treatments-xml/data/03/E3/22/03E3224DFFB92834FC5CFC35FB1FA409.xml

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<mods:title id="7AC566F18C1DCA7B2CD576B8D67A85D6">The genus Parilia Wood-Mason, in Wood-Mason &amp; Alcock, 1891, with description of a new species and establishment of a new genus for P. tuberculata Sakai, 1961 (Crustacea, Brachyura, Leucosiidae)</mods:title>
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<taxonomicName id="4C4AE8D8FFB92837FC5CFC35FAE4A1C4" ID-CoL="4DMD9" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[1009,1300,989,1013]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="301" phylum="Arthropoda" rank="species" species="major">
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<emphasis id="B93E4F49FFB92837FC5CFC35FB77A1C4" bold="true" box="[1009,1159,989,1013]" italics="true" pageId="1" pageNumber="301">Parilia major</emphasis>
<bibRefCitation id="EFDBEEAAFFB92837FB3DFC35FAE4A1C4" author="Sakai T" box="[1168,1300,989,1013]" pageId="1" pageNumber="301" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
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<paragraph id="8BF5935BFFB92837FCEEFC15FA32A624" blockId="1.[835,1474,989,1045]" box="[835,1474,1021,1045]" pageId="1" pageNumber="301">
(
<figureCitation id="13718FDEFFB92837FCE6FC15FC5AA624" box="[843,938,1021,1045]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="1" pageNumber="301">Figs. 1B</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FC15FC15FC29A624" box="[952,985,1021,1045]" captionStart="Fig" captionStartId="3.[136,171,1290,1312]" captionTargetBox="[136,1474,189,1275]" captionTargetId="figure-8@3.[136,1474,189,1275]" captionTargetPageId="3" captionText="Fig. 2. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; B, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; C, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503193" httpUri="https://zenodo.org/record/4503193/files/figure.png" pageId="1" pageNumber="301">2B</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FC4BFC15FBF8A624" box="[998,1032,1021,1045]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="1" pageNumber="301">3B</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FBB8FC15FBA8A624" box="[1045,1112,1021,1045]" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="1" pageNumber="301">4C, D</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FBCBFC15FB58A624" box="[1126,1192,1021,1045]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="1" pageNumber="301">5D, E</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FB18FC15FB26A624" box="[1205,1238,1021,1045]" captionStart="Fig" captionStartId="8.[113,148,1410,1432]" captionTargetBox="[113,1451,189,1393]" captionTargetId="figure-279@8.[113,1451,189,1393]" captionText="Fig. 6. Outer view of right chela. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778841" httpUri="https://zenodo.org/record/4778841/files/figure.png" pageId="1" pageNumber="301">6B</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FB4EFC15FAD7A624" box="[1251,1319,1021,1045]" captionStart="Fig" captionStartId="9.[135,170,950,972]" captionTargetBox="[136,1474,189,933]" captionTargetId="figure-310@9.[136,1474,189,933]" captionText="Fig. 7. Male thoracic sternum and pleon. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778845" httpUri="https://zenodo.org/record/4778845/files/figure.png" pageId="1" pageNumber="301">7C, D</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FA99FC15FAB0A624" box="[1332,1344,1021,1045]" captionStart="Fig" captionStartId="10.[113,148,1974,1996]" captionTargetBox="[115,1453,190,1952]" captionTargetId="figure-8@10.[99,1459,176,1964]" captionText="Fig. 8. Gonopods. A–D, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; E–I, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; J–M, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; N–S, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India. A, E, J, N, left G1 (ventral view); O, left G1 (dorsal view); B, F, K, P, distal part of left G1 (ventral view); C, G, H, L, Q, R, distal part of left G1 (dorsal view); D, I, M, S, left G2. Scales: A, D, E, I, J, M, N, O, S = 2.0 mm; B, C, F–H, K, L, P–R = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4778849" httpUri="https://zenodo.org/record/4778849/files/figure.png" pageId="1" pageNumber="301">8</figureCitation>
E–I, 9C, D)
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<bibRefCitation id="EFDBEEAAFFB92837FC7BFBD4FB75A660" author="Sakai T" box="[982,1157,1084,1106]" pageId="1" pageNumber="301" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961: 137</bibRefCitation>
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, pl. 3 fig. 5; 
<treatmentCitation id="0AEBB54AFFB92837FABDFBD4FA4EA663" author="Sakai T" box="[1296,1470,1084,1106]" page="105" pageId="1" pageNumber="301" year="1976">
<bibRefCitation id="EFDBEEAAFFB92837FABDFBD4FA4EA663" author="Sakai T" box="[1296,1470,1084,1106]" pageId="1" pageNumber="301" pagination="728 - 738" refId="ref10963" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo. Samouelle G (1819) The Entomologist's Useful Compendium, or an Introduction to the Knowledge of the British Insects. London, Thomas Boys, Shih Y-J, Ho P-H, Chan T-Y &amp; Naruse T (2013) New records of leucosiid crabs (Decapoda, Brachyura) from Taiwan. Crustaceana, 86: 728 - 738." type="book chapter" year="1976">Sakai, 1976: 105</bibRefCitation>
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, pl. 31 fig. 1 (
<collectingCountry id="F35DD3CBFFB92837FC5FFBB1FBDEA65E" box="[1010,1070,1113,1135]" name="Japan" pageId="1" pageNumber="301">Japan</collectingCountry>
); Chen, 1989: 233, text fig. 24, pl. II, 1 (
<collectingCountry id="F35DD3CBFFB92837FCDDFB9DFC14A6BA" box="[880,996,1141,1163]" name="Philippines" pageId="1" pageNumber="301">Philippines</collectingCountry>
); Chen, 1990: 180, pl. 3E; 
<bibRefCitation id="EFDBEEAAFFB92837FB53FB9DFA79A6BA" author="Takeda M" box="[1278,1417,1141,1163]" pageId="1" pageNumber="301" refId="ref11144" refString="Takeda M (1993) Catalogue of Crab Specimens (Crustacea: Decapoda: Brachyura) (Part 1). National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993</bibRefCitation>
; 
<treatmentCitation id="0AEBB54AFFB92837FA3BFB9EFC2DA699" author="Tan CGS" page="1046" pageId="1" pageNumber="301" year="1996">
<bibRefCitation id="EFDBEEAAFFB92837FA3BFB9EFC2DA699" author="Tan CGS" pageId="1" pageNumber="301" pagination="1021 - 1058" refId="ref11174" refString="Tan CGS (1996) Leucosiidae of the Albatross expedition to the Philippines, 1907 - 1910 (Crustacea: Brachyura: Decapoda). Journal of Natural History, 30 (7): 1021 - 1058, figs. 1 - 8." type="journal article" year="1996">Tan, 1996: 1046</bibRefCitation>
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, fig. 7f–k; 
<treatmentCitation id="0AEBB54AFFB92837FBFCFB7AFB02A699" author="Ikeda H" box="[1105,1266,1170,1192]" page="86" pageId="1" pageNumber="301" year="1998">
<bibRefCitation id="EFDBEEAAFFB92837FBFCFB7AFB02A699" author="Ikeda H" box="[1105,1266,1170,1192]" pageId="1" pageNumber="301" refId="ref10359" refString="Ikeda H (1998) The Deep-Sea Crabs of Sagami Bay. Hayama Shiosai Museum, Kanagawa, Japan, 180 pp." type="book" year="1998">Ikeda, 1998: 86</bibRefCitation>
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, 87 (
<collectingCountry id="F35DD3CBFFB92837FA84FB7AFA96A699" box="[1321,1382,1170,1192]" name="Japan" pageId="1" pageNumber="301">Japan</collectingCountry>
); 
<treatmentCitation id="0AEBB54AFFB92837FAD6FB7BFC03A6F5" author="Jeng M-S &amp; Shao K &amp; Tzeng C-S &amp; Feng F-L &amp; Wu S-H" page="121" pageId="1" pageNumber="301" year="1998">
<bibRefCitation id="EFDBEEAAFFB92837FAD6FB7BFC03A6F5" author="Jeng M-S &amp; Shao K &amp; Tzeng C-S &amp; Feng F-L &amp; Wu S-H" pageId="1" pageNumber="301" refId="ref10382" refString="Jeng M-S, Shao K - T, Tzeng C-S, Feng F-L &amp; Wu S-H (1998) Survey and monitoring of natural ecological resources in the Northeast Coast National Scenic Area (IV). Northeast Coast National Scenic Area Administration, Bureau of the Ministry of Transportation and Communications, Taipei, 182 pp. [In Chinese]" type="book" year="1998">Jeng et al., 1998: 121</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AEBB54AFFB92837FC52FB58FB3FA6F5" author="Tan SH &amp; Wu S-H &amp; Huang J-F" box="[1023,1231,1199,1221]" page="143" pageId="1" pageNumber="301" year="2000">
<bibRefCitation id="EFDBEEAAFFB92837FC52FB58FB3FA6F5" author="Tan SH &amp; Wu S-H &amp; Huang J-F" box="[1023,1231,1199,1221]" pageId="1" pageNumber="301" pagination="141 - 147" refId="ref11219" refString="Tan SH, Wu S-H &amp; Huang J-F (2000) Seven new records of marine crabs from Taiwan (Crustacea: Decapoda: Brachyura: Calappidae, Lencosiidae and Cancridae). In: Hwang J-S, Wang C-H &amp; Chan T-Y (eds.) Proceedings of the International Symposium on Marine Biology in Taiwan - Crustacean and Zooplankton Taxonomy, Ecology and Living Resources. National Taiwan Museum Special Publication Series, No. 10: 141 - 147." type="book chapter" year="2000">Tan et al., 2000: 143</bibRefCitation>
</treatmentCitation>
, fig. 1d; 
<treatmentCitation id="0AEBB54AFFB92837FA85FB58FC85A6D0" author="Ng PKL &amp; Wang CH &amp; Ho PH &amp; Shih HT" page="9" pageId="1" pageNumber="301" year="2001">
<bibRefCitation id="EFDBEEAAFFB92837FA85FB58FC85A6D0" author="Ng PKL &amp; Wang CH &amp; Ho PH &amp; Shih HT" pageId="1" pageNumber="301" refId="ref10789" refString="Ng PKL, Wang CH, Ho PH &amp; Shih HT (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, Number 11: 1 - 86, 8 colour pls." type="journal volume" year="2001">Ng et al., 2001: 9</bibRefCitation>
</treatmentCitation>
(Taiwan); Chen &amp; Sun, 2002: 365, text fig. 162, pl. III–3; 
<treatmentCitation id="0AEBB54AFFB92837FCC4FB01FBDBA6CF" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[873,1067,1257,1279]" page="92" pageId="1" pageNumber="301" year="2008">
<bibRefCitation id="EFDBEEAAFFB92837FCC4FB01FBDBA6CF" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[873,1067,1257,1279]" pageId="1" pageNumber="301" pagination="1 - 286" refId="ref10633" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 92</bibRefCitation>
</treatmentCitation>
; Galil, 2007: 560; 
<treatmentCitation id="0AEBB54AFFB92837FB47FB01FA32A6CF" author="Galil BS &amp; Ng PKL" box="[1258,1474,1257,1279]" page="87" pageId="1" pageNumber="301" year="2007">
<bibRefCitation id="EFDBEEAAFFB92837FB47FB01FA32A6CF" author="Galil BS &amp; Ng PKL" box="[1258,1474,1257,1279]" pageId="1" pageNumber="301" pagination="79 - 94" refId="ref10077" refString="Galil BS &amp; Ng PKL (2007) Leucosiid crabs from Panglao, Philippines, with descriptions of three new species (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 79 - 94." type="journal article" year="2007">Galil &amp; Ng, 2007: 87</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F35DD3CBFFB92837FCDDFAEDFC12A72A" box="[880,994,1285,1307]" name="Philippines" pageId="1" pageNumber="301">Philippines</collectingCountry>
); Galil, 2015: 4; 
<treatmentCitation id="0AEBB54AFFB92837FB27FAEDFA9CA72A" author="Galil BS &amp; Ng PKL" box="[1162,1388,1285,1307]" page="460" pageId="1" pageNumber="301" year="2015">
<bibRefCitation id="EFDBEEAAFFB92837FB27FAEDFA9CA72A" author="Galil BS &amp; Ng PKL" box="[1162,1388,1285,1307]" pageId="1" pageNumber="301" pagination="451 - 486" refId="ref10159" refString="Galil BS &amp; Ng PKL (2015) Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa, 4027 (4): 451 - 486." type="journal article" year="2015">Galil &amp; Ng, 2015: 460</bibRefCitation>
</treatmentCitation>
, fig. 4B; 
<bibRefCitation id="EFDBEEAAFFB92837FCC4FACBFBD0A706" author="Prema M &amp; Ravichandran S &amp; Ng PKL" box="[873,1056,1314,1336]" pageId="1" pageNumber="301" pagination="111 - 120" refId="ref10835" refString="Prema M, Ravichandran S &amp; Ng PKL (2018) Redescription of Parilia alcocki (Wood-Mason, in Wood-Mason &amp; Alcock, 1891) (Decapoda, Brachyura, Leucosiidae) from southeast India. Zootaxa, 4378 (1): 111 - 120." type="journal article" year="2018">Prema et al., 2018</bibRefCitation>
: figs. 3, 4D, F, 5H–K.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFB92837FCEEFA88FBEDA4E9" pageId="1" pageNumber="301" type="materials_examined">
<paragraph id="8BF5935BFFB92837FCEEFA88FBEDA4E9" blockId="1.[835,1475,1376,1753]" pageId="1" pageNumber="301">
<emphasis id="B93E4F49FFB92837FCEEFA88FBDDA749" bold="true" box="[835,1069,1376,1400]" pageId="1" pageNumber="301">Material examined.</emphasis>
<materialsCitation id="3B229906FFB92837FB9AFA89FBC8A788" ID-GBIF-Occurrence="3115212308" collectingDate="2000-11-05" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Taiwan" location="Port" municipality="Nangfangao" pageId="1" pageNumber="301" specimenCode="ZRC 2001.50" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="9D4C58D2FFB92837FB9AFA89FB77A749" box="[1079,1159,1376,1401]" pageId="1" pageNumber="301" type="male">1 male</specimenCount>
, 
<specimenCount id="9D4C58D2FFB92837FB3AFA89FAF5A749" box="[1175,1285,1376,1401]" pageId="1" pageNumber="301" type="female">2 females</specimenCount>
(
<specimenCode id="DBEC3B20FFB92837FABAFA89FA46A749" box="[1303,1462,1376,1401]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="301">ZRC 2001.50</specimenCode>
), Su-Ao 
<location id="8E95C580FFB92837FC3CFA69FC30A7A9" LSID="urn:lsid:plazi:treatment:03E3224DFFB92834FC5CFC35FB1FA409:8E95C580FFB92837FC3CFA69FC30A7A9" box="[913,960,1409,1432]" country="Taiwan" municipality="Nangfangao" name="Port" pageId="1" pageNumber="301">Port</location>
, 
<collectingMunicipality id="6B910921FFB92837FC61FA68FBA1A7A9" box="[972,1105,1408,1432]" pageId="1" pageNumber="301">Nangfangao</collectingMunicipality>
, northeastern 
<collectingCountry id="F35DD3CBFFB92837FB45FA68FACBA7A9" box="[1256,1339,1408,1432]" name="Taiwan" pageId="1" pageNumber="301">Taiwan</collectingCountry>
, coll. 
<collectorName id="26BFF68DFFB92837FAD1FA69FC96A788" pageId="1" pageNumber="301">P.K.L. Ng</collectorName>
, 
<date id="FFF4B59BFFB92837FCDFFA48FBC8A788" box="[882,1080,1440,1465]" pageId="1" pageNumber="301" value="2000-11-05">
<collectingDate id="EFB04C73FFB92837FCDFFA48FBC8A788" box="[882,1080,1440,1465]" pageId="1" pageNumber="301" value="2000-11-05">5 November 2000</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B229906FFB92837FBE9FA49FB29A7C8" ID-GBIF-Occurrence="3115212310" collectingDate="2004-06" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Philippines" location="Balicasag" municipality="Panglao" pageId="1" pageNumber="301" specimenCode="ZRC 2017.217" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Bohol">
<specimenCount id="9D4C58D2FFB92837FBE9FA49FB7FA789" box="[1092,1167,1440,1465]" pageId="1" pageNumber="301" type="male">1 male</specimenCount>
, 
<specimenCount id="9D4C58D2FFB92837FB36FA49FAF3A789" box="[1179,1283,1440,1465]" pageId="1" pageNumber="301" type="female">3 females</specimenCount>
(
<specimenCode id="DBEC3B20FFB92837FABEFA49FA43A788" box="[1299,1459,1441,1465]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="301">ZRC 2017.217</specimenCode>
), station PN1, 
<location id="8E95C580FFB92837FC7BFA28FBB1A7E9" LSID="urn:lsid:plazi:treatment:03E3224DFFB92834FC5CFC35FB1FA409:8E95C580FFB92837FC7BFA28FBB1A7E9" box="[982,1089,1472,1496]" country="Philippines" municipality="Panglao" name="Balicasag" pageId="1" pageNumber="301" stateProvince="Bohol">Balicasag</location>
, 
<collectingMunicipality id="6B910921FFB92837FBE2FA28FB58A7E9" box="[1103,1192,1472,1496]" pageId="1" pageNumber="301">Panglao</collectingMunicipality>
, 
<collectingRegion id="498E5DB9FFB92837FB1AFA28FB0BA7E9" box="[1207,1275,1472,1496]" country="Philippines" name="Bohol" pageId="1" pageNumber="301">Bohol</collectingRegion>
, 
<collectingCountry id="F35DD3CBFFB92837FAA4FA28FA75A7E9" box="[1289,1413,1472,1496]" name="Philippines" pageId="1" pageNumber="301">Philippines</collectingCountry>
, coll. 
<collectorName id="26BFF68DFFB92837FCEEFA09FC42A7C8" box="[835,946,1505,1529]" pageId="1" pageNumber="301">P.K.L. Ng</collectorName>
from fishermen, 
<date id="FFF4B59BFFB92837FBC4FA09FB29A7C8" box="[1129,1241,1505,1529]" pageId="1" pageNumber="301" value="2004-06">
<collectingDate id="EFB04C73FFB92837FBC4FA09FB29A7C8" box="[1129,1241,1505,1529]" pageId="1" pageNumber="301" value="2004-06">June 2004</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="9D4C58D2FFB92837FB49FA09FAC6A7C9" box="[1252,1334,1504,1528]" pageId="1" pageNumber="301" type="male">7 males</specimenCount>
(largest 70.0 × 69.0 mm), 
<materialsCitation id="3B229906FFB92837FC67F9E9FB63A468" ID-GBIF-Occurrence="3115212311" collectingDate="2001-11-28" collectionCode="ZRC" country="Philippines" location="Panglao" pageId="1" pageNumber="301" specimenCode="ZRC 2001.552" specimenCount="4" specimenCount-female="4" stateProvince="Bohol">
<specimenCount id="9D4C58D2FFB92837FC67F9E9FBC0A429" box="[970,1072,1536,1561]" pageId="1" pageNumber="301" type="female">4 females</specimenCount>
(largest 48.3 × 
<quantity id="4CB23EBEFFB92837FB79F9E9FAC5A428" box="[1236,1333,1537,1561]" metricMagnitude="-2" metricUnit="m" metricValue="4.92" pageId="1" pageNumber="301" unit="mm" value="49.2">49.2 mm</quantity>
), 2 ovigerous females (
<specimenCode id="DBEC3B20FFB92837FC09F9C9FBB9A409" box="[932,1097,1568,1593]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="301">ZRC 2001.552</specimenCode>
), 
<location id="8E95C580FFB92837FBF6F9C8FB44A409" LSID="urn:lsid:plazi:treatment:03E3224DFFB92834FC5CFC35FB1FA409:8E95C580FFB92837FBF6F9C8FB44A409" box="[1115,1204,1568,1592]" country="Philippines" name="Panglao" pageId="1" pageNumber="301" stateProvince="Bohol">Panglao</location>
, 
<collectingRegion id="498E5DB9FFB92837FB12F9C8FAF3A409" box="[1215,1283,1568,1592]" country="Philippines" name="Bohol" pageId="1" pageNumber="301">Bohol</collectingRegion>
, 
<collectingCountry id="F35DD3CBFFB92837FAA3F9C8FA79A409" box="[1294,1417,1568,1592]" name="Philippines" pageId="1" pageNumber="301">Philippines</collectingCountry>
, coll. fishermen, 
<date id="FFF4B59BFFB92837FC11F9A9FB63A468" box="[956,1171,1600,1625]" pageId="1" pageNumber="301" value="2001-11-28">
<collectingDate id="EFB04C73FFB92837FC11F9A9FB63A468" box="[956,1171,1600,1625]" pageId="1" pageNumber="301" value="2001-11-28">28 November 2001</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B229906FFB92837FB0DF9A9FB88A4A8" ID-GBIF-Occurrence="3115212301" collectingDate="2003-11" collectionCode="ZRC" collectorName="J. Arbasto" country="Philippines" location="Maribojoc Bay" municipality="Panglao" pageId="1" pageNumber="301" specimenCode="ZRC 2017.218" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Bohol">
<specimenCount id="9D4C58D2FFB92837FB0DF9A9FAF5A469" box="[1184,1285,1600,1625]" pageId="1" pageNumber="301" type="male">10 males</specimenCount>
, 
<specimenCount id="9D4C58D2FFB92837FABFF9A8FA8CA469" box="[1298,1404,1600,1624]" pageId="1" pageNumber="301" type="female">5 females</specimenCount>
(
<specimenCode id="DBEC3B20FFB92837FA23F9A9FC56A448" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="301">ZRC 2017.218</specimenCode>
), 
<location id="8E95C580FFB92837FC16F988FBABA448" LSID="urn:lsid:plazi:treatment:03E3224DFFB92834FC5CFC35FB1FA409:8E95C580FFB92837FC16F988FBABA448" box="[955,1115,1632,1657]" country="Philippines" municipality="Panglao" name="Maribojoc Bay" pageId="1" pageNumber="301" stateProvince="Bohol">Maribojoc Bay</location>
, 
<collectingMunicipality id="6B910921FFB92837FBC9F988FB4CA449" box="[1124,1212,1632,1656]" pageId="1" pageNumber="301">Panglao</collectingMunicipality>
, 
<collectingRegion id="498E5DB9FFB92837FB68F988FAF8A449" box="[1221,1288,1632,1656]" country="Philippines" name="Bohol" pageId="1" pageNumber="301">Bohol</collectingRegion>
, 
<collectingCountry id="F35DD3CBFFB92837FABCF988FA7AA449" box="[1297,1418,1632,1656]" name="Philippines" pageId="1" pageNumber="301">Philippines</collectingCountry>
, coll. 
<collectorName id="26BFF68DFFB92837FCEEF969FC48A4A9" box="[835,952,1664,1688]" pageId="1" pageNumber="301">J. Arbasto</collectorName>
, 
<date id="FFF4B59BFFB92837FC6BF968FB88A4A8" box="[966,1144,1664,1689]" pageId="1" pageNumber="301" value="2003-11">
<collectingDate id="EFB04C73FFB92837FC6BF968FB88A4A8" box="[966,1144,1664,1689]" pageId="1" pageNumber="301" value="2003-11">November 2003</collectingDate>
</date>
</materialsCitation>
– 
<materialsCitation id="3B229906FFB92837FB36F968FAE8A4A8" ID-GBIF-Occurrence="3115212302" box="[1179,1304,1664,1689]" collectingDate="2004-04" collectionCode="ZRC" collectorName="J. Arbasto" country="Philippines" isTailEnumeration="true" location="Maribojoc Bay" municipality="Panglao" pageId="1" pageNumber="301" specimenCount="1" stateProvince="Bohol">
<date id="FFF4B59BFFB92837FB36F968FAE8A4A8" box="[1179,1304,1664,1689]" pageId="1" pageNumber="301" value="2004-04">
<collectingDate id="EFB04C73FFB92837FB36F968FAE8A4A8" box="[1179,1304,1664,1689]" pageId="1" pageNumber="301" value="2004-04">April 2004</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B229906FFB92837FA88F969FBE9A4E9" ID-GBIF-Occurrence="3115212303" country="Philippines" location="For" pageId="1" pageNumber="301" specimenCount="1">
<location id="8E95C580FFB92837FA88F969FABAA4A9" LSID="urn:lsid:plazi:treatment:03E3224DFFB92834FC5CFC35FB1FA409:8E95C580FFB92837FA88F969FABAA4A9" box="[1317,1354,1665,1688]" country="Philippines" name="For" pageId="1" pageNumber="301">For</location>
additional material from 
<collectingCountry id="F35DD3CBFFB92837FC71F948FBA4A489" box="[988,1108,1696,1720]" name="Philippines" pageId="1" pageNumber="301">Philippines</collectingCountry>
and 
<collectingCountry id="F35DD3CBFFB92837FB25F949FAAAA489" box="[1160,1370,1696,1720]" name="Papua New Guinea" pageId="1" pageNumber="301">Papua New Guinea</collectingCountry>
, see 
<bibRefCitation id="EFDBEEAAFFB92837FA20F948FC3BA4E8" author="Galil BS &amp; Ng PKL" pageId="1" pageNumber="301" pagination="79 - 94" refId="ref10077" refString="Galil BS &amp; Ng PKL (2007) Leucosiid crabs from Panglao, Philippines, with descriptions of three new species (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 79 - 94." type="journal article" year="2007">Galil &amp; Ng (2007</bibRefCitation>
, 2015)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFB92834FCEEF8E8FD4EA409" lastPageId="2" lastPageNumber="302" pageId="1" pageNumber="301" type="diagnosis">
<paragraph id="8BF5935BFFB92834FCEEF8E8FD4EA409" blockId="1.[835,1475,1792,2104]" lastBlockId="2.[113,752,1440,1592]" lastPageId="2" lastPageNumber="302" pageId="1" pageNumber="301">
<emphasis id="B93E4F49FFB92837FCEEF8E8FC4CA529" bold="true" box="[835,956,1792,1816]" pageId="1" pageNumber="301">Diagnosis.</emphasis>
Carapace globose (
<figureCitation id="13718FDEFFB92837FB32F8E8FAF2A528" box="[1183,1282,1792,1817]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="1" pageNumber="301">Figs. 1B</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FABFF8E9FAC4A528" box="[1298,1332,1793,1817]" captionStart="Fig" captionStartId="3.[136,171,1290,1312]" captionTargetBox="[136,1474,189,1275]" captionTargetId="figure-8@3.[136,1474,189,1275]" captionTargetPageId="3" captionText="Fig. 2. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; B, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; C, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503193" httpUri="https://zenodo.org/record/4503193/files/figure.png" pageId="1" pageNumber="301">2B</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FAE9F8E9FA97A528" box="[1348,1383,1793,1817]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="1" pageNumber="301">3B</figureCitation>
); dorsal surface appears almost smooth or with very low granules (
<figureCitation id="13718FDEFFB92837FCE6F8A8FC50A568" box="[843,928,1856,1881]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="1" pageNumber="301">Fig. 3B</figureCitation>
); branchial regions not distinctly swollen laterally and dorsally, lateral margin of carapace smooth (
<figureCitation id="13718FDEFFB92837FA3EF888FC96A5A8" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="1" pageNumber="301">Fig. 3B</figureCitation>
); dome-shape in frontal view (
<figureCitation id="13718FDEFFB92837FB12F868FAE7A5A8" box="[1215,1303,1920,1945]" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="1" pageNumber="301">Fig. 4C</figureCitation>
); frontal region protruding anteriorly with buccal cavity and third maxillipeds not visible in dorsal view, frontal lobes distinctly triangular (
<figureCitation id="13718FDEFFB92837FCE6F808FC50A5C8" box="[843,928,2016,2041]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="1" pageNumber="301">Fig. 3B</figureCitation>
); outer surface of third maxillipeds almost smooth or with low, flattened granules (
<figureCitation id="13718FDEFFB92837FB3BF7E8FAE3AA29" box="[1174,1299,2048,2073]" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="1" pageNumber="301">Figs. 4C, D</figureCitation>
, 
<figureCitation id="13718FDEFFB92837FAB2F7E8FA90AA28" box="[1311,1376,2048,2073]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="1" pageNumber="301">5D, E</figureCitation>
); exopod of third maxilliped in adult broad but not foliaceous (
<figureCitation id="13718FDEFFB92834FA3AF7C8FF40A789" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" lastPageId="2" lastPageNumber="302" pageId="1" pageNumber="301">Fig. 5D, E</figureCitation>
); surfaces of ambulatory legs and chelipeds completely smooth (
<figureCitation id="13718FDEFFBA2834FF7EFA28FEC5A7E9" box="[211,309,1472,1496]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="2" pageNumber="302">Figs. 1B</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FEE9FA28FE96A7E9" box="[324,358,1472,1496]" captionStart="Fig" captionStartId="3.[136,171,1290,1312]" captionTargetBox="[136,1474,189,1275]" captionTargetId="figure-8@3.[136,1474,189,1275]" captionTargetPageId="3" captionText="Fig. 2. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; B, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; C, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503193" httpUri="https://zenodo.org/record/4503193/files/figure.png" pageId="2" pageNumber="302">2B</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FED8FA28FE68A7E9" box="[373,408,1472,1496]" captionStart="Fig" captionStartId="8.[113,148,1410,1432]" captionTargetBox="[113,1451,189,1393]" captionTargetId="figure-279@8.[113,1451,189,1393]" captionText="Fig. 6. Outer view of right chela. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778841" httpUri="https://zenodo.org/record/4778841/files/figure.png" pageId="2" pageNumber="302">6B</figureCitation>
); G1 very elongate, distal part slightly flared, subdistal lateral projection dorso-ventrally flattened (
<figureCitation id="13718FDEFFBA2834FF73F9E8FEEFA429" box="[222,287,1536,1560]" captionStart="Fig" captionStartId="10.[113,148,1974,1996]" captionTargetBox="[115,1453,190,1952]" captionTargetId="figure-8@10.[99,1459,176,1964]" captionText="Fig. 8. Gonopods. A–D, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; E–I, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; J–M, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; N–S, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India. A, E, J, N, left G1 (ventral view); O, left G1 (dorsal view); B, F, K, P, distal part of left G1 (ventral view); C, G, H, L, Q, R, distal part of left G1 (dorsal view); D, I, M, S, left G2. Scales: A, D, E, I, J, M, N, O, S = 2.0 mm; B, C, F–H, K, L, P–R = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4778849" httpUri="https://zenodo.org/record/4778849/files/figure.png" pageId="2" pageNumber="302">Fig. 8</figureCitation>
E–H); female sternopleonal cavity without projection between vulvae (
<figureCitation id="13718FDEFFBA2834FE32F9C8FE07A409" box="[415,503,1568,1592]" captionStart="Fig" captionStartId="11.[136,171,1668,1690]" captionTargetBox="[136,1474,189,1653]" captionTargetId="figure-7@11.[136,1474,189,1653]" captionTargetPageId="11" captionText="Fig. 9. Female pleon and sternopleonal cavity. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; C, D, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; E, F, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India" figureDoi="http://doi.org/10.5281/zenodo.4503199" httpUri="https://zenodo.org/record/4503199/files/figure.png" pageId="2" pageNumber="302">Fig. 9D</figureCitation>
). Western Pacific.
</paragraph>
</subSubSection>
<caption id="DF35C3D3FFBA2834FFDCFB1DFB7AA775" ID-DOI="http://doi.org/10.5281/zenodo.4503191" ID-Zenodo-Dep="4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="2" pageNumber="302" startId="2.[113,148,1269,1291]" targetBox="[113,1451,189,1253]" targetPageId="2">
<paragraph id="8BF5935BFFBA2834FFDCFB1DFB7AA775" blockId="2.[113,1451,1269,1348]" pageId="2" pageNumber="302">
Fig. 1. Overall view. A, 
<taxonomicName id="4C4AE8D8FFBA2834FECFFB1DFC3FA73B" authority="Wood-Mason, in Wood-Mason &amp; Alcock, 1891" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[354,975,1269,1291]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B93E4F49FFBA2834FECFFB1DFE02A73B" box="[354,498,1269,1290]" italics="true" pageId="2" pageNumber="302">Parilia alcocki</emphasis>
Wood-Mason, in 
<bibRefCitation id="EFDBEEAAFFBA2834FD08FB1DFC3FA73B" author="Wood-Mason J &amp; Alcock A" box="[677,975,1269,1291]" pageId="2" pageNumber="302" pagination="258 - 272" refId="ref11528" refString="Wood-Mason J &amp; Alcock A (1891) Natural History Notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;, Commander R. F. Hoskyn, R. N., commanding. No. 21. On the Results of the last Season's Deepsea Dredging. Annals and Magazine of Natural History, March 1891, (6) 7 (39): 258 - 272. [For an elaboration on the authorship of the taxa described in this series of papers, see HUYS et al., 2014.]" type="journal article" year="1891">Wood-Mason &amp; Alcock, 1891</bibRefCitation>
</taxonomicName>
, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, 
<taxonomicName id="4C4AE8D8FFBA2834FF22FAFAFE94A716" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[143,356,1297,1320]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B93E4F49FFBA2834FF22FAFAFF15A716" box="[143,229,1298,1319]" italics="true" pageId="2" pageNumber="302">P. major</emphasis>
<bibRefCitation id="EFDBEEAAFFBA2834FF43FAFAFE94A716" author="Sakai T" box="[238,356,1297,1320]" pageId="2" pageNumber="302" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
</taxonomicName>
, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, 
<taxonomicName id="4C4AE8D8FFBA2834FC07FAFAFB88A716" authority="Chen, 1984" authorityName="Chen" authorityYear="1984" box="[938,1144,1298,1320]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBA2834FC07FAFAFC0CA716" box="[938,1020,1298,1319]" italics="true" pageId="2" pageNumber="302">P. ovata</emphasis>
Chen, 1984
</taxonomicName>
, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, 
<taxonomicName id="4C4AE8D8FFBA2834FED7FAC7FE07A775" authority="Ng &amp; Devi &amp; Kumar, 2018" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[378,503,1327,1348]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="pattersoni" status="sp. nov.">
<emphasis id="B93E4F49FFBA2834FED7FAC7FE07A775" box="[378,503,1327,1348]" italics="true" pageId="2" pageNumber="302">P. pattersoni</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20DF232FFBA2834FDABFAC7FD8DA775" box="[518,637,1326,1348]" pageId="2" pageNumber="302" rank="species">new species</taxonomicNameLabel>
, holotype male (53.0 × 52.6 mm) (DABFUK), India.
</paragraph>
</caption>
<subSubSection id="C350C0D0FFBA2834FFDCF988FD03A5A9" pageId="2" pageNumber="302" type="discussion">
<paragraph id="8BF5935BFFBA2834FFDCF988FD03A5A9" blockId="2.[113,755,1632,1944]" pageId="2" pageNumber="302">
<emphasis id="B93E4F49FFBA2834FFDCF988FF12A449" bold="true" box="[113,226,1632,1656]" pageId="2" pageNumber="302">Remarks.</emphasis>
<bibRefCitation id="EFDBEEAAFFBA2834FF41F988FE8AA449" author="Sakai T" box="[236,378,1632,1656]" pageId="2" pageNumber="302" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai (1961)</bibRefCitation>
described the species from three male and two female specimens from Tosa Bay in 
<collectingCountry id="F35DD3CBFFBA2834FD00F969FD1DA4A9" box="[685,749,1665,1688]" name="Japan" pageId="2" pageNumber="302">Japan</collectingCountry>
. The depository for this material was not specified and it is not known where they are now (see remarks for 
<taxonomicName id="4C4AE8D8FFBA2834FD27F928FF1AA4C9" baseAuthorityName="Sakai" baseAuthorityYear="1961" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B93E4F49FFBA2834FD27F928FF1AA4C9" italics="true" pageId="2" pageNumber="302">Neparilia tuberculata</emphasis>
</taxonomicName>
below). The species, however, is very distinct and there is now no doubt about its identity. Uncertainties about the presence of 
<taxonomicName id="4C4AE8D8FFBA2834FEB9F8C9FE80A509" authorityName="Sakai" authorityYear="1961" box="[276,368,1825,1848]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B93E4F49FFBA2834FEB9F8C9FE80A509" box="[276,368,1825,1848]" italics="true" pageId="2" pageNumber="302">P. major</emphasis>
</taxonomicName>
in the Indian Ocean by 
<bibRefCitation id="EFDBEEAAFFBA2834FDC4F8C8FD00A509" author="Sakai T" box="[617,752,1824,1848]" pageId="2" pageNumber="302" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai (1961)</bibRefCitation>
and 
<bibRefCitation id="EFDBEEAAFFBA2834FF32F8A8FE58A569" author="Tirmizi NM &amp; Serene R" box="[159,424,1856,1880]" pageId="2" pageNumber="302" pagination="21 - 32" refId="ref11302" refString="Tirmizi NM &amp; Serene R (1971) The rediscovery of two species of crabs (Decapoda, Brachyura) with observations on three other species from Pakistan. Crustaceana, 2: 21 - 32, pls. 1, 2." type="book chapter" year="1971">Tirmizi &amp; Serène (1971)</bibRefCitation>
are clarified in this paper with the discovery that this material actually belongs to a new species, 
<taxonomicName id="4C4AE8D8FFBA2834FF63F869FEA9A5A9" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[206,345,1921,1944]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="pattersoni">
<emphasis id="B93E4F49FFBA2834FF63F869FEA9A5A9" box="[206,345,1921,1944]" italics="true" pageId="2" pageNumber="302">P. pattersoni</emphasis>
</taxonomicName>
(see remarks for this species below).
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFBA2834FFDCF828FB1FA409" pageId="2" pageNumber="302" type="distribution">
<paragraph id="8BF5935BFFBA2834FFDCF828FB1FA409" blockId="2.[113,752,1984,2105]" lastBlockId="2.[813,1452,1440,1592]" pageId="2" pageNumber="302">
<emphasis id="B93E4F49FFBA2834FFDCF828FEF4A5E9" bold="true" box="[113,260,1984,2008]" pageId="2" pageNumber="302">Distribution.</emphasis>
Has been reported from 
<collectingCountry id="F35DD3CBFFBA2834FD8DF829FDADA5E9" box="[544,605,1985,2008]" name="Japan" pageId="2" pageNumber="302">Japan</collectingCountry>
and the East 
<collectingCountry id="F35DD3CBFFBA2834FFDCF808FF41A5C9" box="[113,177,2016,2040]" name="China" pageId="2" pageNumber="302">China</collectingCountry>
Sea (
<bibRefCitation id="EFDBEEAAFFBA2834FF5DF808FE84A5C9" author="Sakai T" box="[240,372,2016,2040]" pageId="2" pageNumber="302" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
, 
<bibRefCitation id="EFDBEEAAFFBA2834FE2CF808FE4AA5C9" author="Sakai T" box="[385,442,2016,2040]" pageId="2" pageNumber="302" pagination="728 - 738" refId="ref10963" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo. Samouelle G (1819) The Entomologist's Useful Compendium, or an Introduction to the Knowledge of the British Insects. London, Thomas Boys, Shih Y-J, Ho P-H, Chan T-Y &amp; Naruse T (2013) New records of leucosiid crabs (Decapoda, Brachyura) from Taiwan. Crustaceana, 86: 728 - 738." type="book chapter" year="1976">1976</bibRefCitation>
; 
<bibRefCitation id="EFDBEEAAFFBA2834FE6BF808FDACA5C9" author="Takeda M" box="[454,604,2016,2040]" pageId="2" pageNumber="302" refId="ref11144" refString="Takeda M (1993) Catalogue of Crab Specimens (Crustacea: Decapoda: Brachyura) (Part 1). National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993</bibRefCitation>
; 
<bibRefCitation id="EFDBEEAAFFBA2834FDC4F808FD1BA5C9" author="Ikeda H" box="[617,747,2016,2040]" pageId="2" pageNumber="302" refId="ref10359" refString="Ikeda H (1998) The Deep-Sea Crabs of Sagami Bay. Hayama Shiosai Museum, Kanagawa, Japan, 180 pp." type="book" year="1998">Ikeda, 1998</bibRefCitation>
; 
<bibRefCitation id="EFDBEEAAFFBA2834FFDCF7E9FE6AAA29" author="Marumura M &amp; Kosaka A" box="[113,410,2048,2072]" pageId="2" pageNumber="302" refId="ref10589" refString="Marumura M &amp; Kosaka A (2003) Catalogue of the Brachyuran and Anomuran Crabs Donated by the Late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, Kainan, 73 pp." type="book" year="2003">Marumura &amp; Kosaka, 2003</bibRefCitation>
), mainland 
<collectingCountry id="F35DD3CBFFBA2834FDBAF7E8FDA7AA29" box="[535,599,2048,2072]" name="China" pageId="2" pageNumber="302">China</collectingCountry>
(Chen &amp; Sun, 2002), 
<collectingCountry id="F35DD3CBFFBA2834FF6DF7C8FEE0AA09" box="[192,272,2080,2104]" name="Taiwan" pageId="2" pageNumber="302">Taiwan</collectingCountry>
(
<bibRefCitation id="EFDBEEAAFFBA2834FE8CF7C8FE25AA09" author="Tan SH &amp; Wu S-H &amp; Huang J-F" box="[289,469,2080,2104]" pageId="2" pageNumber="302" pagination="141 - 147" refId="ref11219" refString="Tan SH, Wu S-H &amp; Huang J-F (2000) Seven new records of marine crabs from Taiwan (Crustacea: Decapoda: Brachyura: Calappidae, Lencosiidae and Cancridae). In: Hwang J-S, Wang C-H &amp; Chan T-Y (eds.) Proceedings of the International Symposium on Marine Biology in Taiwan - Crustacean and Zooplankton Taxonomy, Ecology and Living Resources. National Taiwan Museum Special Publication Series, No. 10: 141 - 147." type="book chapter" year="2000">Tan et al., 2000</bibRefCitation>
; 
<bibRefCitation id="EFDBEEAAFFBA2834FE4FF7C9FD7CAA09" author="Ng PKL &amp; Wang CH &amp; Ho PH &amp; Shih HT" box="[482,652,2080,2105]" pageId="2" pageNumber="302" refId="ref10789" refString="Ng PKL, Wang CH, Ho PH &amp; Shih HT (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, Number 11: 1 - 86, 8 colour pls." type="journal volume" year="2001">Ng et al., 2001</bibRefCitation>
; Chen &amp; Sun, 2002), 
<collectingCountry id="F35DD3CBFFBA2834FC1AFA48FBC0A789" box="[951,1072,1440,1464]" name="Philippines" pageId="2" pageNumber="302">Philippines</collectingCountry>
(Chen, 1989; 
<bibRefCitation id="EFDBEEAAFFBA2834FB79FA49FAB8A789" author="Tan CGS" box="[1236,1352,1440,1464]" pageId="2" pageNumber="302" pagination="1021 - 1058" refId="ref11174" refString="Tan CGS (1996) Leucosiidae of the Albatross expedition to the Philippines, 1907 - 1910 (Crustacea: Brachyura: Decapoda). Journal of Natural History, 30 (7): 1021 - 1058, figs. 1 - 8." type="journal article" year="1996">Tan, 1996</bibRefCitation>
; 
<bibRefCitation id="EFDBEEAAFFBA2834FAFBFA48FC60A7E9" author="Galil BS &amp; Ng PKL" pageId="2" pageNumber="302" pagination="79 - 94" refId="ref10077" refString="Galil BS &amp; Ng PKL (2007) Leucosiid crabs from Panglao, Philippines, with descriptions of three new species (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 79 - 94." type="journal article" year="2007">Galil &amp; Ng, 2007</bibRefCitation>
) to Makassar (
<collectingCountry id="F35DD3CBFFBA2834FB87FA28FB65A7E9" box="[1066,1173,1472,1496]" name="Indonesia" pageId="2" pageNumber="302">Indonesia</collectingCountry>
) (Chen, 1990) 
<collectingCountry id="F35DD3CBFFBA2834FA9EFA29FC8CA7C9" name="Papua New Guinea" pageId="2" pageNumber="302">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="EFDBEEAAFFBA2834FC23FA08FBABA7C9" author="Galil BS &amp; Ng PKL" box="[910,1115,1504,1529]" pageId="2" pageNumber="302" pagination="451 - 486" refId="ref10159" refString="Galil BS &amp; Ng PKL (2015) Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa, 4027 (4): 451 - 486." type="journal article" year="2015">Galil &amp; Ng, 2015</bibRefCitation>
) and 
<collectingCountry id="F35DD3CBFFBA2834FB32FA08FAAAA7C9" box="[1183,1370,1504,1528]" name="Solomon Islands" pageId="2" pageNumber="302">Solomon Islands</collectingCountry>
(Galil, 2007). The species occurs at depths between 
<quantity id="4CB23EBEFFBA2834FA83F9E8FA5CA429" box="[1326,1452,1536,1560]" metricMagnitude="2" metricUnit="m" metricValue="2.785" metricValueMax="4.57" metricValueMin="1.0" pageId="2" pageNumber="302" unit="m" value="278.5" valueMax="457.0" valueMin="100.0">100–457 m</quantity>
(Alcock, 1899; Chen, 1989; Galil, 2015).
</paragraph>
</subSubSection>
</treatment>
</document>