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<document id="38BCF892A64A83DF34BE2947010FBA09" ID-CLB-Dataset="55597" ID-DOI="10.1093/zoolinnean/zlab107" ID-GBIF-Dataset="6e437bd5-bce3-4f86-97f6-8d40fff86b84" ID-ISSN="0024-4082" ID-Zenodo-Dep="6530771" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652084627399" checkinUser="plazi" docAuthor="Karameta, Emmanouela, Lymberakis, Petros, Grillitsch, Heinz, Ilgaz, Çetin, Avci, Aziz, Kumlutaş, Yusuf, Candan, Kamil, Wagner, Philipp, Sfenthourakis, Spyros, Pafilis, Panayiotis &amp; Poulakakis, Nikos" docDate="2022" docId="03E587DFFFEEFFC4308F0CC8FA3EFC3D" docLanguage="en" docName="zlab107.pdf" docOrigin="Zoological Journal of the Linnean Society 195 (1)" docSource="https://academic.oup.com/zoolinnean/article/195/1/195/6517658" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Laudakia stellio S. S." docType="treatment" docVersion="7" lastPageNumber="212" masterDocId="FFDCFFA7FFFFFFD530650A38FFAEFFFF" masterDocTitle="The story of a rock-star: multilocus phylogeny and species delimitation in the starred or roughtail rock agama, Laudakia stellio (Reptilia: Agamidae)" masterLastPageNumber="219" masterPageNumber="195" pageNumber="212" updateTime="1699329951700" updateUser="ExternalLinkService">
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<mods:title id="8F83B9EB3309294CE83773C32266AB62">The story of a rock-star: multilocus phylogeny and species delimitation in the starred or roughtail rock agama, Laudakia stellio (Reptilia: Agamidae)</mods:title>
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<treatment id="03E587DFFFEEFFC4308F0CC8FA3EFC3D" ID-DOI="http://doi.org/10.5281/zenodo.6536712" ID-GBIF-Taxon="195298168" ID-Zenodo-Dep="6536712" LSID="urn:lsid:plazi:treatment:03E587DFFFEEFFC4308F0CC8FA3EFC3D" httpUri="http://treatment.plazi.org/id/03E587DFFFEEFFC4308F0CC8FA3EFC3D" lastPageNumber="212" pageId="17" pageNumber="212">
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<paragraph id="8BF336C9FFEEFFC4308F0CC8FD6DF8F8" blockId="17.[234,707,1775,1800]" box="[234,707,1775,1800]" pageId="17" pageNumber="212">
<heading id="D0BB81A5FFEEFFC4308F0CC8FD6DF8F8" box="[234,707,1775,1800]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="212" reason="2">
<taxonomicName id="4C4C4D4AFFEEFFC4308F0CC8FD6DF8F8" authority="(LINNAEUS, 1758) S. S." authorityName="S. S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[234,707,1775,1800]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="212" phylum="Chordata" rank="species" species="stellio">
<emphasis id="B938EADBFFEEFFC4308F0CC8FE6FF8F8" box="[234,449,1776,1799]" italics="true" pageId="17" pageNumber="212">LAUDAKIA STELLIO</emphasis>
(LINNAEUS, 1758)
<emphasis id="B938EADBFFEEFFC432FF0CCBFD6DF8F8" box="[666,707,1776,1799]" italics="true" pageId="17" pageNumber="212">S.S.</emphasis>
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<emphasis id="B938EADBFFEEFFC430C60D00FE49F8B1" box="[163,487,1848,1870]" italics="true" pageId="17" pageNumber="212">
Included taxa:
<taxonomicName id="4C4C4D4AFFEEFFC4313E0D00FE63F8B2" authorityName="Gray" authorityYear="1845" box="[347,461,1848,1869]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="212" phylum="Chordata" rank="genus">Laudakia</taxonomicName>
s.
</emphasis>
<taxonomicName id="4C4C4D4AFFEEFFC4318A0D00FCA8F8B1" authority="(Linnaeus, 1758)" authorityName="S. S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[495,774,1848,1870]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="212" phylum="Chordata" rank="species" species="stellio">
<emphasis id="B938EADBFFEEFFC4318A0D00FD98F8B2" box="[495,566,1848,1869]" italics="true" pageId="17" pageNumber="212">stellio</emphasis>
(Linnaeus, 1758)
</taxonomicName>
,
<emphasis id="B938EADBFFEEFFC430C60D6FFEB0F894" box="[163,286,1878,1900]" italics="true" pageId="17" pageNumber="212">L. s. daani</emphasis>
(
<bibRefCitation id="EFDD4B38FFEEFFC431550D6EFD8BF893" author="Beutler A &amp; Fror E" box="[304,549,1878,1900]" pageId="17" pageNumber="212" pagination="255 - 290" refId="ref15179" refString="Beutler A, Fror E. 1980. Die Amphibien und Reptilien der Nordkykladen (Griechenland). Mitteilungen der Zoologischen Gesellschaft Braunau 3: 255 - 290." type="journal article" year="1980">Beutler &amp; Frör, 1980</bibRefCitation>
) and the subclades from the western Taurus,
<emphasis id="B938EADBFFEEFFC4343D0AFFFB64FF23" box="[1112,1226,199,220]" italics="true" pageId="17" pageNumber="212">SymiKaş</emphasis>
,
<collectingCountry id="F35B7659FFEEFFC434B10AFEFA96FF23" box="[1236,1336,198,220]" name="Turkey" pageId="17" pageNumber="212">Anatolia</collectingCountry>
,
<collectingCountry id="F35B7659FFEEFFC435250AFEFA0DFF23" box="[1344,1443,198,220]" name="Lebanon" pageId="17" pageNumber="212">Lebanon</collectingCountry>
and
<collectingCountry id="F35B7659FFEEFFC433090ADDFC1FFF05" box="[876,945,229,250]" name="Turkey" pageId="17" pageNumber="212">Hatay</collectingCountry>
, as shown in this study (
<figureCitation id="13772A4CFFEEFFC434A90ADDFABFFF05" box="[1228,1297,229,251]" captionStart="Figure 2" captionStartId="7.[163,241,1675,1697]" captionTargetBox="[179,762,196,1636]" captionTargetId="figure-539@7.[421,662,735,1006]" captionTargetPageId="7" captionText="Figure 2. Phylogenetic tree based on mtDNA (ND4-tRNAs and 16S rRNA). Bayesian posterior probabilities (PP) and maximum likelihood bootstrap support (bs) values are represented in the form PP/bs above or beside nodes. (Agama spp. contains Agama agama, A. boensis, A. bottega, A. boueti, A. boulengeri, A. impalearis, A. planices, A. sankaranica and A. spinosa)." figureDoi="http://doi.org/10.5281/zenodo.6530775" httpUri="https://zenodo.org/record/6530775/files/figure.png" pageId="17" pageNumber="212">Figs 2</figureCitation>
,
<figureCitation id="13772A4CFFEEFFC4357B0ADDFA82FF05" box="[1310,1324,229,250]" captionStart="Figure 4" captionStartId="9.[163,242,1649,1671]" captionTargetBox="[229,1373,197,1607]" captionTargetId="figure-91@9.[227,1379,195,1609]" captionTargetPageId="9" captionText="Figure 4. Phylogenetic tree based on the concatenated dataset (mtDNA &amp; nuDNA). Bayesian posterior probabilities (PP) and maximum likelihood bootstrap support (bs) values are represented in the form PP/bs above or beside nodes. (Agama spp. contains Agama agama, A. boensis, A. bottega, A. boueti, A. boulengeri, A. impalearis, A. planices, A. sankaranica and A. spinosa)." figureDoi="http://doi.org/10.5281/zenodo.6530779" httpUri="https://zenodo.org/record/6530779/files/figure.png" pageId="17" pageNumber="212">4</figureCitation>
).
</paragraph>
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<emphasis id="B938EADBFFEEFFC4335E0B1CFC43FEC6" box="[827,1005,292,313]" italics="true" pageId="17" pageNumber="212">Type specimen:</emphasis>
<taxonomicName id="4C4C4D4AFFEEFFC434610B1CFCD1FEA7" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="212" phylum="Chordata" rank="subSpecies" species="stellio" subSpecies="stellio">
<emphasis id="B938EADBFFEEFFC434610B1CFAB5FEC6" box="[1028,1307,292,313]" italics="true" pageId="17" pageNumber="212">Laudakia stellio stellio</emphasis>
(Linnaeus, 1758)
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,
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ZFMK 2063 (
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),
<bibRefCitation id="EFDD4B38FFEEFFC434CD0B7AFA0DFEA7" author="Crochet PA &amp; Lymberakis P &amp; Werner YL" box="[1192,1443,322,344]" pageId="17" pageNumber="212" pagination="461 - 471" refId="ref15632" refString="Crochet PA, Lymberakis P, Werner YL. 2006. The type specimens of Laudakia stellio (Linnaeus) (Reptilia: Agamidae) and its subspecies. Journal of Natural History 40: 461 - 471." type="journal article" year="2006">
Crochet
<emphasis id="B938EADBFFEEFFC435750B7BFAE3FEA8" box="[1296,1357,322,344]" italics="true" pageId="17" pageNumber="212">et al.</emphasis>
(2006)
</bibRefCitation>
(
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,
<collectingMunicipality id="6B97ACB3FFEEFFC4335E0BB8FCD7FE6A" box="[827,889,384,405]" pageId="17" pageNumber="212">Bonn</collectingMunicipality>
,
<collectingCountry id="F35B7659FFEEFFC433E10BB8FC5CFE69" box="[900,1010,384,406]" name="Germany" pageId="17" pageNumber="212">Germany</collectingCountry>
)
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.
</paragraph>
<paragraph id="8BF336C9FFEEFFC4335E0B87FAEBFE2C" blockId="17.[827,1349,446,468]" box="[827,1349,446,468]" pageId="17" pageNumber="212">
<materialsCitation id="3B243C94FFEEFFC4335E0B87FAEFFE2C" ID-GBIF-Occurrence="3765334305" box="[827,1345,446,468]" country="Greece" location="Delos Island" municipality="Cyclades" pageId="17" pageNumber="212" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EADBFFEEFFC4335E0B87FC7EFE2C" box="[827,976,446,468]" italics="true" pageId="17" pageNumber="212">
<typeStatus id="54F7886BFFEEFFC4335E0B87FCDEFE2B" box="[827,880,447,468]" pageId="17" pageNumber="212">Type</typeStatus>
locality:
</emphasis>
<location id="8E936012FFEEFFC433810B86FBDAFE2B" LSID="urn:lsid:plazi:treatment:03E587DFFFEEFFC4308F0CC8FA3EFC3D:8E936012FFEEFFC433810B86FBDAFE2B" box="[996,1140,446,468]" country="Greece" municipality="Cyclades" name="Delos Island" pageId="17" pageNumber="212">Delos Island</location>
,
<collectingMunicipality id="6B97ACB3FFEEFFC4341A0B86FB48FE2B" box="[1151,1254,446,468]" pageId="17" pageNumber="212">Cyclades</collectingMunicipality>
,
<collectingCountry id="F35B7659FFEEFFC434950B86FAEFFE2C" box="[1264,1345,446,467]" name="Greece" pageId="17" pageNumber="212">Greece</collectingCountry>
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.
</paragraph>
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<emphasis id="B938EADBFFEEFFC4335E0BC5FC79FDED" box="[827,983,509,530]" italics="true" pageId="17" pageNumber="212">Distribution:</emphasis>
All localities from
<collectingCountry id="F35B7659FFEEFFC434A90BC5FAB0FDED" box="[1228,1310,509,530]" name="Greece" pageId="17" pageNumber="212">Greece</collectingCountry>
(Cyclades,
<collectingRegion id="4988F82BFFEEFFC4335E0824FCD3FDCE" box="[827,893,540,561]" country="Greece" name="Kriti" pageId="17" pageNumber="212">Crete</collectingRegion>
, Thessaloniki, Corfu and East
<collectingRegion id="4988F82BFFEEFFC4348D0824FA90FDCE" box="[1256,1342,540,561]" country="Greece" name="Voreio Aigaio" pageId="17" pageNumber="212">Aegean</collectingRegion>
islands) and
<collectingCountry id="F35B7659FFEEFFC4330C0802FC17FDAF" box="[873,953,570,592]" name="Turkey" pageId="17" pageNumber="212">Turkey</collectingCountry>
, north-western
<collectingCountry id="F35B7659FFEEFFC434010802FB0EFDAF" box="[1124,1184,570,592]" name="Syria" pageId="17" pageNumber="212">Syria</collectingCountry>
and northern
<collectingCountry id="F35B7659FFEEFFC435590802FA0FFDAF" box="[1340,1441,570,592]" name="Lebanon" pageId="17" pageNumber="212">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566542FFEEFFC4335E0841FA3EFC3D" pageId="17" pageNumber="212" type="discussion">
<paragraph id="8BF336C9FFEEFFC4335E0841FA3EFC3D" blockId="17.[827,1443,633,962]" pageId="17" pageNumber="212">
<emphasis id="B938EADBFFEEFFC4335E0841FC09FD71" box="[827,935,633,654]" italics="true" pageId="17" pageNumber="212">Remarks:</emphasis>
According to the principle of priority of the ICZN, this distinct genetic lineage should be named
<taxonomicName id="4C4C4D4AFFEEFFC4335E088EFC2CFD34" authorityName="S. S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[827,898,694,715]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="212" phylum="Chordata" rank="species" species="stellio">
<emphasis id="B938EADBFFEEFFC4335E088EFC2CFD34" box="[827,898,694,715]" italics="true" pageId="17" pageNumber="212">stellio</emphasis>
</taxonomicName>
because that is the oldest available name. It includes already described morphological subspecies and other cryptic lineages of Clade 2, revealed by this study: the nominal
<emphasis id="B938EADBFFEEFFC43478092BFB39FCD8" box="[1053,1175,786,808]" italics="true" pageId="17" pageNumber="212">
L.
<taxonomicName id="4C4C4D4AFFEEFFC4345E092BFB39FCD8" authorityName="S. S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1083,1175,786,808]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="215" phylum="Chordata" rank="species" species="stellio">s. stellio</taxonomicName>
</emphasis>
and its sister
<emphasis id="B938EADBFFEEFFC4355A092AFA0DFCD8" box="[1343,1443,786,807]" italics="true" pageId="17" pageNumber="212">
<collectingCountry id="F35B7659FFEEFFC4355A092AFA0DFCD8" box="[1343,1443,786,807]" name="Turkey" pageId="17" pageNumber="212">Anatolia</collectingCountry>
</emphasis>
subclade,
<emphasis id="B938EADBFFEEFFC433CD090AFBB4FCB9" box="[936,1050,817,839]" italics="true" pageId="17" pageNumber="212">L. s. daani</emphasis>
and its sister
<emphasis id="B938EADBFFEEFFC434D2090AFA0DFCB8" box="[1207,1443,817,839]" italics="true" pageId="17" pageNumber="212">W TaurusSymiKaş</emphasis>
subclades, as well as the
<emphasis id="B938EADBFFEEFFC4342B0968FB01FC9A" box="[1102,1199,848,869]" italics="true" pageId="17" pageNumber="212">
<collectingCountry id="F35B7659FFEEFFC4342B0968FB01FC9A" box="[1102,1199,848,869]" name="Lebanon" pageId="17" pageNumber="212">Lebanon</collectingCountry>
</emphasis>
and
<emphasis id="B938EADBFFEEFFC434800968FA84FC9A" box="[1253,1322,848,869]" italics="true" pageId="17" pageNumber="212">
<collectingCountry id="F35B7659FFEEFFC434800968FA84FC9A" box="[1253,1322,848,869]" name="Turkey" pageId="17" pageNumber="212">Hatay</collectingCountry>
</emphasis>
subclades. The subspecific status of
<emphasis id="B938EADBFFEEFFC434380957FB78FC7C" box="[1117,1238,878,900]" italics="true" pageId="17" pageNumber="212">
L.
<taxonomicName id="4C4C4D4AFFEEFFC4341E0957FB78FC7C" authorityName="S. S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1147,1238,878,900]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="215" phylum="Chordata" rank="species" species="stellio">s. stellio</taxonomicName>
</emphasis>
and
<emphasis id="B938EADBFFEEFFC435750957FA26FC7C" box="[1296,1416,878,900]" italics="true" pageId="17" pageNumber="212">L. s. daani</emphasis>
is retained, while the elevation of the
<emphasis id="B938EADBFFEEFFC434A309B5FA89FC5D" box="[1222,1319,909,930]" italics="true" pageId="17" pageNumber="212">
<collectingCountry id="F35B7659FFEEFFC434A309B5FA89FC5D" box="[1222,1319,909,930]" name="Lebanon" pageId="17" pageNumber="212">Lebanon</collectingCountry>
</emphasis>
and
<emphasis id="B938EADBFFEEFFC4353809B6FA0CFC5C" box="[1373,1442,910,931]" italics="true" pageId="17" pageNumber="212">
<collectingCountry id="F35B7659FFEEFFC4353809B6FA0CFC5C" box="[1373,1442,910,931]" name="Turkey" pageId="17" pageNumber="212">Hatay</collectingCountry>
</emphasis>
lineages to the subspecies level is under preparation.
</paragraph>
</subSubSection>
</treatment>
</document>