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<document id="645AA1CE9D02A552B7A4C51E10E7DEDC" ID-DOI="10.11646/zootaxa.2539.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10094211" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629684766228" checkinUser="felipe" docAuthor="Kokshoorn, Bas &amp; Gittenberger, Edmund" docDate="2010" docId="03E4E904CE7BEC517EEEFC3FC233CFCC" docLanguage="en" docName="zt02539p062.pdf" docOrigin="Zootaxa 2539 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2539.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Abida secale subsp. peteri Kokshoorn &amp; Gittenberger 2010, subspec. nov." docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="FFDD917CCE6FEC457E79FF9BC701C838" masterDocTitle="Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539" masterLastPageNumber="62" masterPageNumber="1" pageNumber="21" updateTime="1699566659466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E29D9CFDD6CCD1EB20E65213B8034BFF">Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539</mods:title>
</mods:titleInfo>
<mods:name id="206A89875002A2A26BCFEF5D49C82A92" type="personal">
<mods:role id="C8FB05A37E4C61EDFB65A7AC9CC20C27">
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<mods:namePart id="D961484134AFBF2EB928EDD58D95C894">Kokshoorn, Bas</mods:namePart>
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<mods:namePart id="93FD4D08825D24053B50FD54D9EE7BFF">Gittenberger, Edmund</mods:namePart>
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<mods:typeOfResource id="82EF300D98D6EF85D6236D19A60AD2CA">text</mods:typeOfResource>
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<mods:title id="D5DB2A6A02FD98E75E9990E0327D2B0E">Zootaxa</mods:title>
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<mods:date id="493682254D3EEB902E7DF84DE33C68C5">2010</mods:date>
<mods:detail id="E6348A5032CBAEE54477B83901513A0B" type="pubDate">
<mods:number id="E77C8B08DAFA524971B4FDA86EC3EF00">2010-07-16</mods:number>
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<mods:detail id="47E1DD53D30DAABD63B7DF8FBB7847B4" type="volume">
<mods:number id="1403C29C5F37CD375A2B77FA6F0C38E3">2539</mods:number>
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<mods:number id="4C738FE3CAD758032EEBBC2DFE43398F">1</mods:number>
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<mods:start id="30B04616D9E16967EAD49CCA49BC817C">1</mods:start>
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<mods:url id="3712C898C337AF01A1C5954BE0DC8656">https://biotaxa.org/Zootaxa/article/view/zootaxa.2539.1.1</mods:url>
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<mods:classification id="2D69226EEA5BF56681970B561F29E007">journal article</mods:classification>
<mods:identifier id="D4E61BFE8B9023C4ABD386DDAF411A8A" type="DOI">10.11646/zootaxa.2539.1.1</mods:identifier>
<mods:identifier id="2938CE50989C4051329DDB0577C0171B" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="68E50C8828BD9ECBC81C50425B26BDBD" type="Zenodo-Dep">10094211</mods:identifier>
</mods:mods>
<treatment id="03E4E904CE7BEC517EEEFC3FC233CFCC" LSID="urn:lsid:plazi:treatment:03E4E904CE7BEC517EEEFC3FC233CFCC" httpUri="http://treatment.plazi.org/id/03E4E904CE7BEC517EEEFC3FC233CFCC" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="C3570B99CE7BEC517EEEFC3FC5E8CB85" box="[151,745,931,958]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BF25812CE7BEC517EEEFC3FC5E8CB85" blockId="20.[151,1436,931,2037]" box="[151,745,931,958]" pageId="20" pageNumber="21">
<heading id="D0BAEF7ECE7BEC517EEEFC3FC5E8CB85" bold="true" box="[151,745,931,958]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B9398400CE7BEC517EEEFC3FC5E8CB85" bold="true" box="[151,745,931,958]" pageId="20" pageNumber="21">
<taxonomicName id="4C4D2391CE7BEC517EEEFC3FC67BCB86" ID-CoL="5DDYS" authority="Kokshoorn &amp; Gittenberger, 2010" authorityName="Kokshoorn &amp; Gittenberger" authorityYear="2010" box="[151,378,932,958]" class="Gastropoda" family="Chondrinidae" genus="Abida" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="subSpecies" species="secale" status="subsp. nov." subSpecies="peteri">
<emphasis id="B9398400CE7BEC517EEEFC3FC67BCB86" bold="true" box="[151,378,932,958]" italics="true" pageId="20" pageNumber="21">Abida secale peteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A397BCE7BEC517FFBFC38C524CB85" box="[386,549,931,957]" pageId="20" pageNumber="21" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
(pl. 7 figs A–C)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3570B99CE7BEC517EEEFC5DC233CFCC" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BF25812CE7BEC517EEEFC5DC4BBCC6C" blockId="20.[151,1436,931,2037]" pageId="20" pageNumber="21">
Type series.—
<materialsCitation id="3B25524FCE7BEC517F3AFC5DC3CFCBD8" box="[323,1230,966,992]" collectionCode="RMNH" country="Spain" county="Pedraforca" location="Lerida" municipality="Sierra del Cadi" pageId="20" pageNumber="21" specimenCode="RMNH 109890" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F35A1882CE7BEC517F3AFC5DC689CBD8" box="[323,392,966,992]" name="Spain" pageId="20" pageNumber="21">Spain</collectingCountry>
,
<location id="8E920EC9CE7BEC517FEEFC5DC6E7CBD8" LSID="urn:lsid:plazi:treatment:03E4E904CE7BEC517EEEFC3FC233CFCC:8E920EC9CE7BEC517FEEFC5DC6E7CBD8" box="[407,486,966,992]" country="Spain" county="Pedraforca" municipality="Sierra del Cadi" name="Lerida" pageId="20" pageNumber="21">Lérida</location>
,
<collectingMunicipality id="6B96C268CE7BEC517F8CFC5DC5AFCBD8" box="[501,686,966,992]" pageId="20" pageNumber="21">Sierra del Cadí</collectingMunicipality>
,
<collectingCounty id="6293209ECE7BEC517CC4FC5DC43DCBD8" box="[701,828,966,992]" pageId="20" pageNumber="21">Pedraforca</collectingCounty>
mtn.
<typeStatus id="54F6E6B0CE7BEC517DFCFC5DC4F3CBD8" box="[901,1010,966,992]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBEBF069CE7BEC517A7DFC5DC3C6CBD8" box="[1028,1223,966,992]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21" type="Museum">RMNH 109890</specimenCode>
)
</materialsCitation>
:
<materialsCitation id="3B25524FCE7BEC517AA4FC5DC4B7CC6C" collectingDate="2004-08-06" collectionCode="RMNH" collectorName="B. Kokshoorn &amp; P. Lindenburg" country="Spain" elevation="2300" pageId="20" pageNumber="21" specimenCode="RMNH 99135, RMNH 54961, RMNH 99136" specimenCount="1" typeStatus="paratype">
<quantity id="4CB5F5F7CE7BEC517AA4FC5DC241CBD8" box="[1245,1344,966,992]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="21" unit="m" value="2300.0">
<elevation id="0060BF21CE7BEC517AA4FC5DC241CBD8" box="[1245,1344,966,992]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="21" unit="m" value="2300.0">2,300 m</elevation>
</quantity>
alt.,
<collectorName id="26B83DC4CE7BEC517BF8FC5DC618CC3F" pageId="20" pageNumber="21">B. Kokshoorn</collectorName>
&amp;
<collectorName id="26B83DC4CE7BEC517F3BFC76C6E6CC3F" box="[322,487,1005,1031]" pageId="20" pageNumber="21">P. Lindenburg</collectorName>
leg.,
<date id="FFF37ED2CE7BEC517C55FC76C5C3CC3F" box="[556,706,1005,1031]" pageId="20" pageNumber="21" value="2004-08-06">
<collectingDate id="EFB7873ACE7BEC517C55FC76C5C3CC3F" box="[556,706,1005,1031]" pageId="20" pageNumber="21" value="2004-08-06">06-viii-2004</collectingDate>
</date>
.
<typeStatus id="54F6E6B0CE7BEC517CB7FC76C445CC3F" box="[718,836,1005,1031]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
: type locality (
<specimenCode id="DBEBF069CE7BEC517D8CFC76C3A4CC3F" box="[1013,1189,1005,1031]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21" type="Museum">RMNH 99135</specimenCode>
/23);
<quantity id="4CB5F5F7CE7BEC517A98FC76C242CC3F" box="[1249,1347,1005,1031]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="21" unit="m" value="2250.0">
<elevation id="0060BF21CE7BEC517A98FC76C242CC3F" box="[1249,1347,1005,1031]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="21" unit="m" value="2250.0">2,250 m</elevation>
</quantity>
alt.,
<collectorName id="26B83DC4CE7BEC517BF8FC76C618CC16" pageId="20" pageNumber="21">B. Kokshoorn</collectorName>
&amp;
<collectorName id="26B83DC4CE7BEC517F47FB8FC6E3CC16" box="[318,482,1044,1070]" pageId="20" pageNumber="21">P. Lindenburg</collectorName>
leg.,
<date id="FFF37ED2CE7BEC517C5BFB8FC5B5CC16" box="[546,692,1044,1070]" pageId="20" pageNumber="21" value="2004-08-06">
<collectingDate id="EFB7873ACE7BEC517C5BFB8FC5B5CC16" box="[546,692,1044,1070]" pageId="20" pageNumber="21" value="2004-08-06">06-viii-2004</collectingDate>
</date>
(
<specimenCode id="DBEBF069CE7BEC517CBAFB8FC473CC16" box="[707,882,1044,1070]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21" type="Museum">RMNH 54961</specimenCode>
/6); E-side, c.
<quantity id="4CB5F5F7CE7BEC517A6BFB8FC34FCC16" box="[1042,1102,1044,1070]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="50.0">
<elevation id="0060BF21CE7BEC517A6BFB8FC34FCC16" box="[1042,1102,1044,1070]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="50.0">50 m</elevation>
</quantity>
below the coll,
<quantity id="4CB5F5F7CE7BEC517B70FB8FC268CC16" box="[1289,1385,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="21" unit="m" value="2250.0">
<elevation id="0060BF21CE7BEC517B70FB8FC268CC16" box="[1289,1385,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="21" unit="m" value="2250.0">2,250 m</elevation>
</quantity>
alt.,
<collectorName id="26B83DC4CE7BEC517EEEFBA1C63ACC6C" box="[151,315,1082,1108]" pageId="20" pageNumber="21">B. Kokshoorn</collectorName>
&amp;
<collectorName id="26B83DC4CE7BEC517F18FBA1C505CC6C" box="[353,516,1082,1108]" pageId="20" pageNumber="21">P. Lindenburg</collectorName>
leg.,
<date id="FFF37ED2CE7BEC517C3FFBA1C5D9CC6C" box="[582,728,1082,1108]" pageId="20" pageNumber="21" value="2004-08-06">
<collectingDate id="EFB7873ACE7BEC517C3FFBA1C5D9CC6C" box="[582,728,1082,1108]" pageId="20" pageNumber="21" value="2004-08-06">06-viii-2004</collectingDate>
</date>
(
<specimenCode id="DBEBF069CE7BEC517C9EFBA1C496CC6C" box="[743,919,1082,1108]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21" type="Museum">RMNH 99136</specimenCode>
/5)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF25812CE7BEC517EBFFBFAC6EECC43" blockId="20.[151,1436,931,2037]" box="[198,495,1121,1147]" pageId="20" pageNumber="21">
Distribution.—See
<figureCitation id="13764497CE7BEC517FD3FBFAC6EACC43" box="[426,491,1121,1147]" captionStart="FIGURE 2" captionStartId="10.[151,255,1265,1289]" captionTargetBox="[304,1274,206,1237]" captionTargetId="graphics-271@10.[351,1270,218,1237]" captionTargetPageId="10" captionText="FIGURE 2. Distribution of Abida secale subspecies (I) in NE Spain and Andorra." figureDoi="http://doi.org/10.5281/zenodo.10094213" httpUri="https://zenodo.org/record/10094213/files/figure.png" pageId="20" pageNumber="21">fig. 2</figureCitation>
.
</paragraph>
<paragraph id="8BF25812CE7BEC517EBFFB13C39FCD2E" blockId="20.[151,1436,931,2037]" pageId="20" pageNumber="21">
Description (n=35).—Shell slender spindle-shaped, with 7 3/8–8 1/8 slightly inflated whorls, sculptured with regularly placed, vague axial ribs. The sculpture is often partially faded. Body whorl slightly flattened and more or less narrowed, without apparent keel. Palatal side of the aperture slightly protruding in the middle (in lateral view). Umbilicus widely open. Shell height,
<quantity id="4CB5F5F7CE7BEC517D65FB67C4ADCD2E" box="[796,940,1276,1302]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="7.3" metricValueMin="5.1" pageId="20" pageNumber="21" unit="mm" value="6.199999999999999" valueMax="7.3" valueMin="5.1">5.1–7.3 mm</quantity>
; width, 2.0–
<quantity id="4CB5F5F7CE7BEC517A44FB67C39ACD2E" box="[1085,1179,1276,1302]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="21" unit="mm" value="2.2">2.2 mm</quantity>
.
</paragraph>
<paragraph id="8BF25812CE7BEC517EBFFAB8C287CDEF" blockId="20.[151,1436,931,2037]" pageId="20" pageNumber="21">Apertural lip thickened, strongly reflected and discontinous at the parietal side or connected by a thin callus. Angularis not connected with spiralis. Subangularis absent. Columellaris not reaching beyond columella, clearly broader than subcolumellaris and parietalis. Infracolumellaris like columellaris but less prominent. Palatalis inferior and palatalis superior clearly indented and almost reaching the edge of the aperture. At the anterodorsal palatal center a weak suprapalatalis and infrapalatalis are present. A suturalis is usually absent.</paragraph>
<paragraph id="8BF25812CE7BEC517EBFFA7FC667CF0B" blockId="20.[151,1436,931,2037]" pageId="20" pageNumber="21">
Differentiation.—This subspecies shares character states with both
<emphasis id="B9398400CE7BEC517D9DFA7FC390CDC5" box="[996,1169,1508,1533]" italics="true" pageId="20" pageNumber="21">A. s. lilietensis</emphasis>
and
<emphasis id="B9398400CE7BEC517AB4FA7FC295CDC5" box="[1229,1428,1508,1533]" italics="true" pageId="20" pageNumber="21">A. s. brauniopsis</emphasis>
. These two subspecies occur at the foot and along the flanks of the mountain, i.e.
<emphasis id="B9398400CE7BEC517A03F990C244CE1C" box="[1146,1349,1547,1572]" italics="true" pageId="20" pageNumber="21">A. s. brauniopsis</emphasis>
on the south- and westside, and
<emphasis id="B9398400CE7BEC517FB8F9AAC56CCE72" box="[449,621,1585,1610]" italics="true" pageId="20" pageNumber="21">A. s. lilietensis</emphasis>
on the NE-flank. The pronounced columellaris, widely open umbilicus and the strongly reflected apertural lip point to a relation with
<emphasis id="B9398400CE7BEC517DDAF9C3C369CE49" box="[931,1128,1624,1649]" italics="true" pageId="20" pageNumber="21">A. s. brauniopsis</emphasis>
. However, along the eastflank of the mountain, at altitudes from
<quantity id="4CB5F5F7CE7BEC517C00F9E5C448CEA0" box="[633,841,1662,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="2.1" metricValueMin="1.65" pageId="20" pageNumber="21" unit="m" value="1875.0" valueMax="2100.0" valueMin="1650.0">1,650 to 2,100 m</quantity>
, shells that are clearly intermediate between
<emphasis id="B9398400CE7BEC517B1FF9E4C7DACE86" italics="true" pageId="20" pageNumber="21">A. s. peteri</emphasis>
and
<emphasis id="B9398400CE7BEC517F6DF93EC6BECE86" box="[276,447,1701,1726]" italics="true" pageId="20" pageNumber="21">A. s. lilietensis</emphasis>
are found. These specimens have a discontinuous apertural lip and the more slender shape that links these two taxa.
<emphasis id="B9398400CE7BEC517C69F957C591CEDD" box="[528,656,1740,1765]" italics="true" pageId="20" pageNumber="21">A. s. peteri</emphasis>
resembles
<emphasis id="B9398400CE7BEC517D6FF957C4C1CEDD" box="[790,960,1740,1765]" italics="true" pageId="20" pageNumber="21">A. s. cadiensis</emphasis>
and
<emphasis id="B9398400CE7BEC517D83F957C38ACEDD" box="[1018,1163,1740,1765]" italics="true" pageId="20" pageNumber="21">A. s. merijni</emphasis>
, but differs in that it is somewhat smaller and less conical and by its very fine striae that are more pronounced in both
<emphasis id="B9398400CE7BEC517A8DF968C29DCF34" box="[1268,1436,1779,1804]" italics="true" pageId="20" pageNumber="21">A. s. cadiensis</emphasis>
and
<emphasis id="B9398400CE7BEC517EB1F882C656CF0A" box="[200,343,1817,1842]" italics="true" pageId="20" pageNumber="21">A. s. merijni</emphasis>
.
</paragraph>
<paragraph id="8BF25812CE7BEC517EBFF8DBC609CFF6" blockId="20.[151,1436,931,2037]" pageId="20" pageNumber="21">
Derivatio nominis.—This subspecies is named after Mr.
<collectingRegion id="498996F0CE7BEC517D1AF8DBC49ECF62" box="[867,927,1856,1882]" country="Montserrat" name="Saint Peter" pageId="20" pageNumber="21">Peter</collectingRegion>
Lindenburg. Without his help and support, the first fieldwork period of the first author would not have been possible. Together collecting trips to the peaks of the Pedraforca mtn and the Torre del Cadí mtn were made, where material of this new subspecies was collected.
</paragraph>
<paragraph id="8BF25812CE7BEC517EBFF840C233CFCC" blockId="20.[151,1436,931,2037]" box="[198,1330,2010,2037]" pageId="20" pageNumber="21">
<materialsCitation id="3B25524FCE7BEC517EBFF840C482CFCD" accessionNumber="EU395369" box="[198,899,2011,2037]" pageId="20" pageNumber="21" specimenCount="1">
Genetic barcode.—GenBank accession numbers
<accessionNumber id="941EC5F1CE7BEC517D7DF840C482CFCD" box="[772,899,2011,2037]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU395369" pageId="20" pageNumber="21">EU395369</accessionNumber>
</materialsCitation>
–
<materialsCitation id="3B25524FCE7BEC517DE9F841C22FCFCC" accessionNumber="EU395370, EU395373, EU395378" box="[912,1326,2010,2036]" isTailEnumeration="true" pageId="20" pageNumber="21" specimenCount="1">
<accessionNumber id="941EC5F1CE7BEC517DE9F841C313CFCC" box="[912,1042,2010,2036]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU395370" pageId="20" pageNumber="21">EU395370</accessionNumber>
,
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