317 lines
44 KiB
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317 lines
44 KiB
XML
<document id="2DBC0061159551685401A949B83E5843" ID-CLB-Dataset="264223" ID-DOI="10.5852/ejt.2023.888.2237" ID-GBIF-Dataset="c0a2f738-d7c6-4c71-b178-058487b3b26a" ID-ISSN="2118-9773" ID-Zenodo-Dep="8269539" ID-ZooBank="56507FD3-29FE-4CF7-B205-A339EC9D471A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692626437556" checkinUser="felipe" docAuthor="Frau, Camille" docDate="2023" docId="03E23D6D4B1BFFB7396FFC35ED4219AA" docLanguage="en" docName="ejt-888_137-158.pdf" docOrigin="European Journal of Taxonomy 888 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.888.2237" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Protacanthoplites abichi" docType="treatment" docVersion="5" lastPageNumber="141" masterDocId="FFDB45154B19FFB33B4BFF93EC2C1E7E" masterDocTitle="The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900)" masterLastPageNumber="158" masterPageNumber="137" pageNumber="139" updateTime="1699595454097" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4AD6AB514813498C5611525A681CFCAB">The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900)</mods:title>
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<mods:namePart id="5F71FD02882137D7FFE329373D6EC87C">Frau, Camille</mods:namePart>
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<mods:affiliation id="0DD2E1BE8DB5E74514B38149C92AE525">Groupement d’Intérêt Paléontologique, Science et Exposition, 60 bd Georges Richard, 83000 Toulon, France</mods:affiliation>
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<mods:nameIdentifier id="21A175138B963C2D4C45BA30FF54BCB8" type="email">camille_frau@hotmail.fr</mods:nameIdentifier>
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<mods:title id="4D2BE0788A85E3BA855F4B92070A4C3C">European Journal of Taxonomy</mods:title>
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<mods:date id="F74808A5BD75BA8096768C48BC207883">2023</mods:date>
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<treatment id="03E23D6D4B1BFFB7396FFC35ED4219AA" ID-DOI="http://doi.org/10.5281/zenodo.8270488" ID-GBIF-Taxon="212448180" ID-Zenodo-Dep="8270488" LSID="urn:lsid:plazi:treatment:03E23D6D4B1BFFB7396FFC35ED4219AA" httpUri="http://treatment.plazi.org/id/03E23D6D4B1BFFB7396FFC35ED4219AA" lastPageId="4" lastPageNumber="141" pageId="2" pageNumber="139">
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<subSubSection id="C351DFF04B1BFFB1396FFC35E8231DBF" box="[548,1039,934,961]" pageId="2" pageNumber="139" type="nomenclature">
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<paragraph id="8BF48C7B4B1BFFB1396FFC35E8231DBF" blockId="2.[548,1039,934,996]" box="[548,1039,934,961]" pageId="2" pageNumber="139">
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<heading id="D0BC3B174B1BFFB1396FFC35E8231DBF" box="[548,1039,934,961]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="139" reason="2">
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<taxonomicName id="4C4BF7F84B1BFFB1396FFC35E8231DBF" ID-CoL="BTFWV" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[548,1039,934,961]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="2" pageNumber="139" phylum="Mollusca" rank="species" species="abichi">
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<emphasis id="B93F50694B1BFFB1396FFC35EF601DBF" bold="true" box="[548,844,934,961]" italics="true" pageId="2" pageNumber="139">Protacanthoplites abichi</emphasis>
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(
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<bibRefCitation id="EFDAF18A4B1BFFB13817FC34E82B1DBF" author="Anthula" box="[860,1031,935,961]" firstAuthor="Anthula" pageId="2" pageNumber="139" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C351DFF04B1BFFB139FBFC5AEE211A5A" pageId="2" pageNumber="139" type="description">
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<paragraph id="8BF48C7B4B1BFFB139FBFC5AEFAF1D9A" blockId="2.[548,1039,934,996]" box="[688,899,969,996]" pageId="2" pageNumber="139">
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<figureCitation id="137090FE4B1BFFB139FBFC5AEF181D9D" box="[688,820,969,995]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="2" pageNumber="139">Figs 2A–K</figureCitation>
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,
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<figureCitation id="137090FE4B1BFFB1380BFC5AEFAF1D9A" box="[832,899,969,996]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[223,1364,265,1748]" captionTargetId="figure-81@6.[215,929,275,1535]" captionTargetPageId="6" captionText="Fig. 3. Illustration of the ontogenetic stages observed after the ammonitella (not seen), including Royerianum (green), Abichi (yellow), Crassicostatum-like (orange), Tobleri (violet), and Aschiltaensis (blue). A. Protacanthoplites abichi (Anthula, 1900), lectotype (IPUW 1900-3a). B. Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105). C. A. aschiltaensis, paralectotype (PMU.24107). D–J. Schematic evolution of the whorl section of the plexus P. abichi – A. aschiltaensis. D–E. After Bogdanova & Mikhailova (2016: fig. 57), not to scale. F. After IPUW 1900-3a, not to scale. G. After paralectotype (PMU.24108), at scale. H–J. After paralectotype (PMU.24107), at scale." figureDoi="http://doi.org/10.5281/zenodo.8269546" httpUri="https://zenodo.org/record/8269546/files/figure.png" pageId="2" pageNumber="139">3A, F</figureCitation>
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</paragraph>
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<paragraph id="8BF48C7B4B1BFFB13BF6FB9AEE211A5A" blockId="2.[189,525,1033,1060]" box="[189,525,1033,1060]" pageId="2" pageNumber="139">
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<heading id="D0BC3B174B1BFFB13BF6FB9AEE211A5A" bold="true" box="[189,525,1033,1060]" fontSize="11" level="3" pageId="2" pageNumber="139" reason="3">
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<emphasis id="B93F50694B1BFFB13BF6FB9AEE211A5A" bold="true" box="[189,525,1033,1060]" pageId="2" pageNumber="139">Conch shape and ontogeny</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351DFF04B1BFFB13BA1FBA4EE0C1A2F" box="[234,544,1079,1105]" pageId="2" pageNumber="139" type="materials_examined">
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<paragraph id="8BF48C7B4B1BFFB13BA1FBA4EE0C1A2F" blockId="2.[189,1399,1079,1345]" box="[234,544,1079,1105]" pageId="2" pageNumber="139">
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<materialsCitation id="3B2386264B1BFFB13BA1FBA4EE0C1A2F" ID-GBIF-Occurrence="4398934302" box="[234,544,1079,1105]" collectionCode="IPUW" pageId="2" pageNumber="139" specimenCode="IPUW 1900-3" specimenCount="1" typeStatus="lectotype">
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<emphasis id="B93F50694B1BFFB13BA1FBA4EE0C1A2F" bold="true" box="[234,544,1079,1105]" pageId="2" pageNumber="139">
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<typeStatus id="54F032D94B1BFFB13BA1FBA4ED4A1A2F" box="[234,358,1079,1105]" pageId="2" pageNumber="139" type="lectotype">Lectotype</typeStatus>
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<specimenCode id="DBED24004B1BFFB13A26FBA4EE3C1A2F" box="[365,528,1079,1105]" collectionCode="IPUW" country="Austria" httpUri="http://grbio.org/cool/60hn-b4s1" name="Institut fuer Palaeontologie der Universitaet Wien" pageId="2" pageNumber="139">IPUW 1900-3</specimenCode>
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a
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</emphasis>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351DFF04B1BFFB73BF6FBCAE8771D36" lastPageId="4" lastPageNumber="141" pageId="2" pageNumber="139" type="description">
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<paragraph id="8BF48C7B4B1BFFB13BF6FBCAEE881B3F" blockId="2.[189,1399,1079,1345]" pageId="2" pageNumber="139">
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Understanding of the
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<typeStatus id="54F032D94B1BFFB13A88FBCAEE031A0D" box="[451,559,1113,1139]" pageId="2" pageNumber="139" type="lectotype">lectotype</typeStatus>
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was so far based on the hand-drawing of
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<bibRefCitation id="EFDAF18A4B1BFFB13F60FBCAE8F01A0D" author="Anthula" box="[1067,1244,1113,1139]" firstAuthor="Anthula" pageId="2" pageNumber="139" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900</bibRefCitation>
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: pl. IX(VIII) fig. 2a–c) (see
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<figureCitation id="137090FE4B1BFFB13A24FBE8EDDD1AEB" box="[367,497,1147,1173]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="2" pageNumber="139">Fig. 2A–B</figureCitation>
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). It is here properly illustrated for the first time (
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<figureCitation id="137090FE4B1BFFB13F72FBE8E8901AEB" box="[1081,1212,1147,1173]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="2" pageNumber="139">Fig. 2C–G</figureCitation>
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). The specimen corresponds to a small-sized, 3D-preserved internal mould with a calcitic phragmocone and calcareous body chamber with trace of aragonitic shell. The body chamber occupies half of the last whorl, but the peristome is not preserved. Measurements are given in
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<tableCitation id="C6C9B9C04B1BFFB138DEFB72EFDD1A82" box="[917,1009,1249,1276]" captionStart="Table 1" captionStartId="4.[189,259,265,291]" captionTargetBox="[433,1156,377,457]" captionTargetPageId="4" captionText="Table 1. Dimensions of Protacanthoplites abichi (Anthula, 1900), lectotype (IPUW 1900-3a). *measurements taken between ventral ribs. Abbreviations: see Material and methods." httpUri="http://table.plazi.org/id/DF34DCF34B1DFFB73BF6FE9AE88B1F38" pageId="2" pageNumber="139" tableUuid="DF34DCF34B1DFFB73BF6FE9AE88B1F38">Table 1</tableCitation>
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. Shell shape of the
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<typeStatus id="54F032D94B1BFFB13FA7FB72E9741A85" box="[1260,1368,1249,1275]" pageId="2" pageNumber="139" type="lectotype">lectotype</typeStatus>
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is characterised by a discoidal (Ww/D ~ 0.41), weakly depressed (Ww/Wh ~ 1.08), very evolute (U/Wh ~ 1.18) subophiocone coiling (U/D ~ 0.45).
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</paragraph>
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<paragraph id="8BF48C7B4B1BFFB13BF6FAF5E8C51B99" blockId="2.[189,1399,1382,1511]" pageId="2" pageNumber="139">
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The embryonic (Ammonitella) stage is poorly preserved, and remains poorly characterised. The first visible whorls of the shell develop a reniform whorl section, crateriform umbilicus, spaced and flat-topped ribs angulate at shoulders followed by smooth interspaces. This corresponds to the Royerianum stage that is typical of the early ontogeny of the basal
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<taxonomicName id="4C4BF7F84B1BFFB1387FFA5EE8211B99" authorityName="Stoyanow" authorityYear="1949" box="[820,1037,1485,1511]" class="Cephalopoda" family="Acanthohoplitidae" kingdom="Animalia" order="Ammonitida" pageId="2" pageNumber="139" phylum="Mollusca" rank="family">Acanthohoplitidae</taxonomicName>
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(
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<bibRefCitation id="EFDAF18A4B1BFFB13F57FA5EE8F11B99" author="Frau" box="[1052,1245,1485,1511]" etAl="et al." firstAuthor="Frau" pageId="2" pageNumber="139" pagination="1 - 16" refId="ref11154" refString="Frau C. 2020 a. New insight on the age and significance of the Aparein black shales in the Basque- Cantabrian Basin, northern Spain. Newsletters on Stratigraphy 54 (1): 1 - 16. https: // doi. org / 10.1127 / nos / 2020 / 0583" type="journal article" year="2020">
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Frau
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<emphasis id="B93F50694B1BFFB13F10FA5EE8A31B99" box="[1115,1167,1485,1511]" italics="true" pageId="2" pageNumber="139">et al</emphasis>
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. 2020
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</bibRefCitation>
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||
).
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</paragraph>
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<paragraph id="8BF48C7B4B1BFFB13BF6F99EEDAD199D" blockId="2.[187,1400,1549,2019]" pageId="2" pageNumber="139">
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Then, the whorl section progressively becomes depressed, sub-rounded with convex flanks. The umbilical wall is rounded. The ornamentation consists of an alternation of straight to convex, strong primary ribs and variable number, two to four, of sharp atuberculate secondaries, irregular in thickness and irregularly arranged. Primary ribs are gradually thickened on the flank. They develop strong, elongated tubercles, sometimes clavate, located high on the flank, in tight contact with the succeeding whorl. The tuberculate ribs get a distinct hexagonal cross section. The tubercles divide into two, rarely three, branches over the venter. They strengthen as fold-like ribs, notably the posterior branch, and generally join on the tubercles of the other side. A third discrete rib sometimes appears between two ventral ribs. The secondary ribs are simple, rarely bifurcate, and can be coalescent on the primaries forming a polygyrate pattern. Primary ribs can be followed by an approximated and enlarged secondary rib so that they mimic constrictions. By these features, the ontogeny closely resembles the Gargasense stage sensu
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<bibRefCitation id="EFDAF18A4B1BFFB13F17F8F0E9061902" author="Frau" box="[1116,1322,1890,1917]" etAl="et al." firstAuthor="Frau" pageId="2" pageNumber="139" pagination="1 - 16" refId="ref11154" refString="Frau C. 2020 a. New insight on the age and significance of the Aparein black shales in the Basque- Cantabrian Basin, northern Spain. Newsletters on Stratigraphy 54 (1): 1 - 16. https: // doi. org / 10.1127 / nos / 2020 / 0583" type="journal article" year="2020">
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Frau
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<emphasis id="B93F50694B1BFFB13FD2F8F0E8E01902" box="[1177,1228,1890,1916]" italics="true" pageId="2" pageNumber="139">et al</emphasis>
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. (2020)
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</bibRefCitation>
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of the
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<taxonomicName id="4C4BF7F84B1BFFB13BF5F817EDBB19E0" authorityName="Stoyanow" authorityYear="1949" box="[190,407,1924,1950]" class="Cephalopoda" family="Acanthohoplitidae" kingdom="Animalia" order="Ammonitida" pageId="2" pageNumber="139" phylum="Mollusca" rank="family">Acanthohoplitidae</taxonomicName>
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<taxonomicName id="4C4BF7F84B1BFFB13AD6F817EED419E0" baseAuthorityName="d'Orbigny" baseAuthorityYear="1841" box="[413,760,1924,1950]" class="Cephalopoda" family="Deshayesitidae" genus="Colombiceras" kingdom="Animalia" pageId="2" pageNumber="139" phylum="Mollusca" rank="species" species="crassicostatum">
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<emphasis id="B93F50694B1BFFB13AD6F817EED419E0" box="[413,760,1924,1950]" italics="true" pageId="2" pageNumber="139">Colombiceras crassicostatum</emphasis>
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</taxonomicName>
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but differences are seen in the quadratic whorl section with rounded venter, and distinctive strengthened ribbing and tubercles. This stage is here referred to as the Abichi stage.
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</paragraph>
|
||
<caption id="DF34DCF34B1AFFB03BF6F8D2ED3B1997" ID-DOI="http://doi.org/10.5281/zenodo.8269544" ID-Zenodo-Dep="8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="3" pageNumber="140" startId="3.[189,232,1857,1883]" targetBox="[188,1399,225,1818]" targetPageId="3">
|
||
<paragraph id="8BF48C7B4B1AFFB03BF6F8D2ED3B1997" blockId="3.[187,1399,1857,2025]" pageId="3" pageNumber="140">
|
||
<emphasis id="B93F50694B1AFFB03BF6F8D2ED781925" bold="true" box="[189,340,1857,1883]" pageId="3" pageNumber="140">Fig. 2. A–K</emphasis>
|
||
.
|
||
<taxonomicName id="4C4BF7F84B1AFFB03A2CF8D2EF211922" authority="Anthula, 1900" authorityName="Anthula" authorityYear="1900" box="[359,781,1857,1884]" class="Cephalopoda" family="Deshayesitidae" genus="Parahoplites" kingdom="Animalia" pageId="3" pageNumber="140" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1AFFB03A2CF8D2EE7F1925" box="[359,595,1857,1883]" italics="true" pageId="3" pageNumber="140">Parahoplites abichi</emphasis>
|
||
<bibRefCitation id="EFDAF18A4B1AFFB03916F8D2EF211922" author="Anthula" box="[605,781,1857,1884]" firstAuthor="Anthula" pageId="3" pageNumber="140" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93F50694B1AFFB03857F8D2EF781922" bold="true" box="[796,852,1857,1884]" pageId="3" pageNumber="140">A–B</emphasis>
|
||
. Reproduction of the original hand-drawing.
|
||
<emphasis id="B93F50694B1AFFB03BF6F8F7ECD41900" bold="true" box="[189,248,1892,1919]" pageId="3" pageNumber="140">C–G</emphasis>
|
||
. Lectotype (IPUW 1900-3a).
|
||
<emphasis id="B93F50694B1AFFB03919F8F7EE451900" bold="true" box="[594,617,1892,1918]" pageId="3" pageNumber="140">G</emphasis>
|
||
. Focus on its inner whorls.
|
||
<emphasis id="B93F50694B1AFFB038E2F8F6EFCB1901" bold="true" box="[937,999,1893,1919]" pageId="3" pageNumber="140">H–K</emphasis>
|
||
. Syntype (IPUW 1900-3b), which is here excluded from the species. –
|
||
<emphasis id="B93F50694B1AFFB03911F81BEEBD19DC" bold="true" box="[602,657,1928,1954]" pageId="3" pageNumber="140">L–R</emphasis>
|
||
.
|
||
<taxonomicName id="4C4BF7F84B1AFFB039EBF81BE8AE19DC" authority="Anthula, 1900" authorityName="Anthula" authorityYear="1900" box="[672,1154,1928,1954]" class="Cephalopoda" family="Deshayesitidae" genus="Parahoplites" kingdom="Animalia" pageId="3" pageNumber="140" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B1AFFB039EBF81BEFF919DC" box="[672,981,1928,1954]" italics="true" pageId="3" pageNumber="140">Parahoplites aschiltaensis</emphasis>
|
||
<bibRefCitation id="EFDAF18A4B1AFFB03892F81BE8AE19DC" author="Anthula" box="[985,1154,1928,1954]" firstAuthor="Anthula" pageId="3" pageNumber="140" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93F50694B1AFFB03FC3F81BE8EB19DC" bold="true" box="[1160,1223,1928,1954]" pageId="3" pageNumber="140">L–M</emphasis>
|
||
. Reproduction of the original hand-drawing.
|
||
<emphasis id="B93F50694B1AFFB03956F838EE7A19BB" bold="true" box="[541,598,1963,1990]" pageId="3" pageNumber="140">N–R</emphasis>
|
||
. Lectotype (PMU.24105).
|
||
<emphasis id="B93F50694B1AFFB038DEF838EF8619BB" bold="true" box="[917,938,1963,1989]" pageId="3" pageNumber="140">R</emphasis>
|
||
. Focus on its inner whorls. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF34DCF34B1DFFB73BF6FE9AE88B1F38" ID-Table-UUID="DF34DCF34B1DFFB73BF6FE9AE88B1F38" httpUri="http://table.plazi.org/id/DF34DCF34B1DFFB73BF6FE9AE88B1F38" pageId="4" pageNumber="141" startId="4.[189,259,265,291]" targetBox="[433,1156,377,457]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FE9AE88B1F38" blockId="4.[189,1399,265,327]" pageId="4" pageNumber="141">
|
||
<emphasis id="B93F50694B1DFFB73BF6FE9AED071F5D" bold="true" box="[189,299,265,291]" pageId="4" pageNumber="141">Table 1.</emphasis>
|
||
Dimensions of
|
||
<taxonomicName id="4C4BF7F84B1DFFB7394EFE9AE8201F5D" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[517,1036,265,291]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1DFFB7394EFE9AEF1D1F5D" box="[517,817,265,291]" italics="true" pageId="4" pageNumber="141">Protacanthoplites abichi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73800FE9AE82F1F5D" author="Anthula" box="[843,1027,265,291]" firstAuthor="Anthula" pageId="4" pageNumber="141" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype (IPUW 1900-3a). *measurements taken between ventral ribs. Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF48C7B4B1DFFB73A86FEEAE8A81FB6" pageId="4" pageNumber="141">
|
||
<table id="F94B7EDB4B1D004C3AFAFEEAE8A81FB7" box="[433,1156,377,457]" gridcols="5" gridrows="2" pageId="4" pageNumber="141">
|
||
<tr id="357B8E394B1D004C3AFAFEEAE8A81FED" box="[433,1156,377,403]" gridrow="0" pageId="4" pageNumber="141">
|
||
<th id="76AAE7454B1D004C3AFAFEEAEE4E1FED" box="[433,610,377,403]" gridcol="0" gridrow="0" pageId="4" pageNumber="141">Specimen</th>
|
||
<td id="76AAE7454B1D004C39E5FEEAEECD1FED" box="[686,737,377,403]" gridcol="1" gridrow="0" pageId="4" pageNumber="141">D</td>
|
||
<td id="76AAE7454B1D004C387CFEEAEF461FED" box="[823,874,377,403]" gridcol="2" gridrow="0" pageId="4" pageNumber="141">U</td>
|
||
<td id="76AAE7454B1D004C38FFFEEAEFD01FED" box="[948,1020,377,403]" gridcol="3" gridrow="0" pageId="4" pageNumber="141">Wh*</td>
|
||
<td id="76AAE7454B1D004C3F77FEEAE8A81FED" box="[1084,1156,377,403]" gridcol="4" gridrow="0" pageId="4" pageNumber="141">
|
||
<emphasis id="B93F50694B1DFFB73F0BFEEAE8561FED" bold="true" box="[1088,1146,377,403]" pageId="4" pageNumber="141">Ww*</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="357B8E394B1D004C3AFAFE3DE8A81FB7" box="[433,1156,430,457]" gridrow="1" pageId="4" pageNumber="141">
|
||
<th id="76AAE7454B1D004C3AFAFE3DEE4E1FB7" box="[433,610,430,457]" gridcol="0" gridrow="1" pageId="4" pageNumber="141">IPUW 1900-3a</th>
|
||
<td id="76AAE7454B1D004C39E5FE3DEECD1FB7" box="[686,737,430,457]" gridcol="1" gridrow="1" pageId="4" pageNumber="141">33.9</td>
|
||
<td id="76AAE7454B1D004C387CFE3DEF461FB7" box="[823,874,430,457]" gridcol="2" gridrow="1" pageId="4" pageNumber="141">15.2</td>
|
||
<td id="76AAE7454B1D004C38FFFE3DEFD01FB7" box="[948,1020,430,457]" gridcol="3" gridrow="1" pageId="4" pageNumber="141">c.12.9</td>
|
||
<td id="76AAE7454B1D004C3F77FE3DE8A81FB7" box="[1084,1156,430,457]" gridcol="4" gridrow="1" pageId="4" pageNumber="141">c.14.0</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FD8AE87E1D79" blockId="4.[188,1400,537,775]" pageId="4" pageNumber="141">
|
||
As growth proceeds, the number of intercalatories decreases, being mostly one to two, and coalescent ribs become sporadic. Primary ribs are dominantly bifurcate, then simple. They thicken along the flank, especially at the umbilical margin, while the tubercles tend to disappear. Ribs distinctly bend forward on the venter. All ribs are flat-topped or cuneiform over the venter. This stage mimics the Crassicostatum stage sensu
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73A05FD35EE0E1CBE" author="Frau" box="[334,546,678,704]" etAl="et al." firstAuthor="Frau" pageId="4" pageNumber="141" pagination="1 - 16" refId="ref11154" refString="Frau C. 2020 a. New insight on the age and significance of the Aparein black shales in the Basque- Cantabrian Basin, northern Spain. Newsletters on Stratigraphy 54 (1): 1 - 16. https: // doi. org / 10.1127 / nos / 2020 / 0583" type="journal article" year="2020">
|
||
Frau
|
||
<emphasis id="B93F50694B1DFFB73ADBFD34EDEF1CBE" box="[400,451,678,704]" italics="true" pageId="4" pageNumber="141">et al</emphasis>
|
||
. (2020)
|
||
</bibRefCitation>
|
||
of
|
||
<taxonomicName id="4C4BF7F84B1DFFB7391AFD35EF811CBE" baseAuthorityName="d'Orbigny" baseAuthorityYear="1841" box="[593,941,678,704]" class="Cephalopoda" family="Deshayesitidae" genus="Colombiceras" kingdom="Animalia" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="crassicostatum">
|
||
<emphasis id="B93F50694B1DFFB7391AFD35EF811CBE" box="[593,941,678,704]" italics="true" pageId="4" pageNumber="141">Colombiceras crassicostatum</emphasis>
|
||
</taxonomicName>
|
||
, but differences are seen in the whorl section that is broadly rounded, with subhexagonal rib section with a convex venter lacking ventral attenuation. This sequence is here referred to as the Crassicostatum-like stage.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FCBDE8771D36" blockId="4.[189,1115,814,840]" box="[189,1115,814,840]" pageId="4" pageNumber="141">
|
||
The ontogenetic sequence of
|
||
<taxonomicName id="4C4BF7F84B1DFFB7395EFCBDEF1A1D36" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[533,822,814,840]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1DFFB7395EFCBDEF1A1D36" box="[533,822,814,840]" italics="true" pageId="4" pageNumber="141">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
is illustrated on
|
||
<figureCitation id="137090FE4B1DFFB738B1FCBDE87B1D36" box="[1018,1111,814,840]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[223,1364,265,1748]" captionTargetId="figure-81@6.[215,929,275,1535]" captionTargetPageId="6" captionText="Fig. 3. Illustration of the ontogenetic stages observed after the ammonitella (not seen), including Royerianum (green), Abichi (yellow), Crassicostatum-like (orange), Tobleri (violet), and Aschiltaensis (blue). A. Protacanthoplites abichi (Anthula, 1900), lectotype (IPUW 1900-3a). B. Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105). C. A. aschiltaensis, paralectotype (PMU.24107). D–J. Schematic evolution of the whorl section of the plexus P. abichi – A. aschiltaensis. D–E. After Bogdanova & Mikhailova (2016: fig. 57), not to scale. F. After IPUW 1900-3a, not to scale. G. After paralectotype (PMU.24108), at scale. H–J. After paralectotype (PMU.24107), at scale." figureDoi="http://doi.org/10.5281/zenodo.8269546" httpUri="https://zenodo.org/record/8269546/files/figure.png" pageId="4" pageNumber="141">Fig. 3A</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B1DFFB73BA1FCFCEE321DF7" box="[234,542,879,905]" pageId="4" pageNumber="141" type="materials_examined">
|
||
<paragraph id="8BF48C7B4B1DFFB73BA1FCFCEE321DF7" blockId="4.[188,1401,879,1117]" box="[234,542,879,905]" pageId="4" pageNumber="141">
|
||
<emphasis id="B93F50694B1DFFB73BA1FCFCEE321DF7" bold="true" box="[234,542,879,905]" pageId="4" pageNumber="141">Specimen IPUW 1900-3b</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B1DFFB73BF6FC01ED4219AA" pageId="4" pageNumber="141" type="description">
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FC01EF891A23" blockId="4.[188,1401,879,1117]" pageId="4" pageNumber="141">
|
||
The second
|
||
<typeStatus id="54F032D94B1DFFB73A04FC00ED861DD3" box="[335,426,915,941]" pageId="4" pageNumber="141" type="syntype">syntype</typeStatus>
|
||
of
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73A98FC01EEA51DD2" author="Anthula" box="[467,649,914,940]" firstAuthor="Anthula" pageId="4" pageNumber="141" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900)</bibRefCitation>
|
||
is here properly illustrated for the first time (
|
||
<figureCitation id="137090FE4B1DFFB73FE6FC01E91E1DD2" box="[1197,1330,914,940]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="4" pageNumber="141">Fig. 2H–K</figureCitation>
|
||
). It is a small, 3D-preserved internal mould with a calcitic phragmocone and calcareous body chamber with remains of aragonitic shell. It shows a slight distortion of the shell at the beginning of the body chamber. The latter occupies half of the last whorl. The peristome is not preserved. Measurements are given in
|
||
<tableCitation id="C6C9B9C04B1DFFB73BF4FB8CED341A47" box="[191,280,1055,1081]" captionStart="Table 2" captionStartId="5.[189,259,265,291]" captionTargetBox="[432,1145,377,457]" captionTargetPageId="5" captionText="Table 2. Dimensions of Protacanthoplites abichi (Anthula, 1900), specimen IPUW 1900-3b. *measurements taken between ventral ribs. Abbreviations: see Material and methods." httpUri="http://table.plazi.org/id/DF34DCF34B1CFFB63BF6FE9AE88B1F38" pageId="4" pageNumber="141" tableUuid="DF34DCF34B1CFFB63BF6FE9AE88B1F38">Table 2</tableCitation>
|
||
. Shell shape is characterised by a discoidal (Ww/D ~ 0.39), weakly depressed (Ww/Wh ~ 1.02), very evolute (U/Wh ~ 0,91) subvirgacone coiling (U/D ~ 0.35).
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FB17ED771B30" blockId="4.[188,1401,1156,1358]" pageId="4" pageNumber="141">
|
||
The first visible whorl of the shell develops the Royerianum stage, changing into the Abichi stage of the
|
||
<typeStatus id="54F032D94B1DFFB73BF6FB34ED051ABF" box="[189,297,1191,1217]" pageId="4" pageNumber="141" type="lectotype">lectotype</typeStatus>
|
||
<taxonomicName id="4C4BF7F84B1DFFB73A79FB34EE7A1ABF" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[306,598,1191,1217]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1DFFB73A79FB34EE7A1ABF" box="[306,598,1191,1217]" italics="true" pageId="4" pageNumber="141">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
. To the difference, the ribbing is more gracile, with discrete spiny tubercles. The venter is narrower. As growth proceeds, the Crassicostatum-like stage initiates on the body chamber, but it is also much more gracile and regular than the
|
||
<typeStatus id="54F032D94B1DFFB73F40FB7EE85B1B79" box="[1035,1143,1261,1287]" pageId="4" pageNumber="141" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4BF7F84B1DFFB73FEEFB7EED2A1B54" baseAuthorityName="Anthula" baseAuthorityYear="1900" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1DFFB73FEEFB7EED2A1B54" italics="true" pageId="4" pageNumber="141">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
, and develops flexuous primary ribs. By these features, specimen IPUW 1900-3b poorly matches the
|
||
<typeStatus id="54F032D94B1DFFB73BA3FAA7ED7B1B30" box="[232,343,1332,1358]" pageId="4" pageNumber="141" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FAE6ED6B1BF1" blockId="4.[189,327,1397,1423]" box="[189,327,1397,1423]" pageId="4" pageNumber="141">
|
||
<heading id="D0BC3B174B1DFFB73BF6FAE6ED6B1BF1" bold="true" box="[189,327,1397,1423]" fontSize="11" level="3" pageId="4" pageNumber="141" reason="3">
|
||
<emphasis id="B93F50694B1DFFB73BF6FAE6ED6B1BF1" bold="true" box="[189,327,1397,1423]" pageId="4" pageNumber="141">Suture line</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6FA37E8B418EF" blockId="4.[189,1400,1444,1681]" pageId="4" pageNumber="141">
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73BF6FA37ED401BC0" author="Anthula" box="[189,364,1444,1470]" firstAuthor="Anthula" pageId="4" pageNumber="141" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900</bibRefCitation>
|
||
: pl. IX (VIII) fig. 2c) provided a partial suture line drawing for
|
||
<taxonomicName id="4C4BF7F84B1DFFB73F1EFA37E95A1BC0" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[1109,1398,1444,1470]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1DFFB73F1EFA37E95A1BC0" box="[1109,1398,1444,1470]" italics="true" pageId="4" pageNumber="141">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
showing a quinquelobate pattern. The
|
||
<typeStatus id="54F032D94B1DFFB739CFFA54EEDC1B9F" box="[644,752,1479,1505]" pageId="4" pageNumber="141" type="lectotype">lectotype</typeStatus>
|
||
seems to be immature as it does not show any sign of suture approximation at the end of the phragmocone. The suture morphogenesis of that species has subsequently been examined by
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73901F99EEF0A1856" author="Mikhailova" box="[586,806,1549,1576]" firstAuthor="Mikhailova" pageId="4" pageNumber="141" pagination="173 - 182" refId="ref11748" refString="Mikhailova I. A. 1957. On the systematic position of the families Parahoplitidae Spath and Deshayesitidae Stoyanow. Vestnik Moskovskogo Universiteta, Biologiya 3: 173 - 182." type="journal article" year="1957">Mikhailova (1957</bibRefCitation>
|
||
: fig. 5; 1958: fig. 1) and
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73F29F99EE93A1859" author="Tovbina" box="[1122,1302,1549,1575]" firstAuthor="Tovbina" pageId="4" pageNumber="141" pagination="56 - 65" refId="ref12225" refString="Tovbina S. Z. 1970. A new genus of the family Parahoplitidae. Paleontological Journal 3: 56 - 65." type="journal article" year="1970">Tovbina (1970</bibRefCitation>
|
||
: fig. 4). According to these authors, the umbilical lobe is tripartite and rather symmetrical. A wide umbilical saddle is divided into two parts, the outer of which in turn becomes bifid. The ventral and dorsal lobes are bifid and complicated by lateral teeth. The ventral lobe has a low median saddle.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6F92BECC218AC" blockId="4.[189,238,1720,1746]" box="[189,238,1720,1746]" pageId="4" pageNumber="141">
|
||
<heading id="D0BC3B174B1DFFB73BF6F92BECC218AC" bold="true" box="[189,238,1720,1746]" fontSize="11" level="3" pageId="4" pageNumber="141" reason="3">
|
||
<emphasis id="B93F50694B1DFFB73BF6F92BECC218AC" bold="true" box="[189,238,1720,1746]" pageId="4" pageNumber="141">Age</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1DFFB73BF6F974ED4219AA" blockId="4.[187,1399,1767,2004]" pageId="4" pageNumber="141">
|
||
The litho- and biostratigraphy of the Akusha locality was documented by
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73F2EF974E912197F" author="Mordvilko T. A." box="[1125,1342,1767,1793]" pageId="4" pageNumber="141" refId="ref6896" refString="Mordvilko T. A. 1962. Lower Cretaceous Deposits of the Southeast Northern Caucasus and Ciscaucasia), part 2. Akademii Nauk SSSR, Moscow - Leningrad." type="book" year="1962">Mordvilko (1962)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73BF6F899EE73195A" author="Bogdanova & Mikhailova" box="[189,607,1802,1828]" firstAuthor="Bogdanova" pageId="4" pageNumber="141" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova (2016)</bibRefCitation>
|
||
, but none of these authors have reported the
|
||
<typeStatus id="54F032D94B1DFFB73FFEF898E8C4195B" box="[1205,1256,1803,1829]" pageId="4" pageNumber="141">type</typeStatus>
|
||
stratum of
|
||
<taxonomicName id="4C4BF7F84B1DFFB73BF6F8BEEDC41939" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[189,488,1837,1863]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1DFFB73BF6F8BEEDC41939" box="[189,488,1837,1863]" italics="true" pageId="4" pageNumber="141">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
. The species is, however, known to occur in the ‘
|
||
<taxonomicName id="4C4BF7F84B1DFFB73FDBF8BEED791914" baseAuthorityName="Anthula" baseAuthorityYear="1899" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" isUncertain="true" kingdom="Animalia" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B1DFFB73FDBF8BEE91D1939" box="[1168,1329,1837,1863]" italics="true" pageId="4" pageNumber="141">Acanthoplites</emphasis>
|
||
(sic)
|
||
<emphasis id="B93F50694B1DFFB73BF6F8C3ED791914" box="[189,341,1872,1898]" italics="true" pageId="4" pageNumber="141">aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F50694B1DFFB73A1EF8C2EE611914" box="[341,589,1872,1899]" italics="true" pageId="4" pageNumber="141">
|
||
–
|
||
<taxonomicName id="4C4BF7F84B1DFFB73A28F8C3EE611914" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[355,589,1872,1898]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="uhligi">Acanthoplites uhligi</taxonomicName>
|
||
</emphasis>
|
||
Zone’ sensu Mordviko (1962) of
|
||
<collectingRegion id="498F42994B1DFFB73886F8C2E8161915" box="[973,1082,1873,1899]" country="Russia" name="Dagestan" pageId="4" pageNumber="141">Dagestan</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73F0CF8C3ECD719F0" author="Bogdanova & Mikhailova" firstAuthor="Bogdanova" pageId="4" pageNumber="141" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova 2016</bibRefCitation>
|
||
). Correlation of this zone with the SMAZ is still not obtained with certainty, but it may cover part of the
|
||
<taxonomicName id="4C4BF7F84B1DFFB73A46F804EE0019CF" box="[269,556,1943,1969]" class="Cephalopoda" family="Douvilleiceratidae" genus="Epicheloniceras" higherTaxonomySource="CoL" kingdom="Animalia" order="Ammonitida" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="martini">
|
||
<emphasis id="B93F50694B1DFFB73A46F804EE0019CF" box="[269,556,1943,1969]" italics="true" pageId="4" pageNumber="141">Epicheloniceras martini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4BF7F84B1DFFB73921F804EFA419CF" authorityName="Anthula" authorityYear="1899" box="[618,904,1943,1969]" class="Cephalopoda" family="Deshayesitidae" genus="Parahoplites" kingdom="Animalia" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="melchioris">
|
||
<emphasis id="B93F50694B1DFFB73921F804EFA419CF" box="[618,904,1943,1969]" italics="true" pageId="4" pageNumber="141">Parahoplites melchioris</emphasis>
|
||
</taxonomicName>
|
||
standard zones (
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73F18F804E90719CF" author="Luber" box="[1107,1323,1943,1969]" etAl="et al." firstAuthor="Luber" pageId="4" pageNumber="141" pagination="12 - 34" refId="ref11534" refString="Luber T. L., Bulot L. G., Redfern J., Frau C., Arantegui A. & Masrour M. 2017. A revised ammonoid biostratigraphy for the Aptian of NW Africa: Essaouira-Agadir Basin, Morocco. Cretaceous Research 79: 12 - 34. https: // doi. org / 10.1016 / j. cretres. 2017.06.020." type="journal article" year="2017">
|
||
Luber
|
||
<emphasis id="B93F50694B1DFFB73FEEF80BE8F719CF" box="[1189,1243,1943,1969]" italics="true" pageId="4" pageNumber="141">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73E72F804ED2419AA" author="Frau" firstAuthor="Frau" pageId="4" pageNumber="141" pagination="1 - 16" refId="ref11154" refString="Frau C. 2020 a. New insight on the age and significance of the Aparein black shales in the Basque- Cantabrian Basin, northern Spain. Newsletters on Stratigraphy 54 (1): 1 - 16. https: // doi. org / 10.1127 / nos / 2020 / 0583" type="journal article" year="2020" yearSuffix="a">Frau, 2020a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDAF18A4B1DFFB73A58F829ED4E19AA" author="Frau C." box="[275,354,1978,2004]" firstAuthor="Frau" pageId="4" pageNumber="141" pagination="1 - 16" refId="ref11154" refString="Frau C. 2020 a. New insight on the age and significance of the Aparein black shales in the Basque- Cantabrian Basin, northern Spain. Newsletters on Stratigraphy 54 (1): 1 - 16. https: // doi. org / 10.1127 / nos / 2020 / 0583" type="journal article" year="2020" yearSuffix="b">2020b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |