372 lines
48 KiB
XML
372 lines
48 KiB
XML
<document id="D3DF14511198FF18443A925E0B165993" ID-DOI="10.5281/zenodo.191273" ID-GBIF-Dataset="d2982f3f-a5f5-46ff-bbd8-a1e186245a89" ID-ISSN="1175-5326" ID-Zenodo-Dep="191273" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461125997720" checkinUser="plazi" docAuthor="Smetana, Aleš" docDate="2009" docId="03DFE029FFA6F85EFF1949964CF1FBDC" docLanguage="en" docName="zt02286p030.pdf" docOrigin="Zootaxa 2286" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ocypus (Pseudocypus) shiretokensis Hayashi 2008" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFE69851FFAAF850FF8E4A524F65FF8E" masterDocTitle="Contributions to the knowledge of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part 21. The genus Ocypus Leach, 1819, subgenus Pseudocypus Mulsant & Rey, 1876. Section 4" masterLastPageNumber="30" masterPageNumber="1" pageNumber="13" updateTime="1698596766005" updateUser="plazi">
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<mods:title id="805113D9BEF5071CF20F701244149C28">Contributions to the knowledge of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part 21. The genus Ocypus Leach, 1819, subgenus Pseudocypus Mulsant & Rey, 1876. Section 4</mods:title>
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<mods:namePart id="8ADDDFEC7A5F6B365B52F8B60B7A03F5">Smetana, Aleš</mods:namePart>
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<mods:typeOfResource id="DCD4E36DE3415387423C6CA01DE05755">text</mods:typeOfResource>
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<mods:title id="F5AE627DA844F4853B0A5CEEBCEAC047">Zootaxa</mods:title>
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<mods:part id="08D117BB878591E6FC31DA39FF743B7C">
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<mods:date id="72E44B3F4E83DA0ACEADBC6BF7AB0FF7">2009</mods:date>
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<mods:detail id="C2A0E9EA6C82D036D9E6290128BD9674" type="volume">
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<mods:number id="33A69416455541FD5D7B043FC405F407">2286</mods:number>
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<mods:identifier id="FABD0C1A0ECC0EBAC657F62BF884DF93" type="DOI">10.5281/zenodo.191273</mods:identifier>
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<mods:identifier id="E62384911C011C6776364DD350BBF47B" type="GBIF-Dataset">d2982f3f-a5f5-46ff-bbd8-a1e186245a89</mods:identifier>
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<mods:identifier id="026B4D5B5E236BCFEA065298F511775A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8D723184CDDD527D819B6F2B9E26C645" type="Zenodo-Dep">191273</mods:identifier>
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<treatment id="03DFE029FFA6F85EFF1949964CF1FBDC" ID-DOI="http://doi.org/10.5281/zenodo.6217662" ID-GBIF-Taxon="119626388" ID-Zenodo-Dep="6217662" LSID="urn:lsid:plazi:treatment:03DFE029FFA6F85EFF1949964CF1FBDC" httpUri="http://treatment.plazi.org/id/03DFE029FFA6F85EFF1949964CF1FBDC" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="C36C02B4FFA6F85CFF1949964E51FB8A" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BC9513FFFA6F85CFF1949964C66FC53" blockId="12.[151,771,963,1028]" box="[151,771,963,990]" pageId="12" pageNumber="13">
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<heading id="D081E653FFA6F85CFF1949964C66FC53" bold="true" box="[151,771,963,990]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<taxonomicName id="4C762ABCFFA6F85CFF1949964C66FC53" ID-CoL="48MG3" authority="Hayashi, 2008" authorityName="Hayashi" authorityYear="2008" box="[151,771,963,990]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="shiretokensis" subGenus="Pseudocypus">
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<emphasis id="B9028D2DFFA6F85CFF1949964C66FC53" bold="true" box="[151,771,963,990]" pageId="12" pageNumber="13">
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<emphasis id="B9028D2DFFA6F85CFF1949964F94FC50" bold="true" box="[151,241,964,990]" italics="true" pageId="12" pageNumber="13">Ocypus</emphasis>
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(
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<emphasis id="B9028D2DFFA6F85CFE8F49964EF8FC50" bold="true" box="[257,413,964,990]" italics="true" pageId="12" pageNumber="13">Pseudocypus</emphasis>
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)
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<emphasis id="B9028D2DFFA6F85CFE2149964D28FC50" bold="true" box="[431,589,964,990]" italics="true" pageId="12" pageNumber="13">shiretokensis</emphasis>
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<bibRefCitation id="EFE72CCEFFA6F85CFDDA49914C66FC53" author="Hayashi" box="[596,771,963,989]" pageId="12" pageNumber="28" refString="Hayashi, Y. (2008) A new species of Ocypus (Pseudocypus) from the Shiretoko Peninsula, Hokkaido, Japan (Coleoptera: Staphylinidae: Staphylinina). Entomological Review of Japan, 62, 199 - 204." type="journal article" year="2008">Hayashi, 2008</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BC9513FFFA6F85CFF1949B84E51FB8A" blockId="12.[151,771,963,1028]" box="[151,308,1002,1028]" pageId="12" pageNumber="13">
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(
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<figureCitation id="134D4DBAFFA6F85CFF1149B84E48FB8A" box="[159,301,1002,1028]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="12" pageNumber="13">Figs. 29–35</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C02B4FFA6F85CFF194E6A4C5AFBC1" box="[151,831,1080,1103]" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="8BC9513FFFA6F85CFF194E6A4C5AFBC1" blockId="12.[151,831,1080,1103]" box="[151,831,1080,1103]" pageId="12" pageNumber="13">
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<taxonomicName id="4C762ABCFFA6F85CFF194E6A4EA1FBC1" ID-CoL="48MG3" authority="Hayashi, 2008" authorityName="Hayashi" authorityYear="2008" box="[151,452,1080,1103]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="shiretokensis">
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<emphasis id="B9028D2DFFA6F85CFF194E6A4E44FBC1" box="[151,289,1080,1103]" italics="true" pageId="12" pageNumber="13">shiretokensis</emphasis>
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<bibRefCitation id="EFE72CCEFFA6F85CFEA64E6A4EA1FBC1" author="Hayashi" box="[296,452,1080,1103]" pageId="12" pageNumber="28" refString="Hayashi, Y. (2008) A new species of Ocypus (Pseudocypus) from the Shiretoko Peninsula, Hokkaido, Japan (Coleoptera: Staphylinidae: Staphylinina). Entomological Review of Japan, 62, 199 - 204." type="journal article" year="2008">Hayashi, 2008</bibRefCitation>
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</taxonomicName>
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, 199 (
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<taxonomicName id="4C762ABCFFA6F85CFD884E6A4D33FBC1" ID-CoL="8VRWY" box="[518,598,1080,1103]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B9028D2DFFA6F85CFD884E6A4D33FBC1" box="[518,598,1080,1103]" italics="true" pageId="12" pageNumber="13">Ocypus</emphasis>
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</taxonomicName>
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; subg.
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<taxonomicName id="4C762ABCFFA6F85CFD2A4E6A4C4AFBC1" box="[676,815,1080,1103]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B9028D2DFFA6F85CFD2A4E6A4C4AFBC1" box="[676,815,1080,1103]" italics="true" pageId="12" pageNumber="13">Pseudocypus</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C02B4FFA6F85CFF194E2C4E39FA5F" pageId="12" pageNumber="13" type="materials_examined">
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<paragraph id="8BC9513FFFA6F85CFF194E2C4D9DFB68" blockId="12.[151,1437,1150,2030]" pageId="12" pageNumber="13">
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<emphasis id="B9028D2DFFA6F85CFF194E2C4E28FB16" bold="true" box="[151,333,1150,1176]" pageId="12" pageNumber="13">
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<typeStatus id="54CDEF9DFFA6F85CFF194E2C4FB6FB16" box="[151,211,1150,1176]" pageId="12" pageNumber="13">Type</typeStatus>
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material.
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</emphasis>
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<bibRefCitation id="EFE72CCEFFA6F85CFED64E2C4D2BFB16" author="Hayashi" box="[344,590,1150,1176]" pageId="12" pageNumber="28" refString="Hayashi, Y. (2008) A new species of Ocypus (Pseudocypus) from the Shiretoko Peninsula, Hokkaido, Japan (Coleoptera: Staphylinidae: Staphylinina). Entomological Review of Japan, 62, 199 - 204." type="journal article" year="2008">Hayashi (2008: 199)</bibRefCitation>
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described the species from three male specimens from Hokkaido. The
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<typeStatus id="54CDEF9DFFA6F85CFF194EF74F9BFB31" box="[151,254,1189,1215]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
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with the label “Shiretoko pass (Alt.
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<quantity id="4C8EFCDAFFA6F85CFD214EF74D98FB31" box="[687,765,1189,1215]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="13" unit="m" value="730.0">730 m</quantity>
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), Hokkaido,
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<date id="FFC877FFFFA6F85CFC194EF74B7AFB31" box="[919,1055,1189,1215]" pageId="12" pageNumber="13" value="1989-07-29">29.vii.1989</date>
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, H. Nomura leg.”is deposited in the collection of Osaka Museum of Natural History.
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</paragraph>
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<paragraph id="8BC9513FFFA6F85CFF484EA04E41FAD4" blockId="12.[151,1437,1150,2030]" pageId="12" pageNumber="13">
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I have not seen the
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<typeStatus id="54CDEF9DFFA6F85CFE224EA04EBAFA82" box="[428,479,1266,1292]" pageId="12" pageNumber="13">type</typeStatus>
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material, but I have sent one male specimen of the series dealt with in this paper to Hayashi, who compared it with the
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<typeStatus id="54CDEF9DFFA6F85CFDF74F4B4D83FABD" box="[633,742,1305,1331]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C762ABCFFA6F85CFC9C4F4B4CB9FABC" box="[786,988,1305,1331]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="shiretokensis">
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<emphasis id="B9028D2DFFA6F85CFC9C4F4B4C42FABC" box="[786,807,1305,1330]" italics="true" pageId="12" pageNumber="13">O</emphasis>
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.
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<emphasis id="B9028D2DFFA6F85CFCB74F4B4CB9FABC" box="[825,988,1305,1330]" italics="true" pageId="12" pageNumber="13">shiretokensis</emphasis>
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</taxonomicName>
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and confirmed that the specimen is conspecific.
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</paragraph>
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<paragraph id="8BC9513FFFA6F85CFF484F344E39FA5F" blockId="12.[151,1437,1150,2030]" pageId="12" pageNumber="13">
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<emphasis id="B9028D2DFFA6F85CFF484F344D68FA0E" bold="true" box="[198,525,1382,1408]" pageId="12" pageNumber="13">
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Material studied.
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<collectingCountry id="F36111AFFFA6F85CFE2B4F344D6DFA0E" box="[421,520,1382,1408]" name="Japan" pageId="12" pageNumber="13">JAPAN</collectingCountry>
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:
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</emphasis>
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”(Mt. Kurodake) Mts. Daisedtsuzan, Hokkaido,
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<collectingCountry id="F36111AFFFA6F85CFBC34F344BF6FA0E" box="[1101,1171,1382,1408]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
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, VII.21.1981, Coll. N. Yasuda” (4) (
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<collectionCode id="ED67C9FAFFA6F85CFEBD4FC24E15FA24" box="[307,368,1424,1450]" pageId="12" pageNumber="13">ASC</collectionCode>
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, YSC); ”Mt. Kitoushi,
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<quantity id="4C8EFCDAFFA6F85CFD0C4FC24DBBFA24" box="[642,734,1424,1450]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="m" value="1000.0">1000 m</quantity>
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, Rikubetsu, Hokkaido,
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<date id="FFC877FFFFA6F85CFC7D4FC24BC5FA24" box="[1011,1184,1421,1450]" pageId="12" pageNumber="13" value="1993-06-26">June 26th 1993</date>
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, Kôichi Shibata leg.” (3) (
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<collectionCode id="ED67C9FAFFA6F85CFF474FE54E63FA5F" box="[201,262,1463,1489]" pageId="12" pageNumber="13">ASC</collectionCode>
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, YSC).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C02B4FFA6F85CFF484F8F4C86F9CA" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="8BC9513FFFA6F85CFF484F8F4C86F9CA" blockId="12.[151,1437,1150,2030]" pageId="12" pageNumber="13">
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<emphasis id="B9028D2DFFA6F85CFF484F8F4E23FA79" bold="true" box="[198,326,1501,1527]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
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Medium-sized species, sharing most character states with
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<taxonomicName id="4C762ABCFFA6F85CFB9A4F8C4BBFFA79" box="[1044,1242,1501,1527]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inexspectatus">
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<emphasis id="B9028D2DFFA6F85CFB9A4F8C4B4CFA79" box="[1044,1065,1502,1527]" italics="true" pageId="12" pageNumber="13">O</emphasis>
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.
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<emphasis id="B9028D2DFFA6F85CFBB24F8C4BBFFA79" box="[1084,1242,1502,1527]" italics="true" pageId="12" pageNumber="13">inexspectatus</emphasis>
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</taxonomicName>
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, but distinct by base of second abdominal tergite lacking punctation and pubescence, fifth visible abdominal tergite lacking pale apical seem of palisade setae, and by differently shaped aedoeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C02B4FFA6F85EFF484C034CF1FBDC" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BC9513FFFA6F85CFF484C034BE2F888" blockId="12.[151,1437,1150,2030]" pageId="12" pageNumber="13">
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||
<emphasis id="B9028D2DFFA6F85CFF484C034E3AF9E5" bold="true" box="[198,351,1617,1643]" pageId="12" pageNumber="13">Description.</emphasis>
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In all character states similar to
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<taxonomicName id="4C762ABCFFA6F85CFD724C034CA7F9E4" box="[764,962,1617,1643]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inexspectatus">
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<emphasis id="B9028D2DFFA6F85CFD724C034C74F9E4" box="[764,785,1617,1642]" italics="true" pageId="12" pageNumber="13">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA6F85CFCAA4C034CA7F9E4" box="[804,962,1617,1642]" italics="true" pageId="12" pageNumber="13">inexspectatus</emphasis>
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</taxonomicName>
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, but different as follows: average size smaller, body form more slender; pronotum slightly narrower (ratio length / width = 1.12; as long as wide in
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<taxonomicName id="4C762ABCFFA6F85CFF194CCD4E06F936" box="[151,355,1695,1721]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inexspectatus">
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<emphasis id="B9028D2DFFA6F85CFF194CCD4FC9F936" box="[151,172,1695,1720]" italics="true" pageId="12" pageNumber="13">O</emphasis>
|
||
.
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||
<emphasis id="B9028D2DFFA6F85CFF334CCD4E06F936" box="[189,355,1695,1720]" italics="true" pageId="12" pageNumber="13">inexspectatus</emphasis>
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</taxonomicName>
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); punctation of elytra more distinctly granulose and coarser; base of second abdominal tergite (in front of first visible tergit) impunctate and glabrous; fifth visible abdominal tergite lacking pale apical seem of palisade setae; punctation of abdominal tergites coarser and less dense.
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</paragraph>
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<paragraph id="8BC9513FFFA6F85CFF484D404AF9F860" blockId="12.[151,1437,1150,2030]" pageId="12" pageNumber="13">
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Male. Area behind anterior angles of pronotum densely punctate. Sternite 7 with medioapical emargination similar to that of
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<taxonomicName id="4C762ABCFFA6F85CFD894D6B4DAFF8DC" box="[519,714,1849,1875]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inexspectatus">
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||
<emphasis id="B9028D2DFFA6F85CFD894D6B4D79F8DC" box="[519,540,1849,1874]" italics="true" pageId="12" pageNumber="13">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA6F85CFDA24D6B4DAFF8DC" box="[556,714,1849,1874]" italics="true" pageId="12" pageNumber="13">inexspectatus</emphasis>
|
||
</taxonomicName>
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, but narrower and less deep. Genital segment with tergite 10 rather short and wide, evenly narrowed toward subtruncate apex, setose as in
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<figureCitation id="134D4DBAFFA6F85CFBCF4D324BFFF8F4" box="[1089,1178,1888,1914]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
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; sternite 9 with acute basal portion, apical portion with subacute apex, setose as in
|
||
<figureCitation id="134D4DBAFFA6F85CFCE84DD54CDBF82F" box="[870,958,1927,1953]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="12" pageNumber="13">Fig. 30</figureCitation>
|
||
. Aedoeagus (
|
||
<figureCitation id="134D4DBAFFA6F85CFBD54DD54B8DF82F" box="[1115,1256,1927,1953]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="12" pageNumber="13">Figs. 31–34</figureCitation>
|
||
) similar to that of
|
||
<taxonomicName id="4C762ABCFFA6F85CFF364DFC4E1AF849" box="[184,383,1965,1991]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inexspectatus">
|
||
<emphasis id="B9028D2DFFA6F85CFF364DFC4FA8F849" box="[184,205,1966,1991]" italics="true" pageId="12" pageNumber="13">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA6F85CFF534DFC4E1AF849" box="[221,383,1966,1991]" italics="true" pageId="12" pageNumber="13">inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
, but right lateral portion of median lobe in ventral view less conspicuously dilated and apex of median lobe shorter. Paramere situated on median lobe quite asymmetrically, of similar shape (
|
||
<figureCitation id="134D4DBAFFA6F85CFAE24D864AF9F860" box="[1388,1436,2004,2030]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="12" pageNumber="13">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC9513FFFA7F85DFF194ACA4FD9FF71" blockId="13.[151,1437,152,410]" pageId="13" pageNumber="14">
|
||
33) but narrower, especially in basal portion, and more straight, with acute apex almost reaching apex of median lobe; underside of paramere lacking sensory peg setae, but with fine sensory setae situated as in
|
||
<figureCitation id="134D4DBAFFA7F85DFAE34AEC4FDDFF71" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="13" pageNumber="14">Fig. 34</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC9513FFFA7F85DFF484B5E4EFAFEFD" blockId="13.[151,1437,152,410]" pageId="13" pageNumber="14">
|
||
Female. Area behind anterior angles of pronotum sparingly punctate on ground with very fine submeshed microsculpture. Tergite 10 of genital segment rather wide, evenly narrowed toward broadly arcuate apex (
|
||
<figureCitation id="134D4DBAFFA7F85DFEB44B0B4EF7FEFD" box="[314,402,345,371]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="13" pageNumber="14">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9513FFFA7F85DFF484BD24D6BFE14" blockId="13.[151,1437,152,410]" box="[198,526,384,410]" pageId="13" pageNumber="14">Body length 11.0–14 0 mm.</paragraph>
|
||
<caption id="DF0901B7FFA7F85DFF194C904E64F8CF" httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="13" pageNumber="14" targetBox="[159,1420,476,1687]" targetPageId="13">
|
||
<paragraph id="8BC9513FFFA7F85DFF194C904E64F8CF" blockId="13.[151,1437,1730,1857]" pageId="13" pageNumber="14">
|
||
<emphasis id="B9028D2DFFA7F85DFF194C904E00F954" bold="true" box="[151,357,1730,1754]" pageId="13" pageNumber="14">FIGURES 29–35.</emphasis>
|
||
Details of
|
||
<emphasis id="B9028D2DFFA7F85DFE6C4C904DB0F957" box="[482,725,1730,1753]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C762ABCFFA7F85DFE6C4C904DB4F957" box="[482,721,1730,1753]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="shiretokensis">Ocypus shiretokensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
29—tergite 10 of male genital segment; 30—sternite 9 of male genital segment (1.5 mm); 31—aedoeagus, ventral view (1.6 mm); 32—aedoeagus, lateral view (1.6 mm); 33— underside of paramere (1.1 mm); 34—apical portion of underside of paramere; 35—tergite 10 of female genital segment (1.1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9513FFFA7F85EFF484D204C66FF56" blockId="13.[151,1437,1906,2010]" lastBlockId="14.[151,1437,152,1106]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<emphasis id="B9028D2DFFA7F85DFF484D204D4CF802" bold="true" box="[198,553,1906,1932]" pageId="13" pageNumber="14">Recognition and comments.</emphasis>
|
||
<taxonomicName id="4C762ABCFFA7F85DFDBB4D204C55F805" box="[565,816,1906,1931]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA7F85DFDBB4D204C55F805" box="[565,816,1906,1931]" italics="true" pageId="13" pageNumber="14">Ocypus shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
is in general habitus and most external characters (coloration and pubescence of the body, coloration of the appendages) most similar to
|
||
<taxonomicName id="4C762ABCFFA7F85DFB164DCB4A3EF83C" box="[1176,1371,1945,1971]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inexspectatus">
|
||
<emphasis id="B9028D2DFFA7F85DFB164DCB4BC8F83C" box="[1176,1197,1945,1970]" italics="true" pageId="13" pageNumber="14">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA7F85DFB334DCB4A3EF83C" box="[1213,1371,1945,1970]" italics="true" pageId="13" pageNumber="14">inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
, as it has been shown above. However,
|
||
<taxonomicName id="4C762ABCFFA7F85DFDA04D924D88F857" box="[558,749,1984,2009]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA7F85DFDA04D924D88F857" box="[558,749,1984,2009]" italics="true" pageId="13" pageNumber="14">O. shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
differs at once, in addition to the different aedoeagus, by the base of the second abdominal tergite lacking punctation and pubescence, and the fifth visible abdominal tergite lacking the pale apical seem of palisade setae.
|
||
</paragraph>
|
||
<paragraph id="8BC9513FFFA4F85EFF484AB74B24FEA8" blockId="14.[151,1437,152,1106]" pageId="14" pageNumber="15">
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFF484AB74EDCFF71" author="Hayashi" box="[198,441,229,255]" pageId="14" pageNumber="28" refString="Hayashi, Y. (2008) A new species of Ocypus (Pseudocypus) from the Shiretoko Peninsula, Hokkaido, Japan (Coleoptera: Staphylinidae: Staphylinina). Entomological Review of Japan, 62, 199 - 204." type="journal article" year="2008">Hayashi (2008, 199)</bibRefCitation>
|
||
stated that
|
||
<taxonomicName id="4C762ABCFFA4F85EFDCA4AB74C64FF70" box="[580,769,229,255]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA4F85EFDCA4AB74D3CFF70" box="[580,601,229,254]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFDE74AB74C64FF70" box="[617,769,229,254]" italics="true" pageId="14" pageNumber="15">shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
is very similar in general appearance to
|
||
<taxonomicName id="4C762ABCFFA4F85EFB694AB74AF9FF70" box="[1255,1436,229,255]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigroaeneus">
|
||
<emphasis id="B9028D2DFFA4F85EFB694AB74B99FF70" box="[1255,1276,229,254]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFA824AB74AF9FF70" box="[1292,1436,229,254]" italics="true" pageId="14" pageNumber="15">nigroaeneus</emphasis>
|
||
</taxonomicName>
|
||
and at the same time postulated close relationship of the species with
|
||
<taxonomicName id="4C762ABCFFA4F85EFC484B5E4B5FFEAB" box="[966,1082,268,294]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="helleni">
|
||
<emphasis id="B9028D2DFFA4F85EFC484B5E4CBEFEAB" box="[966,987,268,293]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFC644B5E4B5FFEAB" box="[1002,1082,268,293]" italics="true" pageId="14" pageNumber="15">helleni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC9513FFFA4F85EFF484B604BE1FDD5" blockId="14.[151,1437,152,1106]" pageId="14" pageNumber="15">
|
||
The similarity of
|
||
<taxonomicName id="4C762ABCFFA4F85EFE1B4B604D34FEC5" box="[405,593,306,332]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA4F85EFE1B4B604ECFFEC5" box="[405,426,306,331]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFE374B604D34FEC5" box="[441,593,306,331]" italics="true" pageId="14" pageNumber="15">shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C762ABCFFA4F85EFD184B604C2FFEC5" box="[662,842,306,332]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigroaeneus">
|
||
<emphasis id="B9028D2DFFA4F85EFD184B604DCEFEC5" box="[662,683,306,331]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFD344B604C2FFEC5" box="[698,842,306,331]" italics="true" pageId="14" pageNumber="15">nigroaeneus</emphasis>
|
||
</taxonomicName>
|
||
is in fact limited only to the fact that both species belong to the same lineage within
|
||
<taxonomicName id="4C762ABCFFA4F85EFDDF4B0B4D90FEFC" box="[593,757,345,370]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFDDF4B0B4D90FEFC" box="[593,757,345,370]" italics="true" pageId="14" pageNumber="15">Pseudocypus</emphasis>
|
||
</taxonomicName>
|
||
and share the character states of this lineage. But
|
||
<taxonomicName id="4C762ABCFFA4F85EFAF04B0B4E42FE17" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigroaeneus">
|
||
<emphasis id="B9028D2DFFA4F85EFAF04B0B4AF6FEFC" box="[1406,1427,345,370]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFF194BD24E42FE17" box="[151,295,384,409]" italics="true" pageId="14" pageNumber="15">nigroaeneus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C762ABCFFA4F85EFE4E4BD24D19FE17" box="[448,636,384,410]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA4F85EFE4E4BD24EB0FE17" box="[448,469,384,409]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFE6A4BD24D19FE17" box="[484,636,384,409]" italics="true" pageId="14" pageNumber="15">shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
, in addition to the differently shaped aedoeagus and tergite 10 of the female genital segment (
|
||
<figureCitation id="134D4DBAFFA4F85EFE344BF44D78FE4E" box="[442,541,422,448]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="14" pageNumber="15">Figs. 31</figureCitation>
|
||
,
|
||
<figureCitation id="134D4DBAFFA4F85EFDA44BF44D2FFE4E" box="[554,586,422,448]" captionStart="FIGURES 50 – 56" captionStartId="20.[151,271,1807,1831]" captionTargetBox="[151,1434,537,1773]" captionTargetId="figure@20.[151,1436,537,1783]" captionTargetPageId="20" captionText="FIGURES 50 – 56. Details of Ocypus nigroaeneus. 50 — tergite 10 of male genital segment (1.25 mm, left side shows proper shape, right side slightly damaged and unnaturally explanate); 51 — sternite 9 of male genital segment (1.6 mm); 52 — aedoeagus, ventral view (1.8 mm); 53 — aedoeagus, lateral view (1.9 mm); 54 — underside of paramere (1.4 mm); 55 — apical portion of underside of paramere; 56 — tergite 10 of female genital segment (1.45 mm)." httpUri="https://zenodo.org/record/191281/files/figure.png" pageId="14" pageNumber="15">52</figureCitation>
|
||
,
|
||
<figureCitation id="134D4DBAFFA4F85EFDD84BF44D13FE4E" box="[598,630,422,448]" captionStart="FIGURES 29 – 35" captionStartId="13.[151,273,1730,1754]" captionTargetBox="[159,1420,476,1687]" captionTargetId="figure@13.[151,1436,461,1712]" captionTargetPageId="13" captionText="FIGURES 29 – 35. Details of Ocypus shiretokensis. 29 — tergite 10 of male genital segment; 30 — sternite 9 of male genital segment (1.5 mm); 31 — aedoeagus, ventral view (1.6 mm); 32 — aedoeagus, lateral view (1.6 mm); 33 — underside of paramere (1.1 mm); 34 — apical portion of underside of paramere; 35 — tergite 10 of female genital segment (1.1 mm)." httpUri="https://zenodo.org/record/191278/files/figure.png" pageId="14" pageNumber="15">35</figureCitation>
|
||
,
|
||
<figureCitation id="134D4DBAFFA4F85EFD0D4BF44DC5FE4E" box="[643,672,422,448]" captionStart="FIGURES 50 – 56" captionStartId="20.[151,271,1807,1831]" captionTargetBox="[151,1434,537,1773]" captionTargetId="figure@20.[151,1436,537,1783]" captionTargetPageId="20" captionText="FIGURES 50 – 56. Details of Ocypus nigroaeneus. 50 — tergite 10 of male genital segment (1.25 mm, left side shows proper shape, right side slightly damaged and unnaturally explanate); 51 — sternite 9 of male genital segment (1.6 mm); 52 — aedoeagus, ventral view (1.8 mm); 53 — aedoeagus, lateral view (1.9 mm); 54 — underside of paramere (1.4 mm); 55 — apical portion of underside of paramere; 56 — tergite 10 of female genital segment (1.45 mm)." httpUri="https://zenodo.org/record/191281/files/figure.png" pageId="14" pageNumber="15">56</figureCitation>
|
||
), by several important external characters, such as the presence of a variably large impunctate area on the pronotum on each side of the impunctate midline, and the presence of the pale apical seam of palisade setae on the fifth visible abdominal tergite; additional different character states are the dark metallic surface of the fore body, the distinctly double punctation of the dorsal surface of the head and the pronotum, and the darker color of the appendages in
|
||
<taxonomicName id="4C762ABCFFA4F85EFC4748134B18FDD4" box="[969,1149,577,603]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigroaeneus">
|
||
<emphasis id="B9028D2DFFA4F85EFC4748134CBBFDD4" box="[969,990,577,602]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFC6348134B18FDD4" box="[1005,1149,577,602]" italics="true" pageId="14" pageNumber="15">nigroaeneus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC9513FFFA4F85EFF48483A4CF1FBDC" blockId="14.[151,1437,152,1106]" pageId="14" pageNumber="15">
|
||
The postulated close relationship of
|
||
<taxonomicName id="4C762ABCFFA4F85EFD02483A4C2BFD0F" box="[652,846,616,642]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA4F85EFD02483A4DC4FD0F" box="[652,673,616,641]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFD38483A4C2BFD0F" box="[694,846,616,641]" italics="true" pageId="14" pageNumber="15">shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C762ABCFFA4F85EFC12483A4B73FD0F" box="[924,1046,616,642]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="helleni">
|
||
<emphasis id="B9028D2DFFA4F85EFC12483A4CD4FD0F" box="[924,945,616,641]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFC48483A4B73FD0F" box="[966,1046,616,641]" italics="true" pageId="14" pageNumber="15">helleni</emphasis>
|
||
</taxonomicName>
|
||
, based on “similar structure of dentition of mandibles”, is misleading. The presence or absence of mandibular teeth, in addition to some other character states, such as the presence of subbasal indentation in members of the genera
|
||
<taxonomicName id="4C762ABCFFA4F85EFB3B48E74E87FD78" authority="Muller, 1925 (Smetana 2003: 130)" authorityName="Muller, 1925 (Smetana" authorityPageNumber="130" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFB3B48E74A5EFD40" box="[1205,1339,693,718]" italics="true" pageId="14" pageNumber="15">Protocypus</emphasis>
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFACA48E74FB1FD78" author="Muller" pageId="14" pageNumber="29" refString="Muller, J. (1925) Terzo contributo alla conoscenza del genere Staphylinus L. Bolletino della Societa Entomologica Italiana, 57, 40 - 48." type="journal article" year="1925">Müller, 1925</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFF6B488E4EBCFD78" author="Smetana" box="[229,473,732,758]" pageId="14" pageNumber="29" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the " Staphylinus - complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Callocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 135." type="journal article" year="2003">Smetana 2003: 130</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C762ABCFFA4F85EFE7E488E4B05FD78" authority="Smetana, 2006 (Smetana 2006: 40)" authorityName="Smetana, 2006 (Smetana" authorityPageNumber="40" authorityYear="2006" box="[496,1120,732,758]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFE7E488E4DC7FD7B" box="[496,674,732,757]" italics="true" pageId="14" pageNumber="15">Nelmanwaslus</emphasis>
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFD25488E4C04FD78" author="Smetana" box="[683,865,732,758]" pageId="14" pageNumber="30" refString="Smetana, A. (2006) Contributions to the knowledge of the genera of the " Staphylinus - complex (Coleoptera: Staphylinidae) of China. Part 11. The genus Wasmanellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316, 33 - 43." type="journal article" year="2006">Smetana, 2006</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFCFC488E4B3DFD78" author="Smetana" box="[882,1112,732,758]" pageId="14" pageNumber="30" refString="Smetana, A. (2006) Contributions to the knowledge of the genera of the " Staphylinus - complex (Coleoptera: Staphylinidae) of China. Part 11. The genus Wasmanellus Bernhauer, 1920 and Nelmanwaslus gen. nov. Zootaxa, 1316, 33 - 43." type="journal article" year="2006">Smetana 2006: 40</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, or
|
||
<taxonomicName id="4C762ABCFFA4F85EFB1E488E4EE6FC92" authority="Sharp, 1889 (Smetana 2003: 61)" authorityName="Sharp, 1889 (Smetana" authorityPageNumber="61" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Agelosus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFB1E488E4B9BFD7B" box="[1168,1278,732,757]" italics="true" pageId="14" pageNumber="15">Agelosus</emphasis>
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFA89488E4AF9FD78" author="Sharp" box="[1287,1436,732,758]" pageId="14" pageNumber="29" refString="Sharp, D. S. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, (6) 3, 28 - 44, 108 - 121, 249 - 267, 319 - 334, 406 - 419, 463 - 476." type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFF1149504E1CFC92" author="Smetana" box="[159,377,770,796]" pageId="14" pageNumber="29" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the " Staphylinus - complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Callocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 135." type="journal article" year="2003">Smetana 2003: 61</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, belong to the character states important at the generic level for distinguishing genera of the ‘Staphylinus-complex” (
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFE7C497B4C42FCCD" author="Smetana" box="[498,807,809,835]" pageId="14" pageNumber="30" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88." type="journal article" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
). But at the species level, the shape and number of mandibular teeth varies among members of many genus level taxa, and
|
||
<taxonomicName id="4C762ABCFFA4F85EFB9849024BD3FCE7" box="[1046,1206,848,873]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFB9849024BD3FCE7" box="[1046,1206,848,873]" italics="true" pageId="14" pageNumber="15">Pseudocypus</emphasis>
|
||
</taxonomicName>
|
||
is one of them. In addition, the development of mandibular teeth in some members of
|
||
<taxonomicName id="4C762ABCFFA4F85EFC3749254B37FC1E" box="[953,1106,887,912]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFC3749254B37FC1E" box="[953,1106,887,912]" italics="true" pageId="14" pageNumber="15">Pseudocypus</emphasis>
|
||
</taxonomicName>
|
||
is sexually dimorphic (e.g., in
|
||
<taxonomicName id="4C762ABCFFA4F85EFF3649CF4E38FC38" box="[184,349,925,951]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="picipennis">
|
||
<emphasis id="B9028D2DFFA4F85EFF3649CF4FA8FC38" box="[184,205,925,950]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFF5049CF4E38FC38" box="[222,349,925,950]" italics="true" pageId="14" pageNumber="15">picipennis</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFE72CCEFFA4F85EFE1149CF4D31FC39" author="Smetana" box="[415,596,925,951]" pageId="14" pageNumber="29" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the " Staphylinus - complex (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Callocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 135." type="journal article" year="2003">Smetana 2003</bibRefCitation>
|
||
: 62,
|
||
<figureCitation id="134D4DBAFFA4F85EFD1E49CF4C6CFC39" box="[656,777,925,951]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,271,1672,1696]" captionTargetBox="[151,1407,421,1645]" captionTargetId="figure@3.[151,1436,421,1649]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Details of Ocypus picipennis. 1 — tergite 10 of male genital segment (1.7 mm); 2 — sternite 9 of male genital segment (2.45 mm); 3 — aedoeagus, ventral view (2.6 mm); 4 — aedoeagus, lateral view (2.6 mm); 5 — underside of paramere (2.0 mm); 6 — apical portion of underside of paramere; 7 — tergite 10 of female genital segment (1.75 mm)." httpUri="https://zenodo.org/record/191274/files/figure.png" pageId="14" pageNumber="15">Figs. 1–4</figureCitation>
|
||
). It is therefore obvious that any attempt to establish “relationships” among the members of this subgenus, based solely on the development of mandicular teeth, must lead to wrong results. The “relationship” of
|
||
<taxonomicName id="4C762ABCFFA4F85EFD5649B84CF1FB8D" box="[728,916,1002,1028]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA4F85EFD5649B84D88FB8D" box="[728,749,1002,1027]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFD7249B84CF1FB8D" box="[764,916,1002,1027]" italics="true" pageId="14" pageNumber="15">shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C762ABCFFA4F85EFC4249B84B25FB8D" box="[972,1088,1002,1028]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="helleni">
|
||
<emphasis id="B9028D2DFFA4F85EFC4249B84C84FB8D" box="[972,993,1002,1027]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFC7E49B84B25FB8D" box="[1008,1088,1002,1027]" italics="true" pageId="14" pageNumber="15">helleni</emphasis>
|
||
</taxonomicName>
|
||
ends at the level that both are members of the subgenus
|
||
<taxonomicName id="4C762ABCFFA4F85EFE5D4E434D0AFBA4" box="[467,623,1041,1066]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9028D2DFFA4F85EFE5D4E434D0AFBA4" box="[467,623,1041,1066]" italics="true" pageId="14" pageNumber="15">Pseudocypus</emphasis>
|
||
</taxonomicName>
|
||
, but they each belong to a different lineage:
|
||
<taxonomicName id="4C762ABCFFA4F85EFB004E434A35FBA4" box="[1166,1360,1041,1067]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="shiretokensis">
|
||
<emphasis id="B9028D2DFFA4F85EFB004E434BC6FBA4" box="[1166,1187,1041,1066]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFB3A4E434A35FBA4" box="[1204,1360,1041,1066]" italics="true" pageId="14" pageNumber="15">shiretokensis</emphasis>
|
||
</taxonomicName>
|
||
to the
|
||
<emphasis id="B9028D2DFFA4F85EFF194E6A4E64FBDF" box="[151,257,1080,1105]" italics="true" pageId="14" pageNumber="15">fuscatus-</emphasis>
|
||
lineage,
|
||
<taxonomicName id="4C762ABCFFA4F85EFEEA4E6A4EBDFBDF" box="[356,472,1080,1106]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="helleni">
|
||
<emphasis id="B9028D2DFFA4F85EFEEA4E6A4E1CFBDF" box="[356,377,1080,1105]" italics="true" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<emphasis id="B9028D2DFFA4F85EFE064E6A4EBDFBDF" box="[392,472,1080,1105]" italics="true" pageId="14" pageNumber="15">helleni</emphasis>
|
||
</taxonomicName>
|
||
to the
|
||
<taxonomicName id="4C762ABCFFA4F85EFDA64E6A4DC5FBDF" box="[552,672,1080,1105]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="picipennis">
|
||
<emphasis id="B9028D2DFFA4F85EFDA64E6A4DC5FBDF" box="[552,672,1080,1105]" italics="true" pageId="14" pageNumber="15">picipennis</emphasis>
|
||
</taxonomicName>
|
||
-lineage (see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |