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<document id="77CC5C0031657EA6B6002B4C98EBD633" ID-DOI="10.11646/zootaxa.3688.1.1" ID-GBIF-Dataset="151481a0-71e1-4402-ab7e-8262caddf539" ID-ISSN="1175-5326" ID-Zenodo-Dep="284201" ID-ZooBank="DB84C679-C2AA-41CC-816E-83E2F6DCA391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460649331738" checkinUser="plazi" docAuthor="Haddad, Charles R." docDate="2013" docId="03DB8445FFB6FF9FFF2AF9A93023FAE7" docLanguage="en" docName="zt03688p079.pdf" docOrigin="Zootaxa 3688 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Copuetta litipo Haddad, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="46" masterDocId="FFE2FC3DFF9AFFB2FFBDFFE6345DFFF0" masterDocTitle="Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae" masterLastPageNumber="79" masterPageNumber="1" pageNumber="45" updateTime="1698337697679" updateUser="plazi">
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<mods:title id="24942A95036B72D0503FBF3404044FF0">Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae</mods:title>
</mods:titleInfo>
<mods:name id="4687362FCF17D147A2EB33C916ED9A9D" type="personal">
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<mods:namePart id="A8CCC76CA9C20B7A357F379408D45BD9">Haddad, Charles R.</mods:namePart>
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<mods:title id="AB808A1183885EE95C8C8E2AA8932F2F">Zootaxa</mods:title>
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<mods:part id="2E36157501979D19A3FD9E073F0F955E">
<mods:date id="61D1AEF86D3474ADA47B57AD1091358E">2013</mods:date>
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<mods:number id="418060973814F21A274AC805692D502E">3688</mods:number>
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<mods:classification id="24B266082A902AD5AD915794CB2D97FC">journal article</mods:classification>
<mods:identifier id="B64D78223CC0EC90C8F9F835EACC03DB" type="DOI">10.11646/zootaxa.3688.1.1</mods:identifier>
<mods:identifier id="95A3E4124182870B02D7D709A1566B98" type="GBIF-Dataset">151481a0-71e1-4402-ab7e-8262caddf539</mods:identifier>
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<treatment id="03DB8445FFB6FF9FFF2AF9A93023FAE7" ID-DOI="http://doi.org/10.5281/zenodo.6153945" ID-GBIF-Taxon="127664914" ID-Zenodo-Dep="6153945" LSID="urn:lsid:plazi:treatment:03DB8445FFB6FF9FFF2AF9A93023FAE7" httpUri="http://treatment.plazi.org/id/03DB8445FFB6FF9FFF2AF9A93023FAE7" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="C36866D8FFB6FF9EFF2AF9A93507F97A" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BCD3553FFB6FF9EFF2AF9A935F3F999" blockId="44.[151,430,1615,1674]" box="[151,430,1615,1641]" pageId="44" pageNumber="45">
<heading id="D085823FFFB6FF9EFF2AF9A935F3F999" bold="true" box="[151,430,1615,1641]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="B906E941FFB6FF9EFF2AF9A935F3F999" bold="true" box="[151,430,1615,1641]" pageId="44" pageNumber="45">
<taxonomicName id="4C724ED0FFB6FF9EFF2AF9A93514F999" ID-CoL="Y8YH" box="[151,329,1615,1641]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="litipo" status="sp. nov.">
<emphasis id="B906E941FFB6FF9EFF2AF9A93514F999" bold="true" box="[151,329,1615,1641]" italics="true" pageId="44" pageNumber="45">Copuetta litipo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235543AFFB6FF9EFEEDF9A935F3F999" box="[336,430,1615,1641]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD3553FFB6FF9EFF2AF9943507F97A" blockId="44.[151,430,1615,1674]" box="[151,346,1650,1674]" pageId="44" pageNumber="45">
<figureCitation id="134929D6FFB6FF9EFF2AF99434B1F97A" box="[151,236,1650,1674]" captionStart="FIGURES 45 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="44" pageNumber="45">Figs 51</figureCitation>
,
<figureCitation id="134929D6FFB6FF9EFF45F9943507F97A" box="[248,346,1650,1674]" captionStart="FIGURES 128 131" captionStartId="44.[151,264,1209,1232]" captionTargetBox="[175,1425,220,1167]" captionTargetId="figure@44.[151,1436,195,1186]" captionTargetPageId="44" captionText="FIGURES 128 131. Genitalic morphology of Copuetta lesnei sp. nov. (128, 129) and C. litipo sp. nov. (130, 131): 128. Male palp (mirror image of right palp), ventral view. 129. Same, retrolateral view. 130. Female epigyne, ventral view. 131. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284222/files/figure.png" pageId="44" pageNumber="45">130, 131</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB6FF9EFF2AF95D378EF922" box="[151,979,1722,1747]" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8BCD3553FFB6FF9EFF2AF95D378EF922" blockId="44.[151,1436,1722,2034]" box="[151,979,1722,1747]" pageId="44" pageNumber="45">
<emphasis id="B906E941FFB6FF9EFF2AF95D357CF923" bold="true" box="[151,289,1723,1747]" pageId="44" pageNumber="45">Etymology:</emphasis>
The species name is a noun in apposition of the
<typeStatus id="54C98BF1FFB6FF9EFCFFF95D372FF923" box="[834,882,1723,1747]" pageId="44" pageNumber="45">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB6FF9EFF7AF938315FF8CE" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8BCD3553FFB6FF9EFF7AF938315FF8CE" blockId="44.[151,1436,1722,2034]" pageId="44" pageNumber="45">
<emphasis id="B906E941FFB6FF9EFF7AF938351EF907" bold="true" box="[199,323,1758,1783]" pageId="44" pageNumber="45">Diagnosis:</emphasis>
Females of this species can be easily recognised by the epigyne with oval sclerotised rims laterally with the oval copulatory openings situated within them (
<figureCitation id="134929D6FFB6FF9EFCB0F8E43732F8EA" box="[781,879,1794,1818]" captionStart="FIGURES 128 131" captionStartId="44.[151,264,1209,1232]" captionTargetBox="[175,1425,220,1167]" captionTargetId="figure@44.[151,1436,195,1186]" captionTargetPageId="44" captionText="FIGURES 128 131. Genitalic morphology of Copuetta lesnei sp. nov. (128, 129) and C. litipo sp. nov. (130, 131): 128. Male palp (mirror image of right palp), ventral view. 129. Same, retrolateral view. 130. Female epigyne, ventral view. 131. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284222/files/figure.png" pageId="44" pageNumber="45">Fig. 130</figureCitation>
). This differs from the female of
<taxonomicName id="4C724ED0FFB6FF9EFB5FF8E53168F8EA" box="[1250,1333,1795,1818]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="naja" status="sp. nov.">
<emphasis id="B906E941FFB6FF9EFB5FF8E53168F8EA" box="[1250,1333,1795,1818]" italics="true" pageId="44" pageNumber="45">C. naja</emphasis>
</taxonomicName>
<emphasis id="B906E941FFB6FF9EFA81F8E531C8F8EB" bold="true" box="[1340,1429,1795,1819]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="A235543AFFB6FF9EFA81F8E531C8F8EB" box="[1340,1429,1795,1819]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which has inverted comma-shaped sclerotised rims incorporating the copulatory openings (
<figureCitation id="134929D6FFB6FF9EFB29F8C030ABF8CE" box="[1172,1270,1830,1854]" captionStart="FIGURES 145 148" captionStartId="55.[151,264,1734,1757]" captionTargetBox="[154,1416,802,1708]" captionTargetId="figure@55.[151,1436,792,1717]" captionTargetPageId="55" captionText="FIGURES 145 148. Genitalic morphology of Copuetta naja sp. nov.: 145. Male palp, ventral view. 146. Same, retrolateral view. 147. Female epigyne, ventral view. 148. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284227/files/figure.png" pageId="44" pageNumber="45">Fig. 147</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB6FF9EFF7AF8AD3042F876" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8BCD3553FFB6FF9EFF7AF8AD3042F876" blockId="44.[151,1436,1722,2034]" pageId="44" pageNumber="45">
<emphasis id="B906E941FFB6FF9EFF7AF8AD356FF893" bold="true" box="[199,306,1867,1891]" pageId="44" pageNumber="45">Remark:</emphasis>
The female
<typeStatus id="54C98BF1FFB6FF9EFE70F8AC3672F892" box="[461,559,1866,1890]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
is in rather poor condition and five of its legs are missing. However, the epigyne structure is very distinctive and warrants description of the species here.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB6FF9FFF7AF87535D7FB93" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="description">
<paragraph id="8BCD3553FFB6FF9EFF7AF875378AF802" blockId="44.[151,1436,1722,2034]" pageId="44" pageNumber="45">
<emphasis id="B906E941FFB6FF9EFF7AF8753770F85B" bold="true" box="[199,813,1938,1963]" pageId="44" pageNumber="45">
Female (
<typeStatus id="54C98BF1FFB6FF9EFE8DF87435C7F85B" box="[304,410,1938,1963]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
,
<taxonomicName id="4C724ED0FFB6FF9EFE17F87435A8F85B" box="[426,501,1938,1963]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="73" phylum="Arthropoda" rank="species" species="litipo">Litipo</taxonomicName>
, ZMUC). Measurements:
</emphasis>
CL 2.83, CW 2.26, AL 3.70, AW 3.00, TL 6.80, FL 0.30, SL 1.33, SW 1.30, AMEAME 0.05, AMEALE 0.01, ALEALE 0.45, PMEPME 0.13, PMEPLE 0.05, PLEPLE 0.56,
<collectingCountry id="F36575C3FFB6FF9EFEF3F83D35C7F802" box="[334,410,2011,2034]" name="Peru" pageId="44" pageNumber="45">PERW</collectingCountry>
0.83, MOQAW 0.46, MOQPW 0.48, MOQL 0.54.
</paragraph>
<paragraph id="8BCD3553FFB7FF9FFF7AFF713747FF24" blockId="45.[151,1436,151,1303]" pageId="45" pageNumber="46">Length of leg segments: I 2.45 + 1.05 + 2.00 + 1.98 + 1.18 = 8.66; II missing; III 2.25 + 0.98 + 1.74 + 2.06 + 1.03 = 8.06; IV 2.90 + 1.05 + 2.30 + 3.10 + 1.10 = 10.45.</paragraph>
<paragraph id="8BCD3553FFB7FF9FFF7AFF3937C0FBCF" blockId="45.[151,1436,151,1303]" pageId="45" pageNumber="46">
General appearance as in
<figureCitation id="134929D6FFB7FF9FFE4DFF393618FF07" box="[496,581,223,247]" captionStart="FIGURES 45 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="45" pageNumber="46">Fig. 51</figureCitation>
. Carapace bright yellow-orange, eye region grey; broad median black line from PER to posterior slope, broken up by paler line from fovea to posterior margin of carapace; pale striae radiating from fovea, directed between coxae; lateral margins with narrow mottled black fringe, expanded between leg coxae; markings covered with black feathery setae, areas between them covered in yellowish feathery and short straight setae. AME separated by distance equal to 15 their diameter; AME separated from ALE by distance less than 110 AME diameter; clypeus height slightly less than ½ AME diameter; PLE very slightly larger than PME; PME separated by distance equal to ½ their diameter; PME separated from PLE by distance equal to ¼ PME diameter; CW:
<collectingCountry id="F36575C3FFB7FF9FFE86FE0135D6FE0F" box="[315,395,487,511]" name="Peru" pageId="45" pageNumber="46">PERW</collectingCountry>
= 2.72:1. Chelicerae yellow-brown with faint black mottling, yellow prolaterally distally; promargin with two teeth separated by basal width of proximal tooth, distal tooth much larger; retromargin with two teeth separated by slightly less than their basal width, distal tooth slightly larger, close to fang base. Endites pale orange, cream prolaterally distally; labium pale orange, cream distally; sternum yellow, darker around margins. Legs (both leg II missing) yellow with faint black mottling, without distinct markings except faint black spots at spine bases and tibiae III and IV with faint proximal and distal black bands. Leg spination (both leg II missing): femora: I pl 3 do 3 rl 1, III pl 2 do 3 rl 2, IV pl 2 do 3 rl 1; all femora with scattered erect ventral setae; patellae: I, III and IV with long fine distal seta dorsally; tibiae: I pl 1 plv 2 rlv 2 spines, do 1 long fine seta distally, III pl 2 do 1 rl 2 plv 2 rlv 2, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 5 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 1 plv 1; tarsi: pl 1 plv 2 rlv 1. Abdomen with yellow-orange anterior dorsal scutum extending to ½ abdomen length; dorsum cream, sparsely covered in black feathery and short straight setae, feathery setae yellow-brown posteriorly; spinnerets surrounded by narrow faint black ring; venter cream, covered in black feathery setae; epigastric scutum and inframamillary sclerite yellow. Female epigyne with oval sclerotised rim laterally in posterior half of epigyne, with transverse oval copulatory openings situated within them (
<figureCitation id="134929D6FFB7FF9FFCDDFC39379EFC07" box="[864,963,991,1015]" captionStart="FIGURES 128 131" captionStartId="44.[151,264,1209,1232]" captionTargetBox="[175,1425,220,1167]" captionTargetId="figure@44.[151,1436,195,1186]" captionTargetPageId="44" captionText="FIGURES 128 131. Genitalic morphology of Copuetta lesnei sp. nov. (128, 129) and C. litipo sp. nov. (130, 131): 128. Male palp (mirror image of right palp), ventral view. 129. Same, retrolateral view. 130. Female epigyne, ventral view. 131. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284222/files/figure.png" pageId="45" pageNumber="46">Fig. 130</figureCitation>
); copulatory ducts very short, entering ST II posterolaterally; ST II n-shaped with anterior bend, broadly connected to diverging posterior ST I; ST with several folds on their surface, ST I equal in width to ST II (
<figureCitation id="134929D6FFB7FF9FFC93FBC137D2FBCF" box="[814,911,1063,1087]" captionStart="FIGURES 128 131" captionStartId="44.[151,264,1209,1232]" captionTargetBox="[175,1425,220,1167]" captionTargetId="figure@44.[151,1436,195,1186]" captionTargetPageId="44" captionText="FIGURES 128 131. Genitalic morphology of Copuetta lesnei sp. nov. (128, 129) and C. litipo sp. nov. (130, 131): 128. Male palp (mirror image of right palp), ventral view. 129. Same, retrolateral view. 130. Female epigyne, ventral view. 131. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284222/files/figure.png" pageId="45" pageNumber="46">Fig. 131</figureCitation>
).
</paragraph>
<paragraph id="8BCD3553FFB7FF9FFF7AFBAD35D7FB93" blockId="45.[151,1436,151,1303]" box="[199,394,1099,1123]" pageId="45" pageNumber="46">
<emphasis id="B906E941FFB7FF9FFF7AFBAD3550FB93" bold="true" box="[199,269,1099,1123]" pageId="45" pageNumber="46">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB7FF9FFF7AFB8936DCFB3F" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BCD3553FFB7FF9FFF7AFB8936A2FB5B" blockId="45.[151,1436,151,1303]" pageId="45" pageNumber="46">
<emphasis id="B906E941FFB7FF9FFF7AFB893527FB77" bold="true" box="[199,378,1134,1159]" pageId="45" pageNumber="46">
<typeStatus id="54C98BF1FFB7FF9FFF7AFB89355DFB77" box="[199,256,1135,1159]" pageId="45" pageNumber="46">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54C98BF1FFB7FF9FFE35FB8935ADFB77" box="[392,496,1135,1159]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
Ƥ:
<emphasis id="B906E941FFB7FF9FFD9BFB88369EFB77" bold="true" box="[550,707,1134,1159]" pageId="45" pageNumber="46">
<collectingCountry id="F36575C3FFB7FF9FFD9BFB8836E3FB77" box="[550,702,1134,1159]" name="Tanzania" pageId="45" pageNumber="46">TANZANIA</collectingCountry>
:
</emphasis>
Lindi Region, Lindi District,
<taxonomicName id="4C724ED0FFB7FF9FFB91FB89302DFB77" box="[1068,1136,1135,1159]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="73" phylum="Arthropoda" rank="species" species="litipo">Litipo</taxonomicName>
Forest Reserve,
<geoCoordinate id="EE465394FFB7FF9FFA80FB8931CAFB77" box="[1341,1431,1135,1159]" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="-10.033334">10°02'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFB7FF9FFF2AFB7534AFFB5B" box="[151,242,1171,1195]" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="39.483334">39°29'E</geoCoordinate>
, leg. Frontier
<collectingCountry id="F36575C3FFB7FF9FFE2DFB7535AAFB5B" box="[400,503,1171,1195]" name="Tanzania" pageId="45" pageNumber="46">Tanzania</collectingCountry>
,
<date id="FFCC1393FFB7FF9FFDBFFB7536CEFB5B" box="[514,659,1171,1195]" pageId="45" pageNumber="46" value="1993-07" valueMax="1993-09-31" valueMin="1993-07-00">VIIIX.1993</date>
(
<collectionCode id="ED63AD96FFB7FF9FFD1FFB7536ABFB5B" box="[674,758,1171,1195]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="45" pageNumber="46">ZMUC</collectionCode>
)
</paragraph>
<paragraph id="8BCD3553FFB7FF9FFF7AFB5036DCFB3F" blockId="45.[151,1436,151,1303]" box="[199,641,1206,1231]" pageId="45" pageNumber="46">
<emphasis id="B906E941FFB7FF9FFF7AFB50366BFB3F" bold="true" box="[199,566,1206,1231]" pageId="45" pageNumber="46">Additional material examined:</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB7FF9FFF7AFB3C3731FB03" box="[199,876,1242,1268]" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BCD3553FFB7FF9FFF7AFB3C3731FB03" blockId="45.[151,1436,151,1303]" box="[199,876,1242,1268]" pageId="45" pageNumber="46">
<emphasis id="B906E941FFB7FF9FFF7AFB3C353FFB03" bold="true" box="[199,354,1242,1267]" pageId="45" pageNumber="46">Distribution:</emphasis>
Known only from the
<typeStatus id="54C98BF1FFB7FF9FFDDFFB3A36CFFB04" box="[610,658,1244,1268]" pageId="45" pageNumber="46">type</typeStatus>
locality (
<figureCitation id="134929D6FFB7FF9FFD40FB3D3702FB03" box="[765,863,1243,1267]" captionStart="FIGURE 132" captionStartId="46.[151,250,1450,1473]" captionTargetBox="[151,1425,193,1417]" captionTargetId="figure@46.[151,1436,193,1429]" captionTargetPageId="46" captionText="FIGURE 132. Distribution of Copuetta comorica sp. nov. (black circles), C. erecta sp. nov. (white circles), C. kakamega sp. nov. (black star), C. kwamgumi sp. nov. (white star), C. lacustris (Strand, 1916) (black triangles), C. lesnei sp. nov. (white triangle) and C. litipo sp. nov. (cross) in the Afrotropical Region." httpUri="https://zenodo.org/record/284223/files/figure.png" pageId="45" pageNumber="46">Fig. 132</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFB7FF9FFF7AFB183023FAE7" box="[199,1150,1278,1303]" pageId="45" pageNumber="46" type="biology_ecology">
<paragraph id="8BCD3553FFB7FF9FFF7AFB183023FAE7" blockId="45.[151,1436,151,1303]" box="[199,1150,1278,1303]" pageId="45" pageNumber="46">
<emphasis id="B906E941FFB7FF9FFF7AFB183577FAE7" bold="true" box="[199,298,1278,1303]" pageId="45" pageNumber="46">Biology:</emphasis>
Collected in a forest but details of the sampling methodology are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>