantarly projected and immediately caudoplantarly adjacent to the plantar margin of the plantomedial cuboid facet.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5ED581FCD271EDFEF9FE69" blockId="178.[810,1452,1003,2085]" lastBlockId="180.[136,779,171,2085]" lastPageId="180" lastPageNumber="181" pageId="178" pageNumber="179">
|
||
Calcaneal head large and broad; midline in the sagittal plane is slightly medially offset relative to that of the calcaneal tuberosity (
|
||
<figureCitation id="13772A6EFF5ED587FBFF71ADFB3CF9E9" box="[1143,1260,1515,1541]" captionStart="FIGURE 103" captionStartId="179.[136,235,1932,1956]" captionTargetBox="[198,1389,177,1895]" captionTargetId="figure-18@179.[188,1399,167,1905]" captionTargetPageId="179" captionText="FIGURE 103. left tarsal elements of P. otibandus: (a–d, f) surface scan images of partial calcaneus of UCMP 70584 in (a) dorsal, (b) plantar, (c) medial, (d) lateral, and (f) caudodorsal views; (e) calcaneus of UCMP 45247 in cranial view; (g–i) surface scan images of talus of UCMP 70584 in (g) dorsal, (h) cranial, and (i) medial views; and (j–l) surface scan images of cuboid of UCMP 70078 in (j) caudal, (k) cranial, and (l) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994002" httpUri="https://zenodo.org/record/10994002/files/figure.png" pageId="178" pageNumber="179">Fig. 103a</figureCitation>
|
||
). Sustentaculum tali broad, thick and rounded with no distinct peak in medial view; projected medially slightly beyond the medial margin of the medial talar facet, extends caudally well past the caudal margins of the talar facets; flexor groove very broad and deep, accentuated by the medial projection of the plantar surface. Fibular facet small, with semi-distinct cranial and caudal components, separated by a broad groove; caudal component quite small, roughly oval, unprojected caudally, slightly projected laterally, and medially semicontinuous with lateral talar facet; cranial component lower than the caudal component, broad, laterally projected and slightly laterally tilted, with the dorsal surface gently concave. Lateral talar facet broad, quite tall, smoothly convex with distinct medial tapering; a rounded, very shallow fossa is immediately cranial to the lateral talar facet, with a small, low, rounded tubercle on the medial margin of the fossa; medial talar facet caudally displaced relative to the lateral facet; small, oval, dorsally slightly projected, orientated caudomedially to craniolaterally and strongly cranially tilted, with a rounded lip along the caudomedial margin; smoothly continues caudally into a very low, rounded tubercle or ridge merging caudally into the dorsolateral margin of the tuberosity.
|
||
</paragraph>
|
||
<caption id="DF336663FF5FD586FF0073CAFD71F7E9" ID-DOI="http://doi.org/10.5281/zenodo.10994002" ID-Zenodo-Dep="10994002" httpUri="https://zenodo.org/record/10994002/files/figure.png" pageId="179" pageNumber="180" startId="179.[136,235,1932,1956]" targetBox="[198,1389,177,1895]" targetPageId="179" targetType="figure">
|
||
<paragraph id="8BF336EBFF5FD586FF0073CAFD71F7E9" blockId="179.[136,1452,1932,2053]" pageId="179" pageNumber="180">
|
||
<emphasis id="B938EAF9FF5FD586FF0073CAFECDF849" bold="true" box="[136,285,1932,1957]" pageId="179" pageNumber="180">FIGURE 103.</emphasis>
|
||
left tarsal elements of
|
||
<taxonomicName id="4C4C4D68FF5FD586FD8D73CBFD52F849" box="[517,642,1933,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="179" pageNumber="180" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5FD586FD8D73CBFD52F849" box="[517,642,1933,1957]" italics="true" pageId="179" pageNumber="180">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
: (a–d, f) surface scan images of partial calcaneus of UCMP 70584 in (a) dorsal, (b) plantar, (c) medial, (d) lateral, and (f) caudodorsal views; (e) calcaneus of UCMP 45247 in cranial view; (g–i) surface scan images of talus of UCMP 70584 in (g) dorsal, (h) cranial, and (i) medial views; and (j–l) surface scan images of cuboid of UCMP 70078 in (j) caudal, (k) cranial, and (l) medial views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF58D581FF3075CDFE51FCA9" blockId="180.[136,779,171,2085]" pageId="180" pageNumber="181">Talar head facet very small, rounded, abuts the dorsomedial margin of the dorsomedial facet on the medial surface of the calcaneal head. Dorsomedial cuboid facet squarish with a rounded dorsomedial margin, gently convex and slightly dorsally projected; separated from the dorsolateral facet by a deep, bevelled step; small, shallow fossa sits immediately plantarly. Dorsolateral cuboid facet tall, subequal in width to the dorsomedial facet, cranially projected and roughly semicircular, with curved dorsal and lateral margins; extends plantarly and curving medially to be continuous with the plantomedial facet. Plantomedial cuboid facet small, broad, dorsoplantarly short and oblong; extends medially past the midpoint of the dorsomedial facet.</paragraph>
|
||
<paragraph id="8BF336EBFF58D581FF30770DFC71FEC9" blockId="180.[136,779,171,2085]" lastBlockId="180.[810,1452,171,2085]" pageId="180" pageNumber="181">
|
||
The calcaneus of
|
||
<taxonomicName id="4C4C4D68FF58D581FE28770AFDE6FC89" box="[416,566,843,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF58D581FE28770AFDE6FC89" box="[416,566,843,869]" italics="true" pageId="180" pageNumber="181">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from all compared species in having the calcaneal head slightly medially offset relative to the calcaneal tuberosity. It further differs from
|
||
<taxonomicName id="4C4C4D68FF58D581FEF877EAFE14FC29" authorityName=", Owen" authorityYear="1874" box="[368,452,939,965]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF58D581FEF877EAFE14FC29" box="[368,452,939,965]" italics="true" pageId="180" pageNumber="181">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller and lower, with a relatively shorter, slightly broader tuberosity that is more rounded dorsally in cross-section, less straight, more cranially tapered plantar surface, less rounded, less distinct and less caudolaterally projected caudal component of the fibular facet, less concave plantolateral margin of the dorsolateral-plantomedial facet, and a more laterally tilted dorsomedial cuboid facet; from
|
||
<taxonomicName id="4C4C4D68FF58D581FD7D70CAFF2DFB29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF58D581FD7D70CAFF2DFB29" italics="true" pageId="180" pageNumber="181">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF58D581FE8070EDFEB6FB29" bold="true" box="[264,358,1195,1221]" pageId="180" pageNumber="181">
|
||
<taxonomicNameLabel id="A20B5782FF58D581FE8070EDFEB6FB29" box="[264,358,1195,1221]" pageId="180" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and lower, with a slightly narrower, more cranially tapered plantar surface, more rounded sustentaculum tali, less bulbous lateral talar facet, relatively smaller medial talar facet that is more cranially tilted and more caudally displaced relative to the lateral facet, more laterally tilted dorsomedial cuboid facet, and a less medially tilted dorsolateral cuboid facet; from
|
||
<taxonomicName id="4C4C4D68FF58D581FF4C71CAFEF1FA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[196,289,1420,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF58D581FF4C71CAFEF1FA49" box="[196,289,1420,1445]" italics="true" pageId="180" pageNumber="181">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF58D581FEAF71CDFEAFFA49" bold="true" box="[295,383,1419,1445]" pageId="180" pageNumber="181">
|
||
<taxonomicNameLabel id="A20B5782FF58D581FEAF71CDFEAFFA49" box="[295,383,1419,1445]" pageId="180" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, lower, shorter and much more robust, with a lower, more rounded tuberosity in cross-section, relatively narrower dorsomedial cuboid facet, and in having the lateral surface of the calcaneal head beneath the fibular facet dorsoplantarly much shorter and less concave; from
|
||
<taxonomicName id="4C4C4D68FF58D581FE71726AFDAAF9A9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[505,634,1579,1605]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF58D581FE71726AFDAAF9A9" box="[505,634,1579,1605]" italics="true" pageId="180" pageNumber="181">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having a less medially displaced head relative to the tuberosity, more planar plantar surface with less flaring of the medial margin, and a smaller, shallower fossa cranial to the lateral talar facet; from
|
||
<taxonomicName id="4C4C4D68FF58D581FECA72EAFE1DF929" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[322,461,1707,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF58D581FECA72EAFE1DF929" box="[322,461,1707,1733]" italics="true" pageId="180" pageNumber="181">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF58D581FE5B72EDFDFCF929" bold="true" box="[467,556,1707,1733]" pageId="180" pageNumber="181">
|
||
<taxonomicNameLabel id="A20B5782FF58D581FE5B72EDFDFCF929" box="[467,556,1707,1733]" pageId="180" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a broader calcaneal tuberosity; from
|
||
<taxonomicName id="4C4C4D68FF58D581FE37728DFD84F909" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[447,596,1739,1765]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF58D581FE37728DFD84F909" box="[447,596,1739,1765]" italics="true" pageId="180" pageNumber="181">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger and taller, with a more planar, more cranially extensive and less laterally tilted plantar surface, more dorsally projected medial talar facet, and a less caudally projected, less distinct caudal component of the fibular facet; from
|
||
<taxonomicName id="4C4C4D68FF58D581FF00732DFF37F869" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[136,231,1899,1925]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF58D581FF00732DFF37F869" box="[136,231,1899,1925]" italics="true" pageId="180" pageNumber="181">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF58D581FEAC732AFE17F869" authorityName="Desmarest" authorityYear="1817" box="[292,455,1899,1925]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF58D581FEAC732AFE17F869" box="[292,455,1899,1925]" italics="true" pageId="180" pageNumber="181">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being much broader and more robust, with a broadly rounded, domed calcaneal tuberosity in cross-section, more pointed, more medially projected sustentaculum tali, less cranially extensive cranial component of the fibular facet, more bevelled step between the dorsal cuboid facets, and a more curved, less cranially extensive plantar surface that broadens caudally; and from
|
||
<taxonomicName id="4C4C4D68FF58D581FC1C748AFBD7FF09" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[916,1031,203,229]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF58D581FC1C748AFBD7FF09" box="[916,1031,203,229]" italics="true" pageId="180" pageNumber="181">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and relatively broader, with a relatively larger fibular facet and a broader medial talar facet.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF58D581FCD2756DFA9FFC29" blockId="180.[810,1452,171,2085]" pageId="180" pageNumber="181">
|
||
<emphasis id="B938EAF9FF58D581FCD2756DFB98FEA9" box="[858,1096,299,325]" italics="true" pageId="180" pageNumber="181">
|
||
Talus (
|
||
<figureCitation id="13772A6EFF58D581FC39756DFB91FEA9" box="[945,1089,299,325]" captionStart="FIGURE 103" captionStartId="179.[136,235,1932,1956]" captionTargetBox="[198,1389,177,1895]" captionTargetId="figure-18@179.[188,1399,167,1905]" captionTargetPageId="179" captionText="FIGURE 103. left tarsal elements of P. otibandus: (a–d, f) surface scan images of partial calcaneus of UCMP 70584 in (a) dorsal, (b) plantar, (c) medial, (d) lateral, and (f) caudodorsal views; (e) calcaneus of UCMP 45247 in cranial view; (g–i) surface scan images of talus of UCMP 70584 in (g) dorsal, (h) cranial, and (i) medial views; and (j–l) surface scan images of cuboid of UCMP 70078 in (j) caudal, (k) cranial, and (l) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994002" httpUri="https://zenodo.org/record/10994002/files/figure.png" pageId="180" pageNumber="181">
|
||
Fig.
|
||
<quantity id="4CB49B0EFF58D581FC79756DFBFAFEA9" box="[1009,1066,299,325]" metricMagnitude="-1" metricUnit="kg" metricValue="1.03" pageId="180" pageNumber="181" unit="g" value="103.0">103g</quantity>
|
||
–i
|
||
</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: slightly width greater than craniocaudal length. Trochlear crests equal in height, with the medial peak thinner and less rounded. Trochlea is fairly deep and medially skewed (
|
||
<figureCitation id="13772A6EFF58D581FB5A75CDFA96FE4A" box="[1234,1350,395,422]" captionStart="FIGURE 103" captionStartId="179.[136,235,1932,1956]" captionTargetBox="[198,1389,177,1895]" captionTargetId="figure-18@179.[188,1399,167,1905]" captionTargetPageId="179" captionText="FIGURE 103. left tarsal elements of P. otibandus: (a–d, f) surface scan images of partial calcaneus of UCMP 70584 in (a) dorsal, (b) plantar, (c) medial, (d) lateral, and (f) caudodorsal views; (e) calcaneus of UCMP 45247 in cranial view; (g–i) surface scan images of talus of UCMP 70584 in (g) dorsal, (h) cranial, and (i) medial views; and (j–l) surface scan images of cuboid of UCMP 70078 in (j) caudal, (k) cranial, and (l) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994002" httpUri="https://zenodo.org/record/10994002/files/figure.png" pageId="180" pageNumber="181">Fig. 103h</figureCitation>
|
||
). Medial malleolus large, rounded, smoothly convex, dorsal facing, extends medially and slightly cranially, bounded laterally by a deep, rounded malleolar fossa; rounded, plantomedially projected tubercle situated plantar to the medial malleolus on the medial surface of the talar neck. Talar head broad and distinctly plantarly deflected, creating a concave divot between the head and cranial margin of medial trochlear crest. Facet for the navicular broad dorsally and extending caudoplantarly to beneath anterior margin of medial malleolus. Facet for the cuboid very small, slightly convex and lateral facing, with a very slight cranioplantar tilt. Posterior plantar process rugose, very broad and thickened, deep, plantarly projected and rounded. On the plantar surface, the medial calcaneal facet is rounded and concave, abuts a small, deep, rounded mesial fossa, and the lateral calcaneal facet is very large, broad, deeply concave, becomes deeper laterally.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF58D581FCD2778DFC0AF889" blockId="180.[810,1452,171,2085]" pageId="180" pageNumber="181">
|
||
The talus of
|
||
<taxonomicName id="4C4C4D68FF58D581FC60778AFBA2FC09" box="[1000,1138,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF58D581FC60778AFBA2FC09" box="[1000,1138,971,997]" italics="true" pageId="180" pageNumber="181">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF58D581FAD1778AFA7BFC09" authorityName=", Owen" authorityYear="1874" box="[1369,1451,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF58D581FAD1778AFA7BFC09" box="[1369,1451,971,997]" italics="true" pageId="180" pageNumber="181">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with a broader navicular facet that is more obliquely angled in cranial view, larger, more obliquely aligned medial malleolus, and a small tubercle plantar to the medial malleolus on the medial surface; from
|
||
<taxonomicName id="4C4C4D68FF58D581FCEE702AFC25FB69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[870,1013,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF58D581FCEE702AFC25FB69" box="[870,1013,1131,1157]" italics="true" pageId="180" pageNumber="181">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF58D581FC73702DFB83FB69" bold="true" box="[1019,1107,1131,1157]" pageId="180" pageNumber="181">
|
||
<taxonomicNameLabel id="A20B5782FF58D581FC73702DFB83FB69" box="[1019,1107,1131,1157]" pageId="180" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a deeper malleolar fossa, smaller cuboid facet, less medially projected talar head, and a deeper plantar groove between the navicular facet and the posterior plantar tubercle; from
|
||
<taxonomicName id="4C4C4D68FF58D581FCA270AAFC5BFAE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[810,907,1260,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF58D581FCA270AAFC5BFAE9" box="[810,907,1260,1285]" italics="true" pageId="180" pageNumber="181">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF58D581FC1C70ADFC20FAE9" bold="true" box="[916,1008,1259,1285]" pageId="180" pageNumber="181">
|
||
<taxonomicNameLabel id="A20B5782FF58D581FC1C70ADFC20FAE9" box="[916,1008,1259,1285]" pageId="180" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with a shorter, more concave, and more cranially situated malleolar fossa, a dorsal facing medial malleolus, and a deeper plantar groove between the navicular facet and the posterior plantar tubercle; from
|
||
<taxonomicName id="4C4C4D68FF58D581FBB5712AFB67FA69" authorityName="Bartholomai" authorityYear="1978" box="[1085,1207,1388,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF58D581FBB5712AFB67FA69" box="[1085,1207,1388,1413]" italics="true" pageId="180" pageNumber="181">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having a deeper trochlea and a deeper concavity between the posterior plantar tubercle and the talar head; from
|
||
<taxonomicName id="4C4C4D68FF58D581FB7371EDFA5DFA29" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1275,1421,1451,1477]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF58D581FB7371EDFA5DFA29" box="[1275,1421,1451,1477]" italics="true" pageId="180" pageNumber="181">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having a more medially projected medial malleolus, more caudoplantarly extensive navicular facet, a facet for the cuboid present on the navicular head, and a deeper lateral calcaneal facet; from
|
||
<taxonomicName id="4C4C4D68FF58D581FB95726DFBAAF9A9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1053,1146,1579,1605]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF58D581FB95726DFBAAF9A9" box="[1053,1146,1579,1605]" italics="true" pageId="180" pageNumber="181">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being relatively broader and more caudolaterally extensive, with a deeper trochlea and a larger, more rounded posterior plantar process; from
|
||
<taxonomicName id="4C4C4D68FF58D581FCE272CAFBDDF949" authorityName="Desmarest" authorityYear="1817" box="[874,1037,1675,1701]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF58D581FCE272CAFBDDF949" box="[874,1037,1675,1701]" italics="true" pageId="180" pageNumber="181">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being generally larger, relatively broader and more caudolaterally extensive, with a deeper trochlea and a deeper, broader and more rounded posterior plantar process; and from
|
||
<taxonomicName id="4C4C4D68FF58D581FBDD72AAFB1BF8E9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1109,1227,1771,1797]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="180" pageNumber="181" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF58D581FBDD72AAFB1BF8E9" box="[1109,1227,1771,1797]" italics="true" pageId="180" pageNumber="181">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and relatively broader, with a more caudoplantarly extensive navicular facet and a broader, more rounded posterior plantar process.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF58D580FCD2732DFEA1FE29" blockId="180.[810,1452,171,2085]" lastBlockId="181.[136,778,171,2085]" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181">
|
||
<emphasis id="B938EAF9FF58D581FCD2732DFB6CF869" box="[858,1212,1899,1925]" italics="true" pageId="180" pageNumber="181">
|
||
Cuboid (
|
||
<figureCitation id="13772A6EFF58D581FC4A732DFB9DF869" box="[962,1101,1899,1925]" captionStart="FIGURE 103" captionStartId="179.[136,235,1932,1956]" captionTargetBox="[198,1389,177,1895]" captionTargetId="figure-18@179.[188,1399,167,1905]" captionTargetPageId="179" captionText="FIGURE 103. left tarsal elements of P. otibandus: (a–d, f) surface scan images of partial calcaneus of UCMP 70584 in (a) dorsal, (b) plantar, (c) medial, (d) lateral, and (f) caudodorsal views; (e) calcaneus of UCMP 45247 in cranial view; (g–i) surface scan images of talus of UCMP 70584 in (g) dorsal, (h) cranial, and (i) medial views; and (j–l) surface scan images of cuboid of UCMP 70078 in (j) caudal, (k) cranial, and (l) medial views." figureDoi="http://doi.org/10.5281/zenodo.10994002" httpUri="https://zenodo.org/record/10994002/files/figure.png" pageId="180" pageNumber="181">Figs 103j–l</figureCitation>
|
||
,
|
||
<figureCitation id="13772A6EFF58D581FBD7732DFB64F869" box="[1119,1204,1899,1925]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="180" pageNumber="181">104a–d</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: square and robust. Dorsomedial calcaneal facet squarish, caudally projected, smoothly concave and subequal to the dorsolateral calcaneal facet, separated by a bevelled step; dorsolateral calcaneal facet tall, gently concave and extending plantarly and slightly medially to be continuous with the plantar calcaneal facet; plantar calcaneal facet broad, dorsoplantarly short, oblong and gently concave, with the medial margin situated lateral to the transverse midpoint of the dorsomedial facet, situated on the caudal base of the lateral plantar tubercle. Dorsal margin of the medial surface marked by a low, rounded ridge projecting medially over a tall, broad fossa for articulation with the caudodorsal part of the ectocuneiform and the craniodorsal part of the navicular.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF59D580FF30758DFE5AFB89" blockId="181.[136,778,171,2085]" pageId="181" pageNumber="182">Facet for the talar head narrow and indistinct. Facet for the navicular small and narrow on the caudal margin of dorsal part of the medial surface, adjacent to the medial margin of the dorsomedial calcaneal facet. Facet for the ectocuneiform very tall and thin, extends from near the dorsal margin of the medial surface past the midpoint of the cranial margin, with a distinct, rounded second facet on the plantomedial surface of the medial plantar tubercle. Lateral plantar tubercle large, broad and quite plantarly projected, oval to round in plantar view; occasionally deflected caudally (see UCMP 70078). Medial plantar tubercle moderately small, rounded, less plantarly projected than the lateral tubercle and slightly medially deflected; separated from the lateral tubercle by a variably deep, narrow flexor groove. Facet for metatarsal IV broad and very gently concave, with the dorsomedial margin medially projected; not continuous plantarly with the small, rounded, plantarly tilted plantar facet for metatarsal IV. Facet for metatarsal V fairly small, rounded, gently concave, slightly plantolaterally tilted and laterally projected.</paragraph>
|
||
<paragraph id="8BF336EBFF59D580FF30702DFB67FDA9" blockId="181.[136,778,171,2085]" lastBlockId="181.[810,1452,171,2085]" pageId="181" pageNumber="182">
|
||
The cuboid of
|
||
<taxonomicName id="4C4C4D68FF59D580FEDF702AFE0FFB69" box="[343,479,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF59D580FEDF702AFE0FFB69" box="[343,479,1131,1157]" italics="true" pageId="181" pageNumber="182">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF59D580FD32702AFCD9FB69" authorityName=", Owen" authorityYear="1874" box="[698,777,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF59D580FD32702AFCD9FB69" box="[698,777,1131,1157]" italics="true" pageId="181" pageNumber="182">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller and lower, with less plantarly projected plantar tubercles, and the dorsal and plantar metatarsal IV facets distinct from one another; from
|
||
<taxonomicName id="4C4C4D68FF59D580FDFE708AFCDAFB09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[630,778,1227,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF59D580FDFE708AFCDAFB09" box="[630,778,1227,1253]" italics="true" pageId="181" pageNumber="182">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF59D580FF0070ADFF33FAE9" bold="true" box="[136,227,1259,1285]" pageId="181" pageNumber="182">
|
||
<taxonomicNameLabel id="A20B5782FF59D580FF0070ADFF33FAE9" box="[136,227,1259,1285]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with less distinct step between the dorsal calcaneal facets, slightly narrower dorsolateral calcaneal facet, smaller, less distinct talar head facet, deeper medial fossa for the navicular and ectocuneiform, more plantarly projected lateral plantar tubercle, narrower flexor groove, more laterally tapered metatarsal IV facet that is distinct from the plantar metatarsal IV facet, and a smaller, narrower, more concave metatarsal V facet; from
|
||
<taxonomicName id="4C4C4D68FF59D580FF0071AAFF38F9E9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,232,1516,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF59D580FF0071AAFF38F9E9" box="[136,232,1516,1541]" italics="true" pageId="181" pageNumber="182">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF59D580FF7871ADFE9BF9E9" bold="true" box="[240,331,1515,1541]" pageId="181" pageNumber="182">
|
||
<taxonomicNameLabel id="A20B5782FF59D580FF7871ADFE9BF9E9" box="[240,331,1515,1541]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, shorter and relatively broader, with a more plantarly situated navicular facet, less elongate, less plantarly projected plantar tubercles, and a shallower flexor groove; from
|
||
<taxonomicName id="4C4C4D68FF59D580FDAF720AFD49F989" authorityName="Bartholomai" authorityYear="1978" box="[551,665,1612,1637]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF59D580FDAF720AFD49F989" box="[551,665,1612,1637]" italics="true" pageId="181" pageNumber="182">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having less dorsomedially flared dorsomedial section, more level dorsal calcaneal facets in caudal view, more plantarly extensive plantomedial calcaneal facet, much smaller talar facet, and a much larger, more plantomedially projected medial plantar tubercle; from
|
||
<taxonomicName id="4C4C4D68FF59D580FDDE72ADFD3AF8E9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[598,746,1771,1797]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF59D580FDDE72ADFD3AF8E9" box="[598,746,1771,1797]" italics="true" pageId="181" pageNumber="182">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being slightly larger and taller, with a facet for the talar head present, more plantarly projected lateral plantar tubercle, larger, more elongate and more medially situated medial plantar tubercle, and separate dorsal and plantar metatarsal IV facets; from
|
||
<taxonomicName id="4C4C4D68FF59D580FDCD73CDFD74F849" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[581,676,1931,1957]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF59D580FDCD73CDFD74F849" box="[581,676,1931,1957]" italics="true" pageId="181" pageNumber="182">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being relatively broader, particularly across the plantar section, with a relatively broader dorsolateral calcaneal facet, a smaller, less distinct facet for the talar head, less plantarly projected lateral plantar tubercle, more medially situated medial plantar tubercle, broader flexor groove, and a more medially situated plantar metatarsal IV facet; from
|
||
<taxonomicName id="4C4C4D68FF59D580FCA274AAFC00FEE9" authorityName="Desmarest" authorityYear="1817" box="[810,976,235,261]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF59D580FCA274AAFC00FEE9" box="[810,976,235,261]" italics="true" pageId="181" pageNumber="182">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being generally larger and relatively broader, particularly across plantar section, with a smaller, less distinct facet for the talar head, more deeply concave medial fossa for the navicular and ectocuneiform, broader and less plantarly projected lateral plantar tubercle, larger and more medially situated medial plantar tubercle, broader flexor groove, and separate dorsal and plantar metatarsal IV facets; and from
|
||
<taxonomicName id="4C4C4D68FF59D580FB02758AFB2FFE09" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1162,1279,459,485]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF59D580FB02758AFB2FFE09" box="[1162,1279,459,485]" italics="true" pageId="181" pageNumber="182">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and relatively broader, with a less plantarly projected lateral plantar tubercle and a larger, more plantarly projected medial plantar tuberosity.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF59D580FCD2760DFBC1FC29" blockId="181.[810,1452,171,2085]" pageId="181" pageNumber="182">
|
||
<emphasis id="B938EAF9FF59D580FCD2760DFB42FD89" box="[858,1170,587,613]" italics="true" pageId="181" pageNumber="182">
|
||
Navicular (
|
||
<figureCitation id="13772A6EFF59D580FC55760DFB5AFD89" box="[989,1162,587,613]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="181" pageNumber="182">Fig. 104a–b, d</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: tall, roughly crescentic in medial view and transversely compressed dorsally, broadens gently plantarly, particularly around the laterally flared facet for the ectocuneiform; medial surface rugose. Talar facet smoothly concave and dorsally tilted. Facet for the ectocuneiform tall, gently convex and quite narrow, extends from the dorsal margin of the cranial surface just past the midpoint; plantar component broadens and flares laterally. Facet for the entocuneiform quite small and narrow, oval and situated slightly plantar of the midpoint of the cranial surface, level with the plantar margin of the ectocuneiform facet.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF59D580FCD2778DFB17FA69" blockId="181.[810,1452,171,2085]" pageId="181" pageNumber="182">
|
||
The navicular of
|
||
<taxonomicName id="4C4C4D68FF59D580FBA2778AFB69FC09" box="[1066,1209,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF59D580FBA2778AFB69FC09" box="[1066,1209,971,997]" italics="true" pageId="181" pageNumber="182">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF59D580FCA277AAFC10FBE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[810,960,1003,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF59D580FCA277AAFC10FBE9" box="[810,960,1003,1029]" italics="true" pageId="181" pageNumber="182">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF59D580FC4477ADFBFBFBE9" bold="true" box="[972,1067,1003,1029]" pageId="181" pageNumber="182">
|
||
<taxonomicNameLabel id="A20B5782FF59D580FC4477ADFBFBFBE9" box="[972,1067,1003,1029]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, and shallower relative to height, with a more laterally flared dorsal ectocuneiform facet and a less medially tilted facet for the entocuneiform; from
|
||
<taxonomicName id="4C4C4D68FF59D580FB97700AFB51FB89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1055,1153,1100,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF59D580FB97700AFB51FB89" box="[1055,1153,1100,1125]" italics="true" pageId="181" pageNumber="182">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF59D580FB02700DFB37FB89" bold="true" box="[1162,1255,1099,1125]" pageId="181" pageNumber="182">
|
||
<taxonomicNameLabel id="A20B5782FF59D580FB02700DFB37FB89" box="[1162,1255,1099,1125]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and slightly shallower, with a more medially deflected plantar section and a dorsoplantarly shorter facet for the ectocuneiform; from
|
||
<taxonomicName id="4C4C4D68FF59D580FBD170EDFB21FB29" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1113,1265,1195,1221]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF59D580FBD170EDFB21FB29" box="[1113,1265,1195,1221]" italics="true" pageId="181" pageNumber="182">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being taller and relatively shallower; from
|
||
<taxonomicName id="4C4C4D68FF59D580FB10708DFB29FB09" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1176,1273,1227,1253]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF59D580FB10708DFB29FB09" box="[1176,1273,1227,1253]" italics="true" pageId="181" pageNumber="182">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being taller and relatively shallower, with a more expanded plantar section, and a slightly more plantarly situated facet for the entocuneiform; from
|
||
<taxonomicName id="4C4C4D68FF59D580FBA8716AFB14FAA9" authorityName="Desmarest" authorityYear="1817" box="[1056,1220,1323,1349]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF59D580FBA8716AFB14FAA9" box="[1056,1220,1323,1349]" italics="true" pageId="181" pageNumber="182">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger and much taller; and from
|
||
<taxonomicName id="4C4C4D68FF59D580FBA6710AFB74FA89" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1070,1188,1355,1381]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF59D580FBA6710AFB74FA89" box="[1070,1188,1355,1381]" italics="true" pageId="181" pageNumber="182">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a more medially flared plantar section.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF59D580FCD271CDFA80F889" blockId="181.[810,1452,171,2085]" pageId="181" pageNumber="182">
|
||
<emphasis id="B938EAF9FF59D580FCD271CDFB34FA49" box="[858,1252,1419,1445]" italics="true" pageId="181" pageNumber="182">
|
||
Ectocuneiform (
|
||
<figureCitation id="13772A6EFF59D580FB9471CDFB0BFA49" box="[1052,1243,1419,1445]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="181" pageNumber="182">Fig. 104a–b, d</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: tall and quite transversely compressed; dorsal margin forms a thick, rounded-triangular mesial point, accommodating the articulation of the dorsolateral surface with metatarsal IV. Facet for the navicular tall and quite narrow transversely, broadens plantarly, occupies the entire caudal surface. Facet for metatarsal III concave, quite tall and narrow, occupies the dorsal section of the cranial surface. Facet for the mesocuneiform very small and roughly oval, on the cranial section around the midpoint of the medial surface, plantar to the metatarsal III facet. Medial surface has the dorsal section smooth and gently convex, plantar section pitted and rugose. A small, thin eminence is present on the caudal part of the midpoint of the medial surface, probably for articulation with the entocuneiform.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF59D583FCD2732DFD77F849" blockId="181.[810,1452,171,2085]" lastBlockId="182.[136,778,1675,2085]" lastPageId="182" lastPageNumber="183" pageId="181" pageNumber="182">
|
||
The ectocuneiform of
|
||
<taxonomicName id="4C4C4D68FF59D580FBE8732AFB3DF869" box="[1120,1261,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF59D580FBE8732AFB3DF869" box="[1120,1261,1899,1925]" italics="true" pageId="181" pageNumber="182">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from all compared species in its rounded-triangular dorsal margin. It further differs from
|
||
<taxonomicName id="4C4C4D68FF59D580FB9D73EAFB4FF829" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1045,1183,1963,1989]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF59D580FB9D73EAFB4FF829" box="[1045,1183,1963,1989]" italics="true" pageId="181" pageNumber="182">P.mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF59D580FB2C73EDFB2AF829" bold="true" box="[1188,1274,1963,1989]" pageId="181" pageNumber="182">
|
||
<taxonomicNameLabel id="A20B5782FF59D580FB2C73EDFB2AF829" box="[1188,1274,1963,1989]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and relatively slightly dorsoventrally shorter; from
|
||
<taxonomicName id="4C4C4D68FF59D580FAD9738AFA7CF809" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1361,1452,1996,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="181" pageNumber="182" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF59D580FAD9738AFA7CF809" box="[1361,1452,1996,2021]" italics="true" pageId="181" pageNumber="182">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF59D580FCA273ADFC51F7E9" bold="true" box="[810,897,2027,2053]" pageId="181" pageNumber="182">
|
||
<taxonomicNameLabel id="A20B5782FF59D580FCA273ADFC51F7E9" box="[810,897,2027,2053]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being slightly less transversely compressed and relatively taller, with a less deeply concave entocuneiform facet; from
|
||
<taxonomicName id="4C4C4D68FF5AD583FE9872CDFE73F949" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[272,419,1675,1701]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="182" pageNumber="183" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF5AD583FE9872CDFE73F949" box="[272,419,1675,1701]" italics="true" pageId="182" pageNumber="183">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in having a slightly larger and more cranial facing facet for the mesocuneiform; from
|
||
<taxonomicName id="4C4C4D68FF5AD583FD6772EDFF11F909" baseAuthorityName="Desmarest" baseAuthorityYear="1822" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="182" pageNumber="183" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5AD583FD6772EDFF11F909" italics="true" pageId="182" pageNumber="183">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in having a smaller cranial eminence, more plantarly extensive articulation with the navicular, and a less cranially deflected plantar process; from
|
||
<taxonomicName id="4C4C4D68FF5AD583FDC7734AFD3EF8C9" authorityName="Desmarest" authorityYear="1817" box="[591,750,1803,1829]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="182" pageNumber="183" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5AD583FDC7734AFD3EF8C9" box="[591,750,1803,1829]" italics="true" pageId="182" pageNumber="183">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger and taller, with a more plantarly extensive facet for the navicular; and from
|
||
<taxonomicName id="4C4C4D68FF5AD583FE35730AFDE0F889" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[445,560,1867,1893]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="182" pageNumber="183" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5AD583FE35730AFDE0F889" box="[445,560,1867,1893]" italics="true" pageId="182" pageNumber="183">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and slightly less transversely compressed, with a less concave, slightly more plantarly extensive navicular facet.
|
||
</paragraph>
|
||
<caption id="DF336663FF5AD583FF0071EAFDE7F9C9" ID-DOI="http://doi.org/10.5281/zenodo.10994004" ID-Zenodo-Dep="10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="182" pageNumber="183" startId="182.[136,235,1452,1476]" targetBox="[162,1425,207,1420]" targetPageId="182" targetType="figure">
|
||
<paragraph id="8BF336EBFF5AD583FF0071EAFDE7F9C9" blockId="182.[136,1451,1452,1573]" pageId="182" pageNumber="183">
|
||
<emphasis id="B938EAF9FF5AD583FF0071EAFECEFA29" bold="true" box="[136,286,1452,1477]" pageId="182" pageNumber="183">FIGURE 104.</emphasis>
|
||
right tarsal elements of
|
||
<taxonomicName id="4C4C4D68FF5AD583FD9171EBFD47FA29" box="[537,663,1453,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="182" pageNumber="183" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5AD583FD9171EBFD47FA29" box="[537,663,1453,1477]" italics="true" pageId="182" pageNumber="183">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF5AD583FF3073EDFB34F7C9" blockId="182.[136,778,1675,2085]" lastBlockId="182.[810,1452,1675,2085]" pageId="182" pageNumber="183">
|
||
<emphasis id="B938EAF9FF5AD583FF3073EDFDADF829" box="[184,637,1963,1989]" italics="true" pageId="182" pageNumber="183">
|
||
Metatarsals II and III (
|
||
<figureCitation id="13772A6EFF5AD583FE6A73EDFDA5F829" box="[482,629,1963,1989]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="182" pageNumber="183">Fig. 104a–d</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: both very gracile; III longer than II, both slightly shorter than metatarsal IV; II straight, III very slightly bowed laterally. Proximal end of metatarsal III: articulates with the mesocuneiform, ectocuneiform and metatarsals II and IV; tall and narrow with a smoothly convex ectocuneiform facet; entocuneiform facet small, rounded and situated on the plantar part of the medial surface. Proximal end of metatarsal II: articulates with metatarsal III and possibly with both the entocuneiform and mesocuneiform; relatively small and unflared compared to the metatarsal III proximal end. Shafts transversely compressed proximally; metatarsal III shaft rapidly decreases in height from the proximal end, becomes rounded in cross-section and gently tapers distally; metatarsal II broadens slightly and is un-tapered. Distal ends small, bulbous and rounded. The phalanges are not known.
|
||
</paragraph>
|
||
<caption id="DF336663FF5BD582FF00708AFB70FAE9" ID-DOI="http://doi.org/10.5281/zenodo.10994006" ID-Zenodo-Dep="10994006" httpUri="https://zenodo.org/record/10994006/files/figure.png" pageId="183" pageNumber="184" startId="183.[136,235,1228,1252]" targetBox="[162,1425,200,1195]" targetPageId="183" targetType="figure">
|
||
<paragraph id="8BF336EBFF5BD582FF00708AFB70FAE9" blockId="183.[136,1451,1228,1285]" pageId="183" pageNumber="184">
|
||
<emphasis id="B938EAF9FF5BD582FF00708AFEF1FB09" bold="true" box="[136,289,1228,1253]" pageId="183" pageNumber="184">FIGURE 105.</emphasis>
|
||
surface scan images of left (a–d) proximal, (e–h) middle, and (i–l) distal pedal phalanges IV of
|
||
<taxonomicName id="4C4C4D68FF5BD582FAA3708BFA7BFB09" box="[1323,1451,1229,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="183" pageNumber="184" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5BD582FAA3708BFA7BFB09" box="[1323,1451,1229,1253]" italics="true" pageId="183" pageNumber="184">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
specimen UCMP 70584 in: (a, e, i) dorsal, (b, f, j) plantar, (c, g, k) medial, and (d, h, l) proximal views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF5BD582FF30712DFE03F969" blockId="183.[136,778,1387,2085]" pageId="183" pageNumber="184">
|
||
Metatarsals II and III of
|
||
<taxonomicName id="4C4C4D68FF5BD582FE4A712AFD9AFA69" box="[450,586,1387,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="183" pageNumber="184" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF5BD582FE4A712AFD9AFA69" box="[450,586,1387,1413]" italics="true" pageId="183" pageNumber="184">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differ from those of all compared taxa in having shafts that are less laterally bowed. They further differ from those of
|
||
<taxonomicName id="4C4C4D68FF5BD582FDC671EAFD7CFA29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[590,684,1452,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="183" pageNumber="184" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF5BD582FDC671EAFD7CFA29" box="[590,684,1452,1477]" italics="true" pageId="183" pageNumber="184">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF5BD582FD3971EDFCDAFA29" bold="true" box="[689,778,1451,1477]" pageId="183" pageNumber="184">
|
||
<taxonomicNameLabel id="A20B5782FF5BD582FD3971EDFCDAFA29" box="[689,778,1451,1477]" pageId="183" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being slightly more robust and longer relative to the length of metatarsal IV; from
|
||
<taxonomicName id="4C4C4D68FF5BD582FE5E71AAFDA7F9E9" authorityName="Desmarest" authorityYear="1817" box="[470,631,1515,1541]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="183" pageNumber="184" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF5BD582FE5E71AAFDA7F9E9" box="[470,631,1515,1541]" italics="true" pageId="183" pageNumber="184">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF5BD582FD2671ADFCDAF9E9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[686,778,1515,1541]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="183" pageNumber="184" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF5BD582FD2671ADFCDAF9E9" box="[686,778,1515,1541]" italics="true" pageId="183" pageNumber="184">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being much shorter, more robust and longer relative to length of metatarsal IV, with a taller proximal end for articulation with metatarsal III; and from
|
||
<taxonomicName id="4C4C4D68FF5BD582FDF9720AFD39F989" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[625,745,1611,1637]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="183" pageNumber="184" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF5BD582FDF9720AFD39F989" box="[625,745,1611,1637]" italics="true" pageId="183" pageNumber="184">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more robust.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5BD582FF3072CDFB6BF869" blockId="183.[136,778,1387,2085]" lastBlockId="183.[810,1452,1387,2085]" pageId="183" pageNumber="184">
|
||
<emphasis id="B938EAF9FF5BD582FF3072CDFDC8F949" box="[184,536,1675,1701]" italics="true" pageId="183" pageNumber="184">
|
||
Metatarsal IV (
|
||
<figureCitation id="13772A6EFF5BD582FEF672CDFDC0F949" box="[382,528,1675,1701]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="183" pageNumber="184">Fig. 104a–f</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: short and robust. Proximal end dorsal margin flat to gently rounded; dorsal cuboid facet broad, with the lateral section flat to gently convex and the medial section flat to gently concave; dorsal facet separate from the plantar cuboid facet, separated by a broad, very shallow transverse proximal fossa; plantar cuboid facet small, rounded, slightly proximally projected and tilted dorsally and slightly medially, extends plantarly onto the proximal surface of the plantar tubercle. Facet for the dorsolateral section of the ectocuneiform small, semicircular and distinct, situated against the dorsal margin of the medial surface of the proximal end, facing medially. Plantar tubercle quite small, plantarly projected and slightly proximally deflected. Proximal plantar sesamoid facet small, rounded to squarish and laterally tilted. Facet for metatarsal III indistinct, situated in the shallow, rugose metatarsal III fossa, bordered dorsally by a thin ridge extending plantodistally from the dorsomedial corner of the dorsal cuboid facet. Facet for the ectocuneiform very small, tall and thin, abuts the midpoint of the proximal margin of the medial surface of the shaft; smaller facet for articulation with the plantolateral section of the ectocuneiform occasionally present on the proximal section of the medial surface of the plantar tubercle (see UCMP 70584, L metatarsal IV). Facet for metatarsal V tall, quite deep, distally tilted, gently concave and oblong, projects proximally or proximoplantarly to create one or two small, rounded eminences on the lateral margin of proximal fossa in proximal surface.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF5BD58DFCD273CDFE06F929" blockId="183.[810,1452,1387,2085]" lastBlockId="184.[136,778,1643,2085]" lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184">
|
||
Plantar ridge broad, weakly raised (
|
||
<figureCitation id="13772A6EFF5BD582FB5473CDFA98F849" box="[1244,1352,1931,1957]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="183" pageNumber="184">Fig. 104e</figureCitation>
|
||
), rugose, rounded to square in cross-section, extends distally from the distal base of plantar tubercle to slowly merge with plantar shaft. Shaft flat to very slightly rounded dorsally; height tapers distally, broadens gently to distal end. Distal end large with fossae for collateral ligaments deep and rounded; keel rounded and more plantarly projected than medial and lateral eminences.
|
||
</paragraph>
|
||
<caption id="DF336663FF54D58DFF00718AFABDF9E9" ID-DOI="http://doi.org/10.5281/zenodo.10994008" ID-Zenodo-Dep="10994008" httpUri="https://zenodo.org/record/10994008/files/figure.png" pageId="184" pageNumber="185" startId="184.[136,235,1484,1508]" targetBox="[162,1425,178,1428]" targetPageId="184" targetType="figure">
|
||
<paragraph id="8BF336EBFF54D58DFF00718AFABDF9E9" blockId="184.[136,1451,1484,1541]" pageId="184" pageNumber="185">
|
||
<emphasis id="B938EAF9FF54D58DFF00718AFECDFA09" bold="true" box="[136,285,1484,1509]" pageId="184" pageNumber="185">FIGURE 106.</emphasis>
|
||
surface scan images of right (a–d) proximal pedal phalanges V, and (e–h) articulated middle and partial distal pedal phalanges V of
|
||
<taxonomicName id="4C4C4D68FF54D58DFEAF71ABFE74F9E9" box="[295,420,1517,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF54D58DFEAF71ABFE74F9E9" box="[295,420,1517,1541]" italics="true" pageId="184" pageNumber="185">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
specimen UCMP 45247 in: (a, e) dorsal, (b, f) plantar, (c, g) medial, and (d, h) proximal views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF54D58DFF30728DFB24F889" blockId="184.[136,778,1643,2085]" lastBlockId="184.[810,1452,1643,2085]" pageId="184" pageNumber="185">
|
||
The metatarsal IV of
|
||
<taxonomicName id="4C4C4D68FF54D58DFE69728AFDA9F909" box="[481,633,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF54D58DFE69728AFDA9F909" box="[481,633,1739,1765]" italics="true" pageId="184" pageNumber="185">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
cannot be differentiated from that of
|
||
<taxonomicName id="4C4C4D68FF54D58DFE3972AAFDFEF8E9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[433,558,1771,1797]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF54D58DFE3972AAFDFEF8E9" box="[433,558,1771,1797]" italics="true" pageId="184" pageNumber="185">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
. It differs from that of
|
||
<taxonomicName id="4C4C4D68FF54D58DFF2D734AFF25F8C9" authorityName=", Owen" authorityYear="1874" box="[165,245,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF54D58DFF2D734AFF25F8C9" box="[165,245,1803,1829]" italics="true" pageId="184" pageNumber="185">P. anak</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C4D68FF54D58DFE8A734AFE41F8C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[258,401,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF54D58DFE8A734AFE41F8C9" box="[258,401,1803,1829]" italics="true" pageId="184" pageNumber="185">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF54D58DFE1E734DFE3EF8C9" bold="true" box="[406,494,1803,1829]" pageId="184" pageNumber="185">
|
||
<taxonomicNameLabel id="A20B5782FF54D58DFE1E734DFE3EF8C9" box="[406,494,1803,1829]" pageId="184" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF54D58DFDAA734AFD7CF8C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[546,684,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF54D58DFDAA734AFD7CF8C9" box="[546,684,1803,1829]" italics="true" pageId="184" pageNumber="185">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF54D58DFD3A734DFCDAF8C9" bold="true" box="[690,778,1803,1829]" pageId="184" pageNumber="185">
|
||
<taxonomicNameLabel id="A20B5782FF54D58DFD3A734DFCDAF8C9" box="[690,778,1803,1829]" pageId="184" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and more robust, with separate dorsal and plantar cuboid facets (
|
||
<figureCitation id="13772A6EFF54D58DFE4B730DFDE1F88A" box="[451,561,1867,1894]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="184" pageNumber="185">Fig. 104f</figureCitation>
|
||
) and a less raised plantar ridge; from
|
||
<taxonomicName id="4C4C4D68FF54D58DFEF9732AFE05F869" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[369,469,1900,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF54D58DFEF9732AFE05F869" box="[369,469,1900,1925]" italics="true" pageId="184" pageNumber="185">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF54D58DFE69732DFDEFF869" bold="true" box="[481,575,1899,1925]" pageId="184" pageNumber="185">
|
||
<taxonomicNameLabel id="A20B5782FF54D58DFE69732DFDEFF869" box="[481,575,1899,1925]" pageId="184" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and more robust, with separate dorsal and plantar cuboid facets, a less raised plantar ridge and a shaft with a weaker proximal waist; from
|
||
<taxonomicName id="4C4C4D68FF54D58DFE08738AFE23F809" authorityName="Bartholomai" authorityYear="1978" box="[384,499,1996,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF54D58DFE08738AFE23F809" box="[384,499,1996,2021]" italics="true" pageId="184" pageNumber="185">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being slightly shorter and more robust, with a smaller proximal cuboid fossa; from
|
||
<taxonomicName id="4C4C4D68FF54D58DFF4F7C4DFE88F7C9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[199,344,2059,2085]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF54D58DFF4F7C4DFE88F7C9" box="[199,344,2059,2085]" italics="true" pageId="184" pageNumber="185">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being broader, with separate dorsal and plantar cuboid facets, less narrowing of the proximal shaft, and a more raised plantar ridge; from
|
||
<taxonomicName id="4C4C4D68FF54D58DFA9672CDFAABF949" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1310,1403,1675,1701]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF54D58DFA9672CDFAABF949" box="[1310,1403,1675,1701]" italics="true" pageId="184" pageNumber="185">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF54D58DFCA272EAFC1CF929" authorityName="Desmarest" authorityYear="1817" box="[810,972,1707,1733]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF54D58DFCA272EAFC1CF929" box="[810,972,1707,1733]" italics="true" pageId="184" pageNumber="185">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being much shorter, broader and more robust, with separate dorsal and plantar cuboid facets, a lower plantar ridge, and a more plantarly projected proximal plantar tubercle; and from
|
||
<taxonomicName id="4C4C4D68FF54D58DFB46734AFA94F8C9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1230,1348,1803,1829]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="184" pageNumber="185" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF54D58DFB46734AFA94F8C9" box="[1230,1348,1803,1829]" italics="true" pageId="184" pageNumber="185">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, relatively broader and more robust, with a relatively slightly larger plantar tubercle.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF54D58CFCD2732DFEC6FD69" blockId="184.[810,1452,1643,2085]" lastBlockId="185.[136,778,171,2085]" lastPageId="185" lastPageNumber="186" pageId="184" pageNumber="185">
|
||
<emphasis id="B938EAF9FF54D58DFCD2732DFB49F869" box="[858,1177,1899,1925]" italics="true" pageId="184" pageNumber="185">
|
||
Metatarsal V (
|
||
<figureCitation id="13772A6EFF54D58DFB8C732DFB41F869" box="[1028,1169,1899,1925]" captionStart="FIGURE 104" captionStartId="182.[136,235,1452,1476]" captionTargetBox="[162,1425,207,1420]" captionTargetId="figure-27@182.[151,1436,196,1430]" captionTargetPageId="182" captionText="FIGURE 104. right tarsal elements of P. otibandus: (a–d) articulated cuboid, navicular, ectocuneiform, and metatarsals II–IV of UCMP 45247 in (a) dorsal, (b) plantar, (c) lateral, and (d) medial views; (e–f) surface scan images of metatarsal IV of UCMP 70584 in (e) lateral, and (f) proximal views; and (g–k) surface scan images of metatarsal V of UCMP 70585 in (g) dorsal, (h) plantar, (i) medial, (j) lateral, and (k) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994004" httpUri="https://zenodo.org/record/10994004/files/figure.png" pageId="184" pageNumber="185">
|
||
Fig.
|
||
<quantity id="4CB49B0EFF54D58DFBB5732DFBA5F869" box="[1085,1141,1899,1925]" metricMagnitude="-1" metricUnit="kg" metricValue="1.04" pageId="184" pageNumber="185" unit="g" value="104.0">104g</quantity>
|
||
–k
|
||
</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: very short, robust and transversely compressed with a slight medial tilt; curves slightly laterally distally; slightly arched in lateral view; length to distal facet width index ~4–4.7. Proximolateral process blunt, rugose, proximodistally quite short and transversely compressed. Facet for the cuboid very broad, gently concave and slightly raised at distal margin, mostly proximal facing; extends from proximal surface of the medial plantar tubercle across proximal surface of the shaft onto base of dorsomedial surface of proximolateral process and dorsolaterally over dorsal surface of the base of proximolateral process. Facet for metatarsal IV broad, gently convex, slightly raised and rounded to square; proximally abuts facet for the cuboid, extends medially onto dorsomedial surface of the medial plantar tubercle. Lateral plantar tuberosity broad, long and rugose, separated from medial plantar tubercle by narrow, shallow channel. Medial plantar tubercle very small, sometimes indistinct, gently medially projected. Distal end broad; medial fossa for the collateral ligaments larger and deeper than the lateral fossa.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF55D58CFF3076CDFD13FA29" blockId="185.[136,778,171,2085]" pageId="185" pageNumber="186">
|
||
The metatarsal V of
|
||
<taxonomicName id="4C4C4D68FF55D58CFE2776CAFDEEFD49" box="[431,574,651,677]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF55D58CFE2776CAFDEEFD49" box="[431,574,651,677]" italics="true" pageId="185" pageNumber="186">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF55D58CFF2576EAFED5FD29" authorityName=", Owen" authorityYear="1874" box="[173,261,683,709]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF55D58CFF2576EAFED5FD29" box="[173,261,683,709]" italics="true" pageId="185" pageNumber="186">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with a relatively smaller proximolateral process and a relatively smaller, less proximomedially projected medial plantar tubercle; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFF00774AFEC8FCC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,280,779,805]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF55D58CFF00774AFEC8FCC9" box="[136,280,779,805]" italics="true" pageId="185" pageNumber="186">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF55D58CFE96774DFEA7FCC9" bold="true" box="[286,375,779,805]" pageId="185" pageNumber="186">
|
||
<taxonomicNameLabel id="A20B5782FF55D58CFE96774DFEA7FCC9" box="[286,375,779,805]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and narrower, with a smaller, slightly narrower cuboid facet, deeper plantar groove,and a larger lateral plantar tuberosity;from
|
||
<taxonomicName id="4C4C4D68FF55D58CFD27770AFCDAFC89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[687,778,844,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF55D58CFD27770AFCDAFC89" box="[687,778,844,869]" italics="true" pageId="185" pageNumber="186">P.viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF55D58CFF00772DFF36FC69" bold="true" box="[136,230,875,901]" pageId="185" pageNumber="186">
|
||
<taxonomicNameLabel id="A20B5782FF55D58CFF00772DFF36FC69" box="[136,230,875,901]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, shorter and less transversely compressed, with a broader, more medially situated cuboid facet and a more medially extensive metatarsal IV facet that is less distinct from the cuboid facet; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFD06778AFCDAFC09" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[654,778,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF55D58CFD06778AFCDAFC09" box="[654,778,971,997]" italics="true" pageId="185" pageNumber="186">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in being slightly more gracile and more transversely compressed, with dorsoplantarly shorter proximolateral process, proximodistally longer metatarsal IV facet and larger medial plantar tubercle; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFD96700AFD79FB89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[542,681,1099,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF55D58CFD96700AFD79FB89" box="[542,681,1099,1125]" italics="true" pageId="185" pageNumber="186">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF55D58CFD38700DFCDAFB89" bold="true" box="[688,778,1099,1125]" pageId="185" pageNumber="186">
|
||
<taxonomicNameLabel id="A20B5782FF55D58CFD38700DFCDAFB89" box="[688,778,1099,1125]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being shorter, with more dorsally projected cuboid facet and slightly broader distal end; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFDD670CDFD3EFB49" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[606,750,1163,1189]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF55D58CFDD670CDFD3EFB49" box="[606,750,1163,1189]" italics="true" pageId="185" pageNumber="186">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being shorter and broader, lacking slight kink of arch of the shaft immediately proximal to midpoint in lateral view, with a larger medial plantar tubercle and lateral plantar tuberosity, and a deeper, more distinct plantar groove; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFF4F716DFEF5FAA9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[199,293,1323,1349]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF55D58CFF4F716DFEF5FAA9" box="[199,293,1323,1349]" italics="true" pageId="185" pageNumber="186">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF55D58CFED7716AFDD1FAA9" authorityName="Desmarest" authorityYear="1817" box="[351,513,1323,1349]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF55D58CFED7716AFDD1FAA9" box="[351,513,1323,1349]" italics="true" pageId="185" pageNumber="186">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being much shorter, broader, less transversely compressed, and less arched in lateral view, with a larger medial plantar tubercle and a deeper, more distinct plantar groove; and from
|
||
<taxonomicName id="4C4C4D68FF55D58CFD1E71CAFCDAFA49" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[662,778,1419,1445]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF55D58CFD1E71CAFCDAFA49" box="[662,778,1419,1445]" italics="true" pageId="185" pageNumber="186">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, relatively broader and more robust.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF55D58CFF30718DFCB6FF09" blockId="185.[136,778,171,2085]" lastBlockId="185.[810,1452,171,2085]" pageId="185" pageNumber="186">
|
||
<emphasis id="B938EAF9FF55D58CFF30718DFDE6FA09" box="[184,566,1483,1509]" italics="true" pageId="185" pageNumber="186">
|
||
Pedal phalanges (
|
||
<figureCitation id="13772A6EFF55D58CFE0A718DFE30FA09" box="[386,480,1483,1509]" captionStart="FIGURE 105" captionStartId="183.[136,235,1228,1252]" captionTargetBox="[162,1425,200,1195]" captionTargetId="figure-17@183.[151,1436,189,1206]" captionTargetPageId="183" captionText="FIGURE 105. surface scan images of left (a–d) proximal, (e–h) middle, and (i–l) distal pedal phalanges IV of P. otibandus specimen UCMP 70584 in: (a, e, i) dorsal, (b, f, j) plantar, (c, g, k) medial, and (d, h, l) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994006" httpUri="https://zenodo.org/record/10994006/files/figure.png" pageId="185" pageNumber="186">Fig. 105</figureCitation>
|
||
&
|
||
<figureCitation id="13772A6EFF55D58CFD8B718DFDFEFA09" box="[515,558,1483,1509]" captionStart="FIGURE 106" captionStartId="184.[136,235,1484,1508]" captionTargetBox="[162,1425,178,1428]" captionTargetId="figure-27@184.[151,1436,167,1439]" captionTargetPageId="184" captionText="FIGURE 106. surface scan images of right (a–d) proximal pedal phalanges V, and (e–h) articulated middle and partial distal pedal phalanges V of P. otibandus specimen UCMP 45247 in: (a, e) dorsal, (b, f) plantar, (c, g) medial, and (d, h) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994008" httpUri="https://zenodo.org/record/10994008/files/figure.png" pageId="185" pageNumber="186">106</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: Proximal phalanx IV: short, robust and slightly dorsoplantarly compressed, with a slight waist on the shaft. Proximal end domed, with the proximal plantar tubercles low and rounded; proximal articular facet broad and gently concave with a rounded dorsal margin. Distal end with large, very shallow fossae for the collateral ligaments; trochlea gently concave. Middle phalanx IV: short and broad; proximal end broad, with the proximal plantar tubercles very low and projected slightly proximally. Proximal articular surface dorsally rounded, concave and tilted slightly dorsally. Shaft height decreases distally, with a very slight waist. Distal end with quite deep fossae for the collateral ligaments; trochlea narrow and very slightly concave dorsally, deepens and broadens plantarly. Distal phalanx IV: quite short, broad and robust; proximal articular facet concave and roughly pentagonal, forms a rounded, slightly proximodorsally projected dorsal peak. Flexor tubercle large and plantarly projected, oval in plantar view. Shaft with a rounded dorsal peak (
|
||
<figureCitation id="13772A6EFF55D58CFC3F74EDFBCFFF2A" box="[951,1055,171,198]" captionStart="FIGURE 105" captionStartId="183.[136,235,1228,1252]" captionTargetBox="[162,1425,200,1195]" captionTargetId="figure-17@183.[151,1436,189,1206]" captionTargetPageId="183" captionText="FIGURE 105. surface scan images of left (a–d) proximal, (e–h) middle, and (i–l) distal pedal phalanges IV of P. otibandus specimen UCMP 70584 in: (a, e, i) dorsal, (b, f, j) plantar, (c, g, k) medial, and (d, h, l) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994006" httpUri="https://zenodo.org/record/10994006/files/figure.png" pageId="185" pageNumber="186">Fig. 105i</figureCitation>
|
||
), curves gently downward in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF55D58CFCD274ADFC76FC29" blockId="185.[810,1452,171,2085]" pageId="185" pageNumber="186">Proximal phalanx V: quite tall, robust and asymmetrical; shaft laterally deflected relative to the proximal surface. Proximal end slightly broader than tall; plantar tubercles large, rounded and rugose, lateral tubercle projected plantolaterally past the articular surface; proximal articular surface round, gently concave and medially displaced in proximal view. Shaft rounded and triangular in cross-section. Distal end with very shallow fossae for the collateral ligaments; trochlea broad and shallow, and articular surface not extensive dorsally. Middle phalanx V: very short, broad and very dorsoplantarly compressed. Proximal end with very low plantar tubercles; proximal articular surface gently concave and tilted strongly dorsally. Shaft very short, decreases in height distally, with no waist. Distal end with fossae for collateral ligaments very shallow and tall. Distal phalanx V: tall, robust and asymmetrical, with the dorsal peak of the shaft and the proximal end medially displaced; small, deep fossae for the collateral ligaments. Plantar tubercle large and rounded in plantar view. Shaft with a rounded dorsal peak and a strongly convex plantar margin in cross-section. Distal end abraded in available specimens.</paragraph>
|
||
<paragraph id="8BF336EBFF55D58CFCD2778DFCA7F7C9" blockId="185.[810,1452,171,2085]" pageId="185" pageNumber="186">
|
||
The pedal phalanges of
|
||
<taxonomicName id="4C4C4D68FF55D58CFBE9778AFB3AFC09" box="[1121,1258,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF55D58CFBE9778AFB3AFC09" box="[1121,1258,971,997]" italics="true" pageId="185" pageNumber="186">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
differ from those of
|
||
<taxonomicName id="4C4C4D68FF55D58CFCC177AAFC4BFBE9" authorityName=", Owen" authorityYear="1874" box="[841,923,1003,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF55D58CFCC177AAFC4BFBE9" box="[841,923,1003,1029]" italics="true" pageId="185" pageNumber="186">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being slightly smaller, with middle phalanx IV less dorsoplantarly compressed (
|
||
<figureCitation id="13772A6EFF55D58CFB3B704DFACEFBCA" box="[1203,1310,1035,1062]" captionStart="FIGURE 105" captionStartId="183.[136,235,1228,1252]" captionTargetBox="[162,1425,200,1195]" captionTargetId="figure-17@183.[151,1436,189,1206]" captionTargetPageId="183" captionText="FIGURE 105. surface scan images of left (a–d) proximal, (e–h) middle, and (i–l) distal pedal phalanges IV of P. otibandus specimen UCMP 70584 in: (a, e, i) dorsal, (b, f, j) plantar, (c, g, k) medial, and (d, h, l) proximal views." figureDoi="http://doi.org/10.5281/zenodo.10994006" httpUri="https://zenodo.org/record/10994006/files/figure.png" pageId="185" pageNumber="186">Fig. 105h</figureCitation>
|
||
) and lacking large, transversely flared plantar tubercles, distal phalanx IV with a more rounded dorsal peak, and proximal phalanx V with a less distinct waist; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFB3B702AFA96FB69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1203,1350,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF55D58CFB3B702AFA96FB69" box="[1203,1350,1131,1157]" italics="true" pageId="185" pageNumber="186">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF55D58CFAC7702DFA7BFB69" bold="true" box="[1359,1451,1131,1157]" pageId="185" pageNumber="186">
|
||
<taxonomicNameLabel id="A20B5782FF55D58CFAC7702DFA7BFB69" box="[1359,1451,1131,1157]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and shorter, with proximal phalanx IV with a narrower distal end, and middle phalanx IV with a narrower trochlea; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFBE2708AFB1FFB09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1130,1231,1228,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF55D58CFBE2708AFB1FFB09" box="[1130,1231,1228,1253]" italics="true" pageId="185" pageNumber="186">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF55D58CFB54708DFAECFB09" bold="true" box="[1244,1340,1227,1253]" pageId="185" pageNumber="186">
|
||
<taxonomicNameLabel id="A20B5782FF55D58CFB54708DFAECFB09" box="[1244,1340,1227,1253]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being slightly smaller, with proximal phalanx IV shorter with a broader waist, distal phalanx IV more dorsoplantarly compressed with a more rounded dorsal peak and a less plantarly curved shaft, proximal phalanx V with a less concave proximal articular surface, a more laterally deflected shaft, and a narrower trochlea, middle phalanx V shorter with a less medially tilted proximal surface, and distal phalanx V with a more rounded dorsal peak; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFCA271AAFC4DF9E9" authorityName="Bartholomai" authorityYear="1978" box="[810,925,1516,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF55D58CFCA271AAFC4DF9E9" box="[810,925,1516,1541]" italics="true" pageId="185" pageNumber="186">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having relatively narrower middle phalanx IV across the proximal end and a slightly broader distal phalanx IV with a more rounded dorsal peak; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFA1F726AFC4AF989" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF55D58CFA1F726AFC4AF989" italics="true" pageId="185" pageNumber="186">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF55D58CFC2C720DFBD0F989" bold="true" box="[932,1024,1611,1637]" pageId="185" pageNumber="186">
|
||
<taxonomicNameLabel id="A20B5782FF55D58CFC2C720DFBD0F989" box="[932,1024,1611,1637]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having proximal phalanx IV with a less proximodorsally extensive distal articular surface; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFCE372CDFC2FF949" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[875,1023,1675,1701]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF55D58CFCE372CDFC2FF949" box="[875,1023,1675,1701]" italics="true" pageId="185" pageNumber="186">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being shorter, lower, broader and more robust, with broader, shallower trochleae, middle phalanges with more dorsally tilted proximal surfaces, and distal phalanges with much more rounded, less pointed dorsal peaks; from
|
||
<taxonomicName id="4C4C4D68FF55D58CFB80734DFBB7F8C9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1032,1127,1803,1829]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF55D58CFB80734DFBB7F8C9" box="[1032,1127,1803,1829]" italics="true" pageId="185" pageNumber="186">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF55D58CFB2A734AFA95F8C9" authorityName="Desmarest" authorityYear="1817" box="[1186,1349,1803,1829]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF55D58CFB2A734AFA95F8C9" box="[1186,1349,1803,1829]" italics="true" pageId="185" pageNumber="186">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being shorter, lower, broader and much more robust, particularly the middle phalanges, with proximal phalanx IV having a broader waist, the middle phalanges with a more dorsally tilted proximal surface, and distal phalanges with a more rounded, less pointed dorsal peak in cross-section and a more plantarly curved shaft in lateral view; and from
|
||
<taxonomicName id="4C4C4D68FF55D58CFCA273AAFC72F7E9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[810,930,2027,2053]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="185" pageNumber="186" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF55D58CFCA273AAFC72F7E9" box="[810,930,2027,2053]" italics="true" pageId="185" pageNumber="186">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, relatively broader and more robust.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF56D58FFF0074EDFBB5F7E8" pageId="186" pageNumber="187" type="discussion">
|
||
<paragraph id="8BF336EBFF56D58FFF0074EDFED4FF29" blockId="186.[136,260,171,197]" box="[136,260,171,197]" pageId="186" pageNumber="187">
|
||
<heading id="D0BB8187FF56D58FFF0074EDFED4FF29" bold="true" box="[136,260,171,197]" fontSize="11" level="1" pageId="186" pageNumber="187" reason="1">
|
||
<emphasis id="B938EAF9FF56D58FFF0074EDFED4FF29" bold="true" box="[136,260,171,197]" pageId="186" pageNumber="187">Remarks:</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFF0074ACFD4EFEE8" blockId="186.[136,779,234,1029]" box="[136,670,234,261]" pageId="186" pageNumber="187">
|
||
<emphasis id="B938EAF9FF56D58FFF0074ACFD4EFEE8" bold="true" box="[136,670,234,261]" pageId="186" pageNumber="187">
|
||
<emphasis id="B938EAF9FF56D58FFF0074ACFEF0FEE8" bold="true" box="[136,288,234,261]" italics="true" pageId="186" pageNumber="187">
|
||
<typeStatus id="54F78849FF56D58FFF0074ACFED1FEE9" box="[136,257,234,261]" pageId="186" pageNumber="187" type="holotype">Holotypes</typeStatus>
|
||
of
|
||
</emphasis>
|
||
<taxonomicName id="4C4C4D68FF56D58FFEA074ADFE2DFEE8" authorityName="Bartholomai" authorityYear="1973" box="[296,509,234,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">P. chinchillaensis</taxonomicName>
|
||
<emphasis id="B938EAF9FF56D58FFD8C74ACFDE2FEE9" bold="true" box="[516,562,234,261]" italics="true" pageId="186" pageNumber="187">and</emphasis>
|
||
<taxonomicName id="4C4C4D68FF56D58FFDB174ADFD4EFEE8" authorityName=", Bartholomai" authorityYear="1973" box="[569,670,234,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="devisi">P. devisi</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFF00754DFE3EFBE9" blockId="186.[136,779,234,1029]" pageId="186" pageNumber="187">
|
||
During the printing of
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFE23754DFD43FEC9" author="Bartholomai, A." box="[427,659,267,293]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
, an error occurred wherein the images of the
|
||
<typeStatus id="54F78849FF56D58FFDD9756DFD6EFEA9" box="[593,702,299,325]" pageId="186" pageNumber="187" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF56D58FFD7D756AFEE0FE89" authorityName="Bartholomai" authorityYear="1973" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFD7D756AFEE0FE89" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF56D58FFEEE750AFE13FE89" authorityName=", Bartholomai" authorityYear="1973" box="[358,451,331,357]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF56D58FFEEE750AFE13FE89" box="[358,451,331,357]" italics="true" pageId="186" pageNumber="187">P. devisi</emphasis>
|
||
</taxonomicName>
|
||
and their respective captions were inadvertently switched. This resulted in the image of the
|
||
<typeStatus id="54F78849FF56D58FFF3975CDFEC3FE49" box="[177,275,395,421]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
for
|
||
<taxonomicName id="4C4C4D68FF56D58FFEC975CAFDD4FE49" authorityName="Bartholomai" authorityYear="1973" box="[321,516,395,421]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFEC975CAFDD4FE49" box="[321,516,395,421]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFD9B75CDFD32FE49" author="Bartholomai, A." box="[531,738,395,421]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai 1973</bibRefCitation>
|
||
, pl. 23, figs 3 & 4) being labelled as the
|
||
<typeStatus id="54F78849FF56D58FFD9075EDFDAAFE29" box="[536,634,427,453]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
for
|
||
<taxonomicName id="4C4C4D68FF56D58FFD2F75EAFCD2FE29" authorityName=", Bartholomai" authorityYear="1973" box="[679,770,427,453]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF56D58FFD2F75EAFCD2FE29" box="[679,770,427,453]" italics="true" pageId="186" pageNumber="187">P. devisi</emphasis>
|
||
</taxonomicName>
|
||
, and the image of the
|
||
<typeStatus id="54F78849FF56D58FFEF3758DFE0DFE09" box="[379,477,459,485]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
for
|
||
<taxonomicName id="4C4C4D68FF56D58FFD86758AFDBCFE09" authorityName=", Bartholomai" authorityYear="1973" box="[526,620,459,485]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF56D58FFD86758AFDBCFE09" box="[526,620,459,485]" italics="true" pageId="186" pageNumber="187">P. devisi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFDF4758DFF13FDE9" author="Bartholomai, A." pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai 1973</bibRefCitation>
|
||
, pl. 21, figs 3 & 4), being labelled as the
|
||
<typeStatus id="54F78849FF56D58FFD2075ADFCDAFDE9" box="[680,778,491,517]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
for
|
||
<taxonomicName id="4C4C4D68FF56D58FFF3D764AFEAFFDC9" authorityName="Bartholomai" authorityYear="1973" box="[181,383,523,549]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFF3D764AFEAFFDC9" box="[181,383,523,549]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
. Bartholomai (1975) included an erratum correcting this mistake. During I.A.R.K.’s visit to the palaeontological collection of the
|
||
<collectingRegion id="4988F809FF56D58FFD0E760DFCDAFD89" box="[646,778,587,613]" country="Australia" name="Queensland" pageId="186" pageNumber="187">Queensland</collectingRegion>
|
||
Museum in late 2018, the curator, Dr Scott Hocknull, and I.A.R.K. noted that the registration numbers of each
|
||
<typeStatus id="54F78849FF56D58FFF0076EDFF3AFD29" box="[136,234,683,709]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
reflected those assigned to them incorrectly in plates 21 & 23 of
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFE0E768DFDBBFD09" author="Bartholomai, A." box="[390,619,715,741]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
. Dr Hocknull reviewed the situation, and, informed by the apparent intent of
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFF7B774DFE03FCC9" author="Bartholomai, A." box="[243,467,779,805]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
and by the descriptions of the specimens in the fossil register of the
|
||
<collectingRegion id="4988F809FF56D58FFD0D776DFCD9FCA9" box="[645,777,811,837]" country="Australia" name="Queensland" pageId="186" pageNumber="187">Queensland</collectingRegion>
|
||
Museum, altered the number of each type to its original designation in the species diagnosis of
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFDF6772DFF06FC49" author="Bartholomai, A." pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
. The official stance of the museum is that the
|
||
<typeStatus id="54F78849FF56D58FFF0077EDFF3AFC29" box="[136,234,939,965]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF56D58FFE8777EAFE02FC29" authorityName="Bartholomai" authorityYear="1973" box="[271,466,939,965]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFE8777EAFE02FC29" box="[271,466,939,965]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
has the registration number QM F5246 and that the
|
||
<typeStatus id="54F78849FF56D58FFE3A778DFDC4FC09" box="[434,532,971,997]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF56D58FFDCD778AFD78FC09" authorityName=", Bartholomai" authorityYear="1973" box="[581,680,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF56D58FFDCD778AFD78FC09" box="[581,680,971,997]" italics="true" pageId="186" pageNumber="187">P. devisi</emphasis>
|
||
</taxonomicName>
|
||
has the registration number QM F4710.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFF00706CFD1BFBA8" blockId="186.[136,779,1066,2085]" box="[136,715,1066,1093]" pageId="186" pageNumber="187">
|
||
<emphasis id="B938EAF9FF56D58FFF00706CFD1BFBA8" bold="true" box="[136,715,1066,1093]" pageId="186" pageNumber="187">
|
||
<emphasis id="B938EAF9FF56D58FFF00706CFEC8FBA8" bold="true" box="[136,280,1066,1093]" italics="true" pageId="186" pageNumber="187">Identities of</emphasis>
|
||
<taxonomicName id="4C4C4D68FF56D58FFEA8706DFE6AFBA8" box="[288,442,1066,1093]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">P. otibandus</taxonomicName>
|
||
<emphasis id="B938EAF9FF56D58FFE49706CFE3FFBA9" bold="true" box="[449,495,1066,1093]" italics="true" pageId="186" pageNumber="187">and</emphasis>
|
||
<taxonomicName id="4C4C4D68FF56D58FFE7E706DFD1BFBA8" authority="Prior" authorityName="Prior" box="[502,715,1066,1093]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">P. chinchillaensis</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFF00700DFDEDF96A" blockId="186.[136,779,1066,2085]" pageId="186" pageNumber="187">
|
||
Prior to this study, there was some cause to doubt the separate taxonomic identities of
|
||
<taxonomicName id="4C4C4D68FF56D58FFD91702AFD7CFB69" box="[537,684,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFD91702AFD7CFB69" box="[537,684,1131,1157]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF56D58FFD7D702AFEE0FB49" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFD7D702AFEE0FB49" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C4C4D68FF56D58FFECF70CDFD8DFB49" box="[327,605,1163,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFECF70CDFD8DFB49" box="[327,605,1163,1189]" italics="true" pageId="186" pageNumber="187">Protemnodon otibandus</emphasis>
|
||
</taxonomicName>
|
||
was the first species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFE8870EDFE47FB29" authorityName="Owen" authorityYear="1874" box="[256,407,1195,1221]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFE8870EDFE47FB29" box="[256,407,1195,1221]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
of undoubted Pliocene age to be described, and the first from New
|
||
<collectingCountry id="F35B767BFF56D58FFD9E708DFDB7FB09" box="[534,615,1227,1253]" name="Guinea-Bissau" pageId="186" pageNumber="187">Guinea</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFDF0708DFD2DFB0A" author="Plane, M. D." box="[632,765,1227,1254]" pageId="186" pageNumber="187" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane 1967</bibRefCitation>
|
||
). Later,
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFF5870ADFEB1FAE9" author="Plane, M." box="[208,353,1259,1285]" pageId="186" pageNumber="187" pagination="33 - 36" refId="ref153379" refString="Plane, M. (1972) A New Guinea fossil macropodid (Marsupialia) from the marine Pliocene of Victoria, Australia. Memoirs of the National Museum of Victoria, 33, 33 - 36. https: // doi. org / 10.24199 / j. mmv. 1972.33.03" type="journal article" year="1972">Plane (1972)</bibRefCitation>
|
||
tentatively allocated a partial maxilla from Lake Tyers, southeastern Victoria, to
|
||
<taxonomicName id="4C4C4D68FF56D58FFDFE714AFCD2FAC9" box="[630,770,1291,1317]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFDFE714AFCD2FAC9" box="[630,770,1291,1317]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
. Despite this, when
|
||
<taxonomicName id="4C4C4D68FF56D58FFEE9716AFDF5FAA9" box="[353,549,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFEE9716AFDF5FAA9" box="[353,549,1323,1349]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
was described from the Pliocene material of the Chinchilla Sand,
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFDF6710DFF03FA69" author="Bartholomai, A." pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
made comparisons with all species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFDFA712DFCD9FA69" authorityName="Owen" authorityYear="1874" box="[626,777,1387,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFDFA712DFCD9FA69" box="[626,777,1387,1413]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
recognised at the time with the exception of
|
||
<emphasis id="B938EAF9FF56D58FFDF171CAFCD9FA49" box="[633,777,1419,1445]" italics="true" pageId="186" pageNumber="187">
|
||
<taxonomicName id="4C4C4D68FF56D58FFDF171CAFCD5FA49" box="[633,773,1419,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">P. otibandus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the species with which it was most similar.
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFD0E71EDFEF7FA09" author="Flannery, T. F. & Archer, M." pageId="186" pageNumber="187" pagination="357 - 383" refId="ref146359" refString="Flannery, T. F. & Archer, M. (1984) The macropodoids (Marsupialia) of the early Pliocene Bow Local Fauna, central eastern New South Wales. The Australian Zoologist, 21 (4), 357 - 383." type="journal article" year="1984">Flannery & Archer (1984)</bibRefCitation>
|
||
noted the lack of a urocrista on the anterior upper molars of
|
||
<taxonomicName id="4C4C4D68FF56D58FFEC971AAFDD5F9E9" box="[321,517,1515,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFEC971AAFDD5F9E9" box="[321,517,1515,1541]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
, the presence of which was implied therein to be autapomorphic in
|
||
<taxonomicName id="4C4C4D68FF56D58FFDF0724AFCD2F9C9" box="[632,770,1547,1573]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFDF0724AFCD2F9C9" box="[632,770,1547,1573]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
, though
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFF68726DFE49F9A9" author="Flannery, T. F." box="[224,409,1579,1605]" pageId="186" pageNumber="187" pagination="39 - 48" refId="ref146287" refString="Flannery, T. F. (1994) The fossil land mammal record of New Guinea: a review. Science in New Guinea, 20 (1), 39 - 48." type="journal article" year="1994">Flannery (1994)</bibRefCitation>
|
||
stated ‘
|
||
<taxonomicName id="4C4C4D68FF56D58FFE71726DFCD9F9A9" box="[505,777,1579,1605]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFE71726DFCD9F9A9" box="[505,777,1579,1605]" italics="true" pageId="186" pageNumber="187">Protemnodon otibandus</emphasis>
|
||
</taxonomicName>
|
||
is extremely similar in morphology to
|
||
<taxonomicName id="4C4C4D68FF56D58FFDB6720AFCD3F989" box="[574,771,1611,1637]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFDB6720AFCD3F989" box="[574,771,1611,1637]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
. They may even be conspecific’ (p. 45).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFF3072CDFC00FB5A" blockId="186.[136,779,1066,2085]" lastBlockId="186.[810,1452,171,2052]" pageId="186" pageNumber="187">
|
||
The New Guinean material of
|
||
<taxonomicName id="4C4C4D68FF56D58FFDA972CAFD7FF949" box="[545,687,1675,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFDA972CAFD7FF949" box="[545,687,1675,1701]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
is very restricted in geographic area and time, contrasting with other species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFEC2728DFE31F909" authorityName="Owen" authorityYear="1874" box="[330,481,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFEC2728DFE31F909" box="[330,481,1739,1765]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
, which are sampled over longer time intervals across wider geographic areas. With the exception of a single maxilla of probable Pliocene age preserving P3–M3, from near Koroba in the
|
||
<collectingRegion id="4988F809FF56D58FFD5D736DFF3CF889" country="Papua New Guinea" name="Hela" pageId="186" pageNumber="187">Hela Province</collectingRegion>
|
||
of montane central
|
||
<collectingCountry id="F35B767BFF56D58FFE78730DFDF8F889" box="[496,552,1867,1893]" name="Papua New Guinea" pageId="186" pageNumber="187">PNG</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFDB7730DFD2DF889" author="Flannery, T. F." box="[575,765,1867,1893]" pageId="186" pageNumber="187" refId="ref146103" refString="Flannery, T. F. (1990 a) Mammals of New Guinea. Robert Brown and Associates, Brisbane, 568 pp." type="book" year="1990">Flannery 1990a</bibRefCitation>
|
||
), all New Guinean material of
|
||
<taxonomicName id="4C4C4D68FF56D58FFE6E732AFDA4F869" box="[486,628,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFE6E732AFDA4F869" box="[486,628,1899,1925]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
comes from the Awe LF of the Otibanda Fm. in
|
||
<collectingRegion id="4988F809FF56D58FFDB373CDFCD5F849" box="[571,773,1931,1957]" country="Papua New Guinea" name="Morobe" pageId="186" pageNumber="187">Morobe Province</collectingRegion>
|
||
, eastern
|
||
<collectingCountry id="F35B767BFF56D58FFF6873EDFEC8F829" box="[224,280,1963,1989]" name="Papua New Guinea" pageId="186" pageNumber="187">PNG</collectingCountry>
|
||
(
|
||
<figureCitation id="13772A6EFF56D58FFEA273EDFEBCF829" box="[298,364,1963,1989]" captionStart="FIGURE 1" captionStartId="5.[136,235,1388,1412]" captionTargetBox="[151,1436,178,1363]" captionTargetId="figure-17@5.[151,1436,178,1363]" captionTargetPageId="5" captionText="FIGURE 1. map showing locations of Pliocene deposits inAustralia and New Guinea yielding material of species of Protemnodon, including that not confidently referable to species level. 1, Mogorafugwa, Hela Province; 2, Lower Kikori River, Gulf Province; 3, Otibanda, Morobe Province; 4, Toolapinna Waterhole, Warburton River; 5, Site 4, Lake Kanunka; 6, Lawson–Daily Quarry, Lake Palankarinna; 7, Bluff Downs, Allingham; 8, Rifle Range/Sand Scree locality/Chinchilla, Chinchilla Sands; 9, Bow, Hunter Valley; 10, Big Sink, Wellington Valley; 11, Hamilton, Glenelg Shire; 12, Lake Tyers, East Gippsland Shire." figureDoi="http://doi.org/10.5281/zenodo.10993777" httpUri="https://zenodo.org/record/10993777/files/figure.png" pageId="186" pageNumber="187">Fig. 1</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFE0073EDFDDCF829" author="Plane, M. D." box="[392,524,1963,1989]" pageId="186" pageNumber="187" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFD9273EDFD13F829" author="Flannery, T. F." box="[538,707,1963,1989]" pageId="186" pageNumber="187" pagination="39 - 48" refId="ref146287" refString="Flannery, T. F. (1994) The fossil land mammal record of New Guinea: a review. Science in New Guinea, 20 (1), 39 - 48." type="journal article" year="1994">Flannery 1994</bibRefCitation>
|
||
). This sample, by the standards of the better-sampled
|
||
<taxonomicName id="4C4C4D68FF56D58FFD26738AFCD2F809" authorityName=", Owen" authorityYear="1874" box="[686,770,1995,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF56D58FFD26738AFCD2F809" box="[686,770,1995,2021]" italics="true" pageId="186" pageNumber="187">P. anak</emphasis>
|
||
</taxonomicName>
|
||
, shows tightly clustered dental dimensions and a more uniform craniodental and postcranial morphology, insofar as both are known. That said, various features that have been described as diagnostic of
|
||
<taxonomicName id="4C4C4D68FF56D58FFB16748AFAFBFF09" box="[1182,1323,203,229]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFB16748AFAFBFF09" box="[1182,1323,203,229]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
are in fact variable within the type series.
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFB0374AAFACCFEEB" author="Plane, M. D." box="[1163,1308,236,263]" pageId="186" pageNumber="187" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane (1967)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C4C4D68FF56D58FFA1F74ABFC48FECA" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFA1F74ABFC48FECA" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
as having a distinct buccal cuspid in the talonid basin of the m1. However, this cuspid is variable within the
|
||
<typeStatus id="54F78849FF56D58FFCD97508FC6CFE84" box="[849,956,334,360]" pageId="186" pageNumber="187" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF56D58FFC567508FBB6FE8B" box="[990,1126,333,359]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFC567508FBB6FE8B" box="[990,1126,333,359]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
—for
|
||
<emphasis id="B938EAF9FF56D58FFB217508FB01FE8B" box="[1193,1233,334,359]" italics="true" pageId="186" pageNumber="187">e.g.</emphasis>
|
||
, extremely reduced in UCMP 69896 and absent from UCMP 45246—and thus its absence from the
|
||
<typeStatus id="54F78849FF56D58FFBDF75C9FB69FE45" box="[1111,1209,399,425]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF56D58FFB6D75C9FA7BFE45" box="[1253,1451,399,425]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFB6D75C9FA7BFE45" box="[1253,1451,399,425]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
is not considered taxonomically significant. The dental morphology of
|
||
<taxonomicName id="4C4C4D68FF56D58FFC787597FB6FFE06" box="[1008,1215,464,490]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFC787597FB6FFE06" box="[1008,1215,464,490]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFCA275B6FBD9FDE6" author="Bartholomai, A." box="[810,1033,496,522]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
in the species diagnosis as differing from other members of the genus in having relatively curved molar links, a shallow mandibular ramus and swollen anterolingual protolophid bases. It is not clear whether the phrase ‘molar links’ refers to the upper or lower dentition, or which crista(e) and/or cristid(s) in particular are more curved in this taxon. The paracristid and cristid obliqua on the lower molars were described as curved in the species description, but this is a feature of all species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFC6A7753FBA9FCC3" authorityName="Owen" authorityYear="1874" box="[994,1145,789,815]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFC6A7753FBA9FCC3" box="[994,1145,789,815]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
, and neither the paracristid nor cristid obliqua in any specimens referred to
|
||
<taxonomicName id="4C4C4D68FF56D58FFA1F7771FC02FC9C" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFA1F7771FC02FC9C" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFC777710FB0FFC9C" author="Bartholomai, A." box="[1023,1247,854,880]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
appear especially curved. The postprotocrista in the upper molars was described as curved in the species description, but this does not appear to be especially the case in any specimens of
|
||
<taxonomicName id="4C4C4D68FF56D58FFCC0779FFBDCFC1E" box="[840,1036,984,1010]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFCC0779FFBDCFC1E" box="[840,1036,984,1010]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
examined, including the two figured partial maxillae (QM F5239 and QM F4719;
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFA9777BFFCB5FBD8" author="Bartholomai, A." pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai 1973</bibRefCitation>
|
||
, pl. 20). The curvature of the postparacrista was not mentioned, though this was observed to regularly vary even within individuals of species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFA85701DFA74FB99" authorityName="Owen" authorityYear="1874" box="[1293,1444,1115,1141]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFA85701DFA74FB99" box="[1293,1444,1115,1141]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
, including a specimen figured by
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFB13703DFAA2FB79" author="Bartholomai, A." box="[1179,1394,1147,1173]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973</bibRefCitation>
|
||
; QM F4719, pl. 20).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFCD270FBFB9AFA76" blockId="186.[810,1452,171,2052]" pageId="186" pageNumber="187">
|
||
Dentary height is quite variable within species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFCA2709BFC11FB1B" authorityName="Owen" authorityYear="1874" box="[810,961,1245,1271]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFCA2709BFC11FB1B" box="[810,961,1245,1271]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13772A6EFF56D58FFC47709BFBFFFB1B" box="[975,1071,1245,1271]" captionStart="FIGURE 110" captionStartId="194.[136,235,1036,1060]" captionTargetBox="[171,1416,176,1007]" captionTargetId="figure-27@194.[164,1423,169,1014]" captionTargetPageId="194" captionText="FIGURE 110. scatterplot of lower (mandibular) diastema length versus height of the mandibular corpus beneath m1 in specimens of P.anak, P. mamkurra sp. nov., P. viator sp. nov., P. dawsonae sp. nov., P. snewini, P. otibandus, P. tumbuna, and four extrageneric taxa." figureDoi="http://doi.org/10.5281/zenodo.10994016" httpUri="https://zenodo.org/record/10994016/files/figure.png" pageId="186" pageNumber="187">Fig. 110</figureCitation>
|
||
). Within the sample of
|
||
<taxonomicName id="4C4C4D68FF56D58FFAA47098FAACFB1B" authorityName=", Owen" authorityYear="1874" box="[1324,1404,1245,1271]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF56D58FFAA47098FAACFB1B" box="[1324,1404,1245,1271]" italics="true" pageId="186" pageNumber="187">P. anak</emphasis>
|
||
</taxonomicName>
|
||
, the spread in the measurements of this feature encompasses that of all Pliocene species of
|
||
<taxonomicName id="4C4C4D68FF56D58FFB397158FA98FAD4" authorityName="Owen" authorityYear="1874" box="[1201,1352,1310,1336]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF56D58FFB397158FA98FAD4" box="[1201,1352,1310,1336]" italics="true" pageId="186" pageNumber="187">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
, so the relatively small differences in dentary height between the small samples of
|
||
<taxonomicName id="4C4C4D68FF56D58FFC7A7126FB67FA95" box="[1010,1207,1375,1401]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFC7A7126FB67FA95" box="[1010,1207,1375,1401]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF56D58FFB787126FAABFA95" box="[1264,1403,1375,1401]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFB787126FAABFA95" box="[1264,1403,1375,1401]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
can be effectively discounted.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF56D58FFCD271E7FBB5F7E8" blockId="186.[810,1452,171,2052]" pageId="186" pageNumber="187">
|
||
It is unclear what
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFBCD71E6FAFBFA56" author="Bartholomai, A." box="[1093,1323,1440,1466]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
meant by ‘swollen anterolingual protolophid base’ (p. 352), as no swelling in this part of the lower molars of the
|
||
<typeStatus id="54F78849FF56D58FFAC171A7FA7BFA17" box="[1353,1451,1505,1531]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
and referred specimens of
|
||
<taxonomicName id="4C4C4D68FF56D58FFBC27245FADBF9F0" box="[1098,1291,1538,1564]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFBC27245FADBF9F0" box="[1098,1291,1538,1564]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
was identified in this study. Unpublished research suggests that this trait was not evident in any specimen (Lyndall Dawson, pers. comm. 2001). It is possible that the use of the term protolophid by
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFC5772C2FB11F972" author="Bartholomai, A." box="[991,1217,1668,1694]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
was the result of a
|
||
<emphasis id="B938EAF9FF56D58FFCA272E3FC13F953" box="[810,963,1701,1727]" italics="true" pageId="186" pageNumber="187">lapsus calami</emphasis>
|
||
and that the author instead intended to refer to the anterolingual base of the protoloph of the upper molars, as this area of the upper molar is indeed distinctly convex, with the degree of swelling decreasing toward the posterior of the molar row. Regardless, this trait is shared by specimens of
|
||
<taxonomicName id="4C4C4D68FF56D58FFC57730EFBB6F88D" box="[991,1126,1863,1889]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFC57730EFBB6F88D" box="[991,1126,1863,1889]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, as the
|
||
<typeStatus id="54F78849FF56D58FFAA67301FA40F88D" box="[1326,1424,1863,1889]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF56D58FFCA2732FFC3BF86E" box="[810,1003,1896,1922]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFCA2732FFC3BF86E" box="[810,1003,1896,1922]" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
, a dentary, has no features to confidently distinguish it from the type material of
|
||
<taxonomicName id="4C4C4D68FF56D58FFB6173CFFAA3F84E" box="[1257,1395,1928,1954]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFB6173CFFAA3F84E" box="[1257,1395,1928,1954]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
, and nor indeed does any other referred element, we find
|
||
<taxonomicName id="4C4C4D68FF56D58FFA1F73ECFB78F808" authority="Bartholomai, 1973" authorityName="Bartholomai" authorityYear="1973" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF56D58FFA1F73ECFC02F808" italics="true" pageId="186" pageNumber="187">P. chinchillaensis</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFC5F738CFB78F808" author="Bartholomai, A." box="[983,1192,1994,2020]" pageId="186" pageNumber="187" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
to be a junior synonym of
|
||
<taxonomicName id="4C4C4D68FF56D58FFCC073ADFBB0F7E8" authority="Plane, 1967" authorityName="Plane" authorityYear="1967" box="[840,1120,2026,2052]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF56D58FFCC073ADFC02F7E8" box="[840,978,2026,2052]" italics="true" pageId="186" pageNumber="187">P. otibandus</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF56D58FFC5173ACFBB0F7E8" author="Plane, M. D." box="[985,1120,2026,2052]" pageId="186" pageNumber="187" pagination="1 - 64" refId="ref153431" refString="Plane, M. D. (1967) Stratigraphy and vertebrate fauna of the Otibanda Formation, New Guinea. Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, Australia, 86, 1 - 64." type="journal article" year="1967">Plane, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |