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<document id="42FCC0AE58E048AEFF33917C13453DD5" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58539FF8D5991FE6B510CFB74" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) trisignatum Harold 1881" docType="treatment" docVersion="10" lastPageNumber="463" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="462" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="C8CE04669C2A9485C53FE7B95CBCEB6F">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="97C61803C99BE8A4FD6F4B6BB7559554">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="EA8F0D7B709AD9C07192101341C447CB">Louzada, Júlio</mods:namePart>
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<mods:namePart id="9D059836ED40F26171DF6A12842A5178">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:title id="1A8430B2F40A002929D39A353C4BE4F9">Zootaxa</mods:title>
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<mods:date id="449D790B4F9D06085D55288BA3F31882">2015</mods:date>
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<taxonomicName id="4C774D408539FF8A5991FE6B5650FE78" ID-CoL="34KB5" authority="Harold 1881" authorityName="Harold" authorityYear="1881" box="[151,799,402,428]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum" subGenus="Aganhyboma">
<emphasis id="B903EAD18539FF8A5991FE6B5650FE78" bold="true" box="[151,799,402,428]" pageId="11" pageNumber="462">
<emphasis id="B903EAD18539FF8A5991FE6B5462FE78" bold="true" box="[151,301,402,428]" italics="true" pageId="11" pageNumber="462">Deltochilum</emphasis>
(
<emphasis id="B903EAD18539FF8A583BFE6B5491FE78" bold="true" box="[317,478,402,428]" italics="true" pageId="11" pageNumber="462">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18539FF8A58F6FE6B5732FE78" bold="true" box="[496,637,402,428]" italics="true" pageId="11" pageNumber="462">trisignatum</emphasis>
<bibRefCitation id="EFE64B328539FF8A5B85FE6B5650FE78" author="Harold" box="[643,799,402,428]" pageId="11" pageNumber="462" refString="Harold, E. V. (1881) Einige neue coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 149 - 171." type="journal article" year="1881">Harold 1881</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BC836C38539FF8A5991FE4C5770FE1A" blockId="11.[151,799,402,462]" box="[151,575,437,462]" pageId="11" pageNumber="462">
(
<figureCitation id="134C2A468539FF8A5999FE4C55A8FE1A" box="[159,231,437,462]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="11" pageNumber="462">Figs 9</figureCitation>
, 27, 35, 44, 52, 66, 69, 89, 95)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488539FF8A5991FE04574FFCB0" pageId="11" pageNumber="462" type="reference_group">
<paragraph id="8BC836C38539FF8A5991FE04548FFDE6" blockId="11.[151,1305,509,868]" pageId="11" pageNumber="462">
<treatmentCitationGroup id="AB6711ED8539FF8A5991FE04548FFDE6" pageId="11" pageNumber="462">
<taxonomicName id="4C774D408539FF8A5991FE04575AFDC7" ID-CoL="34KB5" authority="Harold 1881" authorityName="Harold" authorityYear="1881" box="[151,533,509,531]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum">
<emphasis id="B903EAD18539FF8A5991FE0454C0FDC7" box="[151,399,509,531]" italics="true" pageId="11" pageNumber="462">Deltochilum trisignatum</emphasis>
<bibRefCitation id="EFE64B328539FF8A5893FE04575AFDC7" author="Harold" box="[405,533,509,531]" pageId="11" pageNumber="462" refString="Harold, E. V. (1881) Einige neue coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 149 - 171." type="journal article" year="1881">Harold 1881</bibRefCitation>
</taxonomicName>
[1880]: 150 (original description)
<treatmentCitation id="0AD610D28539FF8A5991FDE45477FDE6" author="Heyne" box="[151,312,540,562]" page="61" pageId="11" pageNumber="462" year="1900">
<bibRefCitation id="EFE64B328539FF8A5991FDE45459FDE6" author="Heyne" box="[151,278,540,562]" pageId="11" pageNumber="462" refString="Heyne, A. (1900) pars 9: Scarabaeidae (Lamellicornia): Coprini. In: Heyne, A. &amp; Taschenberg, O. (Eds.), 1893 - 1902, Die Exotischen Kafer in Wort und Bild. J. F. Schreiber, Esslingen &amp; Munchen, pp. 61." type="journal article" year="1900">Heyne 1900</bibRefCitation>
: 61
</treatmentCitation>
(description)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C38539FF8A5991FDC354E8FD84" blockId="11.[151,1305,509,868]" box="[151,423,570,592]" pageId="11" pageNumber="462">
<bibRefCitation id="EFE64B328539FF8A5991FDC35443FD84" author="Gillet" box="[151,268,570,592]" pageId="11" pageNumber="462" refString="Gillet, J. J. E. (1911) Scarabaeidae: Coprinae I. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. W Junk, Berlin, pp. 1 - 100." type="book chapter" year="1911">Gillet 1911</bibRefCitation>
: 36 (catalogue)
</paragraph>
<paragraph id="8BC836C38539FF8A5991FDA354BBFD5A" blockId="11.[151,1305,509,868]" pageId="11" pageNumber="462">
<bibRefCitation id="EFE64B328539FF8A5991FDA35423FDBB" author="Pessoa" box="[151,364,601,623]" pageId="11" pageNumber="462" refString="Pessoa, S. B. &amp; Lane, F. (1941) Coleopteros necrofagos de interesse medico-legal. Ensaio monografico sobre a familia Scarabaeidae de Sao Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo, 2, 389 - 504." type="journal article" year="1941">Pessôa &amp; Lane 1941</bibRefCitation>
: 428, 431 and 432 (description and identification key)
<bibRefCitation id="EFE64B328539FF8A5991FD81541CFD5A" author="Blackwelder" box="[151,339,632,654]" pageId="11" pageNumber="462" refString="Blackwelder, R. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 265." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 203 (checklist)
</paragraph>
<paragraph id="8BC836C38539FF8A5991FD6F5471FD1F" blockId="11.[151,1305,509,868]" pageId="11" pageNumber="462">
<treatmentCitationGroup id="AB6711ED8539FF8A5991FD6F5471FD1F" pageId="11" pageNumber="462">
<taxonomicName id="4C774D408539FF8A5991FD6F5471FD1F" ID-CoL="34KB5" authority="Kolbe 1893: 192 Shipp 1897: 195" authorityName="Kolbe 1893: 192 Shipp" authorityPageNumber="195" authorityYear="1897" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum" subGenus="Aganhyboma">
<emphasis id="B903EAD18539FF8A5991FD6F545BFD78" box="[151,276,662,684]" italics="true" pageId="11" pageNumber="462">Deltochilum</emphasis>
(
<emphasis id="B903EAD18539FF8A5825FD6F54E6FD78" box="[291,425,662,684]" italics="true" pageId="11" pageNumber="462">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18539FF8A58B1FD615763FD79" box="[439,556,664,685]" italics="true" pageId="11" pageNumber="462">trisignatum</emphasis>
<treatmentCitation id="0AD610D28539FF8A5B34FD6F5793FD78" author="Kolbe" box="[562,732,662,684]" page="192" pageId="11" pageNumber="462" year="1893">
<bibRefCitation id="EFE64B328539FF8A5B34FD6F57E2FD78" author="Kolbe" box="[562,685,662,684]" pageId="11" pageNumber="462" refString="Kolbe, H. J. (1893) Beitrage zur kenntniss der mistkafer, Lamellicornia onthophila. Stettiner entomologische Zeitung, 54, 188 - 202." type="journal article" year="1893">Kolbe 1893</bibRefCitation>
: 192
</treatmentCitation>
<treatmentCitation id="0AD610D28539FF8A5991FD4C5471FD1F" author="Shipp" box="[151,318,693,715]" page="195" pageId="11" pageNumber="462" year="1897">
<bibRefCitation id="EFE64B328539FF8A5991FD4C5441FD1F" author="Shipp" box="[151,270,693,715]" pageId="11" pageNumber="462" refString="Shipp, J. W. (1897) On the synonymy of the genus Deltochilum. Entomologische Nachrichten, 23, 194 - 196." type="journal article" year="1897">Shipp 1897</bibRefCitation>
: 195
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C38539FF8A5991FD2D5716FCDC" blockId="11.[151,1305,509,868]" pageId="11" pageNumber="462">
<bibRefCitation id="EFE64B328539FF8A5991FD2D5450FD3E" author="Paulian" box="[151,287,724,746]" pageId="11" pageNumber="462" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
: 240, 252, 253, 256,
<figureCitation id="134C2A468539FF8A58F5FD2D577CFD3E" box="[499,563,724,746]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1892,1915]" captionTargetBox="[168,1431,864,1864]" captionTargetId="figure@3.[151,1436,855,1871]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Male genital sclerites in Deltochilum (Aganhyboma). 1. Position and nomenclature of endophallic sclerites (modified from Medina et al. 2013). 2. Nomenclature of the main parts of the genital segment (D. amandaarcanjoae new species). 3. Nomenclature of the main parts of the superior right peripheral sclerite (SRP) (D. paresi new species). Figures scale = 1 mm." httpUri="https://zenodo.org/record/244796/files/figure.png" pageId="11" pageNumber="462">Figs 2</figureCitation>
,
<figureCitation id="134C2A468539FF8A5B39FD2D5715FD3E" box="[575,602,724,746]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="11" pageNumber="462">18</figureCitation>
, map No 2 (description, taxonomic comments and identification key)
<bibRefCitation id="EFE64B328539FF8A5991FD0A5449FCDC" author="Lane" box="[151,262,754,776]" pageId="11" pageNumber="462" refString="Lane, F. (1946) Sobre os tipos e a sinonimia de alguns Canthonini (Col. Scarabaeidae). Papeis Avulsos do Departamento de Zoologia, 7, 171 - 179." type="journal article" year="1946">Lane 1946</bibRefCitation>
: 173, 174 (taxonomic comments)
</paragraph>
<paragraph id="8BC836C38539FF8A5991FCE85488FCF3" blockId="11.[151,1305,509,868]" box="[151,455,785,807]" pageId="11" pageNumber="462">
<bibRefCitation id="EFE64B328539FF8A5991FCE854D8FCF3" author="Pereira" box="[151,407,785,807]" pageId="11" pageNumber="462" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956</bibRefCitation>
: 121
</paragraph>
<paragraph id="8BC836C38539FF8A5991FCC95778FC92" blockId="11.[151,1305,509,868]" box="[151,567,816,838]" pageId="11" pageNumber="462">
<bibRefCitation id="EFE64B328539FF8A5991FCC954C0FC92" author="Vulcano" box="[151,399,816,838]" pageId="11" pageNumber="462" refString="Vulcano, M. A. &amp; Pereira, F. S. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the western hemisphere. Entomologischen Arbeiten aus dem Museum G. Frey, 15, 569 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964</bibRefCitation>
: 642 (catalogue)
</paragraph>
<paragraph id="8BC836C38539FF8A5991FCB7574FFCB0" blockId="11.[151,1305,509,868]" box="[151,512,846,868]" pageId="11" pageNumber="462">Vaz-de-Mello 2000: 192 (checklist)</paragraph>
</subSubSection>
<subSubSection id="C36D65488539FF8A5991FC6854E8F932" pageId="11" pageNumber="462" type="description">
<paragraph id="8BC836C38539FF8A5991FC6854E8F932" blockId="11.[151,1437,913,2018]" pageId="11" pageNumber="462">
<emphasis id="B903EAD18539FF8A5991FC68543CFC7E" bold="true" box="[151,371,913,938]" pageId="11" pageNumber="462">Description. Body.</emphasis>
Head black with violet hue, pronotum yellowish brown with black spots, elytra reddish brown (
<figureCitation id="134C2A468539FF8A5999FC4C55B9FC1A" box="[159,246,949,974]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="11" pageNumber="462">Fig. 95</figureCitation>
). Ventral surface black with green reflections. Length
<quantity id="4C8F9B268539FF8A5A75FC4F5140FC19" box="[883,1039,950,974]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="11" pageNumber="462" unit="mm" value="18.0" valueMax="19.0" valueMin="17.0">17 to 19 mm</quantity>
.
<emphasis id="B903EAD18539FF8A5D1BFC4C512BFC1A" bold="true" box="[1053,1124,949,974]" pageId="11" pageNumber="462">Head.</emphasis>
Dorsal surface completely covered by closely spaced to confluent foveiform punctures resulting in rugosity, anterior punctures smaller than those at center. Punctures margin well-defined (impressed). Margin between clypeal teeth and genae curved outward near clypeogenal suture.
<emphasis id="B903EAD18539FF8A5B15FBD8573CFBEE" bold="true" box="[531,627,1057,1082]" pageId="11" pageNumber="462">Thorax.</emphasis>
Surface of pronotum with deep and dense foveiform punctures forming rugosity. Anteromedial punctures much smaller than posteromedial punctures. Margin of pronotum (close to lateral angles), curved inward, lacking emargination. Anterior margin with three black spots, one covering each angle and one at center. Pronotum with small black spot close to lateral angles and dark spot in posterior part of disc. Margin between lateral and posterior angles distinctly convex (curved outward).
<emphasis id="B903EAD18539FF8A5AFEFB485103FB1E" bold="true" box="[1016,1100,1201,1226]" pageId="11" pageNumber="462">Elytra.</emphasis>
Striae weakly marked, with almost inconspicuous punctures. Lateral margins evenly curved outward (
<figureCitation id="134C2A468539FF8A5AE5FB2C5176FB3A" box="[995,1081,1237,1262]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="11" pageNumber="462">Fig. 95</figureCitation>
). Interstrial surface with weak, medium-size foveiform punctures intermingled with dense microtubercles, some of which are confluent, shining and forming a glossy zone. First interstria brighter than rest of elytra. Basal carina of seventh interstria weak and short (
<figureCitation id="134C2A468539FF8A59D9FAB8547CFA8E" box="[223,307,1345,1370]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="11" pageNumber="462">Fig. 95</figureCitation>
), approximately the same length as metatibial spur. Length of apical carina of seventh interstria 1.5 times that of apex of sixth interstria.
<emphasis id="B903EAD18539FF8A5B41FA9C578CFAAA" bold="true" box="[583,707,1381,1406]" pageId="11" pageNumber="462">Abdomen.</emphasis>
Punctures large and conspicuous, concentrated on anterior and posterior margins of sternites and reduced to a narrow belt along midline of abdomen. Pygidium almost completely covered by nearly confluent, large, foveiform punctures, leaving only a short, smooth medial line from base to about midlength of pygidial surface. Apex of pygidium rounded.
<emphasis id="B903EAD18539FF8A5A99FA285691FA3E" bold="true" box="[927,990,1489,1514]" pageId="11" pageNumber="462">Legs.</emphasis>
Protibia sharply widened along inner margin from basal third to apex (Fig. 66). Middle protibial tooth closer to apical tooth than to basal tooth (Fig. 66). Apical third of metatibia strongly curved inward (Fig. 67). Inner margin of metatibia with a longitudinal row of small tubercles. Apex of metatibia with six clearly visible longitudinal carinae. Anterior margin of profemur effaced at middle portion (Fig. 69). Most foveiform punctures of the profemur elliptical. Metatarsus approximately one-third as long as metatibia.
<emphasis id="B903EAD18539FF8A58EBF97C5607F94A" bold="true" box="[493,840,1669,1694]" pageId="11" pageNumber="462">Secondary sexual characters.</emphasis>
<collectingCountry id="F36076538539FF8A5A49F97C56C7F94A" box="[847,904,1669,1694]" name="Malta" pageId="11" pageNumber="462">Male</collectingCountry>
abdomen slightly more longitudinally flattened than that of female. Apical margin of pygidium more dilated in male.
<collectingCountry id="F36076538539FF8A5AAFF95056ADF916" box="[937,994,1705,1730]" name="Malta" pageId="11" pageNumber="462">Male</collectingCountry>
protibial spur rounded at apex. Female spur acuminate apically.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488539FF8D59C1F9085771FC88" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462" type="materials_examined">
<paragraph id="8BC836C38539FF8A59C1F908578FF836" blockId="11.[151,1437,913,2018]" pageId="11" pageNumber="462">
<emphasis id="B903EAD18539FF8A59C1F908576FF8DE" bold="true" box="[199,544,1777,1802]" pageId="11" pageNumber="462">
Material studied.
<typeStatus id="54CC88618539FF8A589CF908575FF8DE" box="[410,528,1777,1802]" pageId="11" pageNumber="462" type="lectotype">Lectotype</typeStatus>
(
</emphasis>
<collectionCode id="ED66AE068539FF8A5B26F90B5738F8DE" box="[544,631,1778,1802]" pageId="11" pageNumber="462">MNHU</collectionCode>
)
<emphasis id="B903EAD18539FF8A5B83F908561CF8DE" bold="true" box="[645,851,1777,1802]" pageId="11" pageNumber="462">[here designated]</emphasis>
♂: 1. [old white label, handwritten]. 8915; 2. [green label, handwritten].
<taxonomicName id="4C774D408539FF8A5881F8EC5783F8FA" authority="Harold Brasilia" authorityName="Harold Brasilia" box="[391,716,1813,1838]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum">
trisignatum Harold
<collectingCountry id="F36076538539FF8A5B77F8EC5783F8FA" box="[625,716,1813,1838]" name="Brazil" pageId="11" pageNumber="462">Brasilia</collectingCountry>
</taxonomicName>
; 3. [red label, bordered and printed in black, handwritten].
<typeStatus id="54CC88618539FF8A5991F8C35448F886" box="[151,263,1850,1874]" pageId="11" pageNumber="462" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C774D408539FF8A580BF8C0573BF886" authority="Harold" authorityName="Harold" box="[269,628,1849,1874]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum">Deltochilum trisignatum Harold</taxonomicName>
des. F.Z. Vaz-de-Mello, 2014.
<emphasis id="B903EAD18539FF8A5ACAF8C05133F886" bold="true" box="[972,1148,1849,1874]" pageId="11" pageNumber="462">
<typeStatus id="54CC88618539FF8A5ACAF8C05133F886" box="[972,1148,1849,1874]" pageId="11" pageNumber="462" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
(
<collectionCode id="ED66AE068539FF8A5D8CF8C351ACF886" box="[1162,1251,1850,1874]" pageId="11" pageNumber="462">MNHU</collectionCode>
)
<emphasis id="B903EAD18539FF8A5DF4F8C0504FF886" bold="true" box="[1266,1280,1849,1874]" pageId="11" pageNumber="462">2</emphasis>
♀♀: ♀ 1. [old white label, handwritten]. 8915; 2. [green label, handwritten].
<collectingCountry id="F36076538539FF8A5A65F8A456F1F8A2" box="[867,958,1885,1910]" name="Brazil" pageId="11" pageNumber="462">Brasilia</collectingCountry>
; 3. [yellow label, bordered and printed in black, handwritten].
<typeStatus id="54CC88618539FF8A5886F8785757F84E" box="[384,536,1921,1946]" pageId="11" pageNumber="462" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C774D408539FF8A5B26F87856C6F84E" authority="Harold" authorityName="Harold" box="[544,905,1921,1946]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum">Deltochilum trisignatum Harold</taxonomicName>
des. F.Z. Vaz-de-Mello, 2014./ ♀ 1. [old white label, handwritten]. 8915; 2. [yellow label, bordered and printed in black, handwritten].
<typeStatus id="54CC88618539FF8A5D77F85C5046F86A" box="[1137,1289,1957,1982]" pageId="11" pageNumber="462" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C774D408539FF8A5C16F85C5424F836" authority="Harold" authorityName="Harold" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="trisignatum">Deltochilum trisignatum Harold</taxonomicName>
des. F.Z. Vaz-de-Mello, 2014.
</paragraph>
<paragraph id="8BC836C3853EFF8D59C1FF6E5725FF2C" blockId="12.[151,1437,151,1184]" pageId="12" pageNumber="463">
<emphasis id="B903EAD1853EFF8D59C1FF6E54DFFF64" bold="true" box="[199,400,151,176]" pageId="12" pageNumber="463">Taxonomic notes</emphasis>
. A
<typeStatus id="54CC8861853EFF8D58BCFF6E576EFF64" box="[442,545,151,176]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
is here designated in order to maintain nomenclatural stability by choosing the only male (presenting characters that are easier to use for distinguishing from other species) among the available
<typeStatus id="54CC8861853EFF8D5991FF1955B6FF2C" box="[151,249,224,248]" pageId="12" pageNumber="463" type="syntype">syntypes</typeStatus>
as the single name-bearing
<typeStatus id="54CC8861853EFF8D5B34FF195729FF2C" box="[562,614,224,248]" pageId="12" pageNumber="463">type</typeStatus>
.
</paragraph>
<paragraph id="8BC836C3853EFF8D59C1FEFD57A7FDE8" blockId="12.[151,1437,151,1184]" pageId="12" pageNumber="463">
<emphasis id="B903EAD1853EFF8D59C1FEFD57E8FEC9" bold="true" box="[199,679,260,285]" pageId="12" pageNumber="463">
Non-type material [30]:
<collectingCountry id="F3607653853EFF8D58FAFEFD57E8FEC9" box="[508,679,260,285]" name="Argentina" pageId="12" pageNumber="463">ARGENTINA</collectingCountry>
</emphasis>
: TUCUMAN [probably mislabelled] (1 ♀
<collectionCode id="ED66AE06853EFF8D5DB5FEFD5041FEC8" LSID="urn:lsid:biocol.org:col:34988" box="[1203,1294,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="463">MNHN</collectionCode>
);
<emphasis id="B903EAD1853EFF8D5C21FEFD50D4FEC9" bold="true" box="[1319,1435,260,285]" pageId="12" pageNumber="463">
<collectingCountry id="F3607653853EFF8D5C21FEFD50D9FEC9" box="[1319,1430,260,285]" name="Brazil" pageId="12" pageNumber="463">BRAZIL</collectingCountry>
:
</emphasis>
<collectionCode id="ED66AE06853EFF8D5991FED155BBFE94" box="[151,244,296,320]" pageId="12" pageNumber="463">BAHIA</collectionCode>
: Santa Teresinha, Pedra Branca, (
<geoCoordinate id="EE435004853EFF8D5B86FEDE57B1FE94" box="[640,766,295,320]" direction="south" orientation="latitude" pageId="12" pageNumber="463" precision="9" value="-12.8555">12°51.33'S</geoCoordinate>
,
<geoCoordinate id="EE435004853EFF8D5A08FEDE56E9FE94" box="[782,934,295,320]" direction="west" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="-39.47625">39°28.575'W</geoCoordinate>
),
<date id="FFC91003853EFF8D5ABBFED15169FE94" box="[957,1062,296,320]" pageId="12" pageNumber="463" value="2009-02-02">2.II.2009</date>
, L.R.M. Oliveira (3 MZUEFS); ESPÍRITO
<collectionCode id="ED66AE06853EFF8D581BFEB55431FEB0" box="[285,382,332,356]" pageId="12" pageNumber="463">SANTO</collectionCode>
: P. Cachoeira, (2 ♀
<collectionCode id="ED66AE06853EFF8D5B70FEB55797FEB0" box="[630,728,332,356]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="463">MZUSP</collectionCode>
); Conceição da Barra,
<date id="FFC91003853EFF8D5AECFEB551C3FEB1" box="[1002,1164,332,357]" pageId="12" pageNumber="463" value="1968-10-22" valueMax="1968-10-28" valueMin="1968-10-22">22–28.X.1968</date>
, C. &amp; C.T. Elias (1 ♀
<collectionCode id="ED66AE06853EFF8D5991FE8955AAFE5C" box="[151,229,368,392]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="12" pageNumber="463">DZUP</collectionCode>
); Linhares,
<date id="FFC91003853EFF8D586DFE9654F3FE5C" box="[363,444,367,392]" pageId="12" pageNumber="463" value="1964-02">II.1964</date>
(
<specimenCount id="9D71FD4A853EFF8D58D7FE8954B5FE53" box="[465,506,368,392]" pageId="12" pageNumber="463" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE06853EFF8D5B05FE89571EFE5C" box="[515,593,368,392]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="12" pageNumber="463">DZUP</collectionCode>
); same,
<date id="FFC91003853EFF8D5BA9FE96565AFE5C" box="[687,789,367,392]" pageId="12" pageNumber="463" value="1967-12">XII.1967</date>
(1 ♀
<collectionCode id="ED66AE06853EFF8D5A5DFE8956E6FE5C" box="[859,937,368,392]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="12" pageNumber="463">DZUP</collectionCode>
); Linhares, Lagoa do Macuco, (
<geoCoordinate id="EE435004853EFF8D5C13FE9650D8FE5C" box="[1301,1431,367,392]" direction="south" orientation="latitude" pageId="12" pageNumber="463" precision="15" value="-19.063889">19°03'50&quot;S</geoCoordinate>
,
<geoCoordinate id="EE435004853EFF8D5991FE6D546BFE79" box="[151,292,404,429]" direction="west" orientation="longitude" pageId="12" pageNumber="463" precision="15" value="-39.97861">39°58'43&quot;W</geoCoordinate>
),
<date id="FFC91003853EFF8D583BFE6D549BFE79" box="[317,468,404,429]" pageId="12" pageNumber="463" value="2000-01-26" valueMax="2000-01-27" valueMin="2000-01-26">26–27.I.2000</date>
, F. Génier &amp; S. Ide (
<specimenCount id="9D71FD4A853EFF8D5BDCFE6D5649FE78" box="[730,774,404,428]" pageId="12" pageNumber="463" type="generic">1 ♂</specimenCount>
3 ♀
<collectionCode id="ED66AE06853EFF8D5A4AFE6D56D4FE78" box="[844,923,404,428]" pageId="12" pageNumber="463">CEMT</collectionCode>
, 2 ♀
<collectionCode id="ED66AE06853EFF8D5AE2FE6D5166FE78" LSID="urn:lsid:biocol.org:col:34935" box="[996,1065,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="12" pageNumber="463">CMN</collectionCode>
); Linhares, Povoação, (
<geoCoordinate id="EE435004853EFF8D5C3BFE6D50D8FE79" box="[1341,1431,404,429]" direction="south" orientation="latitude" pageId="12" pageNumber="463" precision="925" value="-19.6">19°36'S</geoCoordinate>
,
<geoCoordinate id="EE435004853EFF8D5991FE4155B2FE04" box="[151,253,440,464]" direction="west" orientation="longitude" pageId="12" pageNumber="463" precision="925" value="-39.8">39°48'W</geoCoordinate>
),
<date id="FFC91003853EFF8D5817FE4154C0FE04" box="[273,399,440,464]" pageId="12" pageNumber="463" value="2000-12-03">3.XII.2000</date>
, G. Schiffler (
<specimenCount id="9D71FD4A853EFF8D5B2BFE415700FE04" box="[557,591,440,464]" pageId="12" pageNumber="463" type="generic">1♂</specimenCount>
<collectionCode id="ED66AE06853EFF8D5B50FE4157E6FE04" box="[598,681,440,464]" pageId="12" pageNumber="463">CEMT</collectionCode>
);
<collectionCode id="ED66AE06853EFF8D5BBBFE415658FE04" box="[701,791,440,464]" pageId="12" pageNumber="463">MINAS</collectionCode>
GERAIS: Mar de Hespanha,
<date id="FFC91003853EFF8D5D6EFE4151AAFE04" box="[1128,1253,440,464]" pageId="12" pageNumber="463" value="1910-12-09">9.XII.1910</date>
, J.F. Zikán (1 ♀
<collectionCode id="ED66AE06853EFF8D5991FE2555A9FE20" box="[151,230,476,500]" pageId="12" pageNumber="463">CEMT</collectionCode>
,
<specimenCount id="9D71FD4A853EFF8D59F7FE255455FE20" box="[241,282,476,500]" pageId="12" pageNumber="463" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE06853EFF8D5824FE2554CBFE20" box="[290,388,476,500]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="463">MZUSP</collectionCode>
); Marliéria, Parque Estadual do Rio Doce, (
<geoCoordinate id="EE435004853EFF8D5A68FE25515BFE20" box="[878,1044,476,500]" direction="south" orientation="latitude" pageId="12" pageNumber="463" precision="1" value="-19.702562">19°42'09.22&quot;S</geoCoordinate>
,
<geoCoordinate id="EE435004853EFF8D5D19FE255180FE21" box="[1055,1231,476,501]" direction="west" orientation="longitude" pageId="12" pageNumber="463" precision="1" value="-40.515057">40°30'54.21&quot;W</geoCoordinate>
),
<date id="FFC91003853EFF8D5DE5FE25501FFE21" box="[1251,1360,476,501]" pageId="12" pageNumber="463" value="2005-01-11">11.I.2005</date>
, S.S.P. Almeida &amp; M.R. Pereira (
<specimenCount id="9D71FD4A853EFF8D58BFFDF954BEFDC3" box="[441,497,512,536]" pageId="12" pageNumber="463" type="generic">10 ♂</specimenCount>
1 ♀
<collectionCode id="ED66AE06853EFF8D5B2EFDF95734FDCC" box="[552,635,512,536]" pageId="12" pageNumber="463">CEMT</collectionCode>
); RIO
<collectingCountry id="F3607653853EFF8D5BC3FDF957A5FDC3" box="[709,746,512,535]" name="South Georgia and the South Sandwich Islands" pageId="12" pageNumber="463">DE</collectingCountry>
JANEIRO: Sto. Antonio dos Brotos, Districto de San Fidelis,
<date id="FFC91003853EFF8D5991FDDD5457FDE9" box="[151,280,548,573]" pageId="12" pageNumber="463" value="1876" valueMax="1882">1876–1882</date>
, Auguste Vincent de Lyon (
<specimenCount id="9D71FD4A853EFF8D5B54FDDD5734FDE8" box="[594,635,548,572]" pageId="12" pageNumber="463" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE06853EFF8D5B87FDDD5793FDE8" LSID="urn:lsid:biocol.org:col:34988" box="[641,732,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="463">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BC836C3853EFF8D59C1FDBE5428FD19" blockId="12.[151,1437,151,1184]" pageId="12" pageNumber="463">
<emphasis id="B903EAD1853EFF8D59C1FDBE54E3FDB4" bold="true" box="[199,428,583,608]" pageId="12" pageNumber="463">Sampling methods.</emphasis>
Millipede carrion baited pitfall trap [
<specimenCount id="9D71FD4A853EFF8D5A54FDB156ABFDB4" box="[850,996,583,608]" pageId="12" pageNumber="463" type="generic">3 specimens</specimenCount>
]; Flight interception trap [1]; un-baited pitfall trap [11]. The un-baited pitfall traps contained formalin solution (as a preservative) may act as an attractant for beetles predacious on millipedes, since millipedes often release defence secretions containing formaldehydebased compounds.
</paragraph>
<paragraph id="8BC836C3853EFF8D59C1FD2E5771FC88" blockId="12.[151,1437,151,1184]" pageId="12" pageNumber="463">
<emphasis id="B903EAD1853EFF8D59C1FD2E5465FD24" bold="true" box="[199,298,727,752]" pageId="12" pageNumber="463">Habitat.</emphasis>
<collectingCountry id="F3607653853EFF8D5834FD2E5439FD24" box="[306,374,727,752]" name="Brazil" pageId="12" pageNumber="463">Brazil</collectingCountry>
(southern Bahia, Espirito Santo and Minas Gerais) (
<figureCitation id="134C2A46853EFF8D5ACBFD2E516EFD24" box="[973,1057,727,752]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="12" pageNumber="463">Fig. 89</figureCitation>
), from available data this species inhabits lowland Atlantic forest, with sandy soil. A single specimen was collected from “cabruca” vegetation (agricultural area where the cacao is planted in the midst of the natural vegetation of Atlantic forest). The known elevation range is
<quantity id="4C8F9B26853EFF8D5865FCBD54A5FC88" box="[355,490,836,860]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="13.0" metricValueMin="1.0" pageId="12" pageNumber="463" unit="m" value="70.0" valueMax="130.0" valueMin="10.0">10 to 130 m</quantity>
<emphasis id="B903EAD1853EFF8D58F7FCBD5778FC8F" box="[497,567,836,859]" italics="true" pageId="12" pageNumber="463">
<collectionCode id="ED66AE06853EFF8D58F7FCBD5778FC8F" box="[497,567,836,859]" pageId="12" pageNumber="463">AMSL</collectionCode>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6548853EFF8D59C1FC9E510CFB74" pageId="12" pageNumber="463" type="diagnosis">
<paragraph id="8BC836C3853EFF8D59C1FC9E510CFB74" blockId="12.[151,1437,151,1184]" pageId="12" pageNumber="463">
<emphasis id="B903EAD1853EFF8D59C1FC9E5491FC54" bold="true" box="[199,478,871,896]" pageId="12" pageNumber="463">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D40853EFF8D5B6AFC9157A2FCAB" box="[620,749,872,895]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="trisignatum">
<emphasis id="B903EAD1853EFF8D5B6AFC9157A2FCAB" box="[620,749,872,895]" italics="true" pageId="12" pageNumber="463">trisignatum</emphasis>
</taxonomicName>
complex): head black with violet hue, pronotum yellowishbrown with three distinct black spots, elytra reddishbrown; dorsal surface of head completely covered by closely spaced to confluent foveiform punctures resulting in rugosity; margin of pronotum (close to lateral angles) curved inward. The following aedeagus characters are also diagnostic: parameres shorter than phallobase, rounded at apex, in lateral view forming an angle of approximately 30° with phallobase (
<figureCitation id="134C2A46853EFF8D5AE7FC0E5166FBC4" box="[993,1065,1015,1040]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="12" pageNumber="463">Fig. 9</figureCitation>
). Lamella copulatrix: right lobe &quot;scaphopode&quot; shaped (Fig. 35). Superior right peripheral sclerite (SRP) as in Fig. 44. Fronto-lateral peripheral sclerite (FLP) as in Fig. 52. Center of genital segment with a well-sclerotized portion and &quot;M&quot;-shaped (Fig. 27). There is some variation in the pronotum of four specimens examined, which display yellow coloration. The dark spot on posterior portion of pronotal disc can be difficult to see in some individuals.
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