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<mods:title id="9F2D213035523784164A1642956B1E0F">On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)</mods:title>
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<mods:namePart id="1A16B129D4546DDDDD3243F31B6FBAA9">Li, Yan-Da</mods:namePart>
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<mods:namePart id="F98B90C0AEF4A62A2558DC73336C752D">Tihelka, Erik</mods:namePart>
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<mods:affiliation id="F7D99AE2CD19E8EBA562FB0B3DC3D752">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355</mods:affiliation>
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<mods:namePart id="F6C7A46AACA0BFC68ACA792468B0BB06">Dahan, Loïc</mods:namePart>
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<mods:namePart id="54D903CB3D6541F04C53B10B677FD862">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="D7482AF9563A59B0E9DB9B4316996950">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
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<treatment id="03D787A2FFEDEE79FF52F97F8E296FB7" ID-DOI="http://doi.org/10.5281/zenodo.5788245" ID-GBIF-Taxon="191654780" ID-Zenodo-Dep="5788245" LSID="urn:lsid:plazi:treatment:03D787A2FFEDEE79FF52F97F8E296FB7" httpUri="http://treatment.plazi.org/id/03D787A2FFEDEE79FF52F97F8E296FB7" lastPageId="14" lastPageNumber="237" pageId="13" pageNumber="236">
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<paragraph id="8BC136B4FFEDEE7AFF52F97F89BD6AFD" blockId="13.[151,708,1749,1812]" box="[151,708,1749,1775]" pageId="13" pageNumber="236">
<heading id="D08981D8FFEDEE7AFF52F97F89BD6AFD" bold="true" box="[151,708,1749,1775]" fontSize="11" level="1" pageId="13" pageNumber="236" reason="1">
<emphasis id="B90AEAA6FFEDEE7AFF52F97F89BD6AFD" bold="true" box="[151,708,1749,1775]" pageId="13" pageNumber="236">
Genus
<taxonomicName id="4C7E4D37FFEDEE7AFF2BF97F89BD6AFD" authority="Tihelka, Huang &amp; Cai, 2020" authorityName="Tihelka, Huang &amp; Cai" authorityYear="2020" box="[238,708,1749,1775]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAA6FFEDEE7AFF2BF97F8A246AFD" bold="true" box="[238,349,1749,1775]" italics="true" pageId="13" pageNumber="236">Basinosa</emphasis>
Tihelka, Huang &amp; Cai, 2020
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<subSubSection id="C364653FFFEDEE7AFF52F9538A5E6B06" box="[151,295,1785,1812]" pageId="13" pageNumber="236" type="description">
<paragraph id="8BC136B4FFEDEE7AFF52F9538A5E6B06" blockId="13.[151,708,1749,1812]" box="[151,295,1785,1812]" pageId="13" pageNumber="236">
(
<figureCitation id="13452A31FFEDEE7AFF5AF9538B8F6B06" box="[159,246,1785,1812]" captionStart="FIGURE 15" captionStartId="16.[151,250,1352,1376]" captionTargetBox="[151,1436,453,1329]" captionTargetId="figure-166@16.[151,1436,453,1329]" captionTargetPageId="16" captionText="FIGURE 15. General habitus of Basinosa pengweii Tihelka et al., holotype, NIGP173119, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5788265" httpUri="https://zenodo.org/record/5788265/files/figure.png" pageId="13" pageNumber="236">Figs 15</figureCitation>
,
<figureCitation id="13452A31FFEDEE7AFEC7F9538A666B06" box="[258,287,1785,1812]" captionStart="FIGURE 16" captionStartId="17.[151,250,1067,1091]" captionTargetBox="[151,1436,181,1043]" captionTargetId="figure-18@17.[151,1436,181,1043]" captionTargetPageId="17" captionText="FIGURE 16. Details of Basinosa pengweii Tihelka et al., holotype, NIGP173119. A, Weighted average stacking of micro-CT slices, showing the left antenna and antennal groove on head. BF, confocal images. B, C, Antenna, ventral view. D, Mandibles, dorsal view. E, Prosternal process, ventral view. F, Metatarsus.Abbreviations: ag, antennal groove; an, antenna; an311, antennomeres 311; md, mandible; mtt14, metatarsomeres 14; mttb, metatibia; ps, prosternum. Scale bars: 1500 μm in A, 400 μm in BF." figureDoi="http://doi.org/10.5281/zenodo.5788267" httpUri="https://zenodo.org/record/5788267/files/figure.png" pageId="13" pageNumber="236">16</figureCitation>
)
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<paragraph id="8BC136B4FFEDEE7AFF52F894886F6B44" blockId="13.[151,790,1853,1878]" box="[151,790,1853,1878]" pageId="13" pageNumber="236">
<emphasis id="B90AEAA6FFEDEE7AFF52F8948A5A6B47" bold="true" box="[151,291,1853,1878]" pageId="13" pageNumber="236">
<typeStatus id="54C58816FFEDEE7AFF52F8948BB56B44" box="[151,204,1854,1878]" pageId="13" pageNumber="236">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C7E4D37FFEDEE7AFEEFF894886B6B44" authority="Tihelka, Huang &amp; Cai, 2020" authorityName="Tihelka, Huang &amp; Cai" authorityYear="2020" box="[298,786,1854,1878]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="pengweii">
<emphasis id="B90AEAA6FFEDEE7AFEEFF8948A916B47" box="[298,488,1854,1877]" italics="true" pageId="13" pageNumber="236">Basinosa pengweii</emphasis>
Tihelka, Huang &amp; Cai, 2020
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFEDEE7AFF52F82E883A6B8C" box="[151,835,1924,1950]" pageId="13" pageNumber="236" type="materials_examined">
<paragraph id="8BC136B4FFEDEE7AFF52F82E883A6B8C" blockId="13.[151,1436,1924,2022]" box="[151,835,1924,1950]" pageId="13" pageNumber="236">
<emphasis id="B90AEAA6FFEDEE7AFF52F82E8AF86B8C" bold="true" box="[151,385,1924,1950]" pageId="13" pageNumber="236">Material examined.</emphasis>
<typeStatus id="54C58816FFEDEE7AFE42F82E8A966B8C" box="[391,495,1924,1950]" pageId="13" pageNumber="236" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C7E4D37FFEDEE7AFDD0F82F89926B8C" authority=", NIGP" authorityName="NIGP" box="[533,747,1924,1950]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="pengweii">
<emphasis id="B90AEAA6FFEDEE7AFDD0F82F89E06B8C" box="[533,665,1925,1950]" italics="true" pageId="13" pageNumber="236">B. pengweii</emphasis>
, NIGP
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173119.
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<paragraph id="8BC136B4FFEDEE79FF02F80288C06CEB" blockId="13.[151,1436,1924,2022]" lastBlockId="14.[151,1437,151,933]" lastPageId="14" lastPageNumber="237" pageId="13" pageNumber="236">
<emphasis id="B90AEAA6FFEDEE7AFF02F8028A426BD0" bold="true" box="[199,315,1960,1986]" pageId="13" pageNumber="236">Remarks.</emphasis>
The amber piece containing the
<typeStatus id="54C58816FFEDEE7AFD6CF80288726BD0" box="[681,779,1960,1986]" pageId="13" pageNumber="236" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7E4D37FFEDEE7AFCF4F80388CC6BD0" box="[817,949,1961,1986]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="pengweii">
<emphasis id="B90AEAA6FFEDEE7AFCF4F80388CC6BD0" box="[817,949,1961,1986]" italics="true" pageId="13" pageNumber="236">B. pengweii</emphasis>
</taxonomicName>
is somewhat nontransparent, making it difficult to determine many of the characters using normal optical methods. Partly due to tomographic artefacts, some characters were not clearly visualized in previous micro-CT reconstructions. For example, one of the key diagnostic characters for
<taxonomicName id="4C7E4D37FFEEEE79FEFDFF118A9D6CC7" authorityName="Erichson" authorityYear="1846" box="[312,484,187,213]" class="Insecta" family="Nosodendridae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="family">Nosodendridae</taxonomicName>
, the 11-segmented antenna with 3-segmented club, could not be observed in
<bibRefCitation id="EFEF4B45FFEEEE79FA8DFF118A496CEB" author="Tihelka, E. &amp; Huang, D. &amp; Cai, C." pageId="14" pageNumber="237" pagination="104701" refId="ref8229" refString="Tihelka, E., Huang, D. &amp; Cai, C. (2021 a) A new genus of wounded-tree beetles from mid-Cretaceous northern Myanmar amber (Coleoptera: Nosodendridae). Cretaceous Research, 119, 104701. https: // doi. org / 10.1016 / j. cretres. 2020.104701" type="journal article" year="2021">
Tihelka
<emphasis id="B90AEAA6FFEEEE79FF52FF758BB66CEB" box="[151,207,223,249]" italics="true" pageId="14" pageNumber="237">et al.</emphasis>
(2021a)
</bibRefCitation>
, although other diagnostic characters could be confirmed.
</paragraph>
<paragraph id="8BC136B4FFEEEE79FF02FEA989166EE3" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="237">
Here we reexamine the original CT data for the
<typeStatus id="54C58816FFEEEE79FD2BFEA988296D0F" box="[750,848,259,285]" pageId="14" pageNumber="237" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7E4D37FFEEEE79FCBCFEA988866D0E" box="[889,1023,259,284]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="species" species="pengweii">
<emphasis id="B90AEAA6FFEEEE79FCBCFEA988866D0E" box="[889,1023,259,284]" italics="true" pageId="14" pageNumber="237">B. pengweii</emphasis>
</taxonomicName>
. It turns out that the majority of the beetle body has high absorption (white), while the antennae (especially the left one) appear to have lower absorption (black) (
<figureCitation id="13452A31FFEEEE79FEE2FEE18AF76D74" box="[295,398,331,358]" captionStart="FIGURE 16" captionStartId="17.[151,250,1067,1091]" captionTargetBox="[151,1436,181,1043]" captionTargetId="figure-18@17.[151,1436,181,1043]" captionTargetPageId="17" captionText="FIGURE 16. Details of Basinosa pengweii Tihelka et al., holotype, NIGP173119. A, Weighted average stacking of micro-CT slices, showing the left antenna and antennal groove on head. BF, confocal images. B, C, Antenna, ventral view. D, Mandibles, dorsal view. E, Prosternal process, ventral view. F, Metatarsus.Abbreviations: ag, antennal groove; an, antenna; an311, antennomeres 311; md, mandible; mtt14, metatarsomeres 14; mttb, metatibia; ps, prosternum. Scale bars: 1500 μm in A, 400 μm in BF." figureDoi="http://doi.org/10.5281/zenodo.5788267" httpUri="https://zenodo.org/record/5788267/files/figure.png" pageId="14" pageNumber="237">Fig. 16A</figureCitation>
). Therefore, the antennae were not properly reconstructed in the simple reconstruction, where only regions with a density higher than a selected threshold were extracted. Confocal microscopy confirms the 3-segmented club of
<taxonomicName id="4C7E4D37FFEEEE79FE4CFE3989766DBE" box="[393,527,403,428]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="species" species="pengweii">
<emphasis id="B90AEAA6FFEEEE79FE4CFE3989766DBE" box="[393,527,403,428]" italics="true" pageId="14" pageNumber="237">B. pengweii</emphasis>
</taxonomicName>
(
<figureCitation id="13452A31FFEEEE79FDE4FE3989F36DBC" box="[545,650,403,430]" captionStart="FIGURE 16" captionStartId="17.[151,250,1067,1091]" captionTargetBox="[151,1436,181,1043]" captionTargetId="figure-18@17.[151,1436,181,1043]" captionTargetPageId="17" captionText="FIGURE 16. Details of Basinosa pengweii Tihelka et al., holotype, NIGP173119. A, Weighted average stacking of micro-CT slices, showing the left antenna and antennal groove on head. BF, confocal images. B, C, Antenna, ventral view. D, Mandibles, dorsal view. E, Prosternal process, ventral view. F, Metatarsus.Abbreviations: ag, antennal groove; an, antenna; an311, antennomeres 311; md, mandible; mtt14, metatarsomeres 14; mttb, metatibia; ps, prosternum. Scale bars: 1500 μm in A, 400 μm in BF." figureDoi="http://doi.org/10.5281/zenodo.5788267" httpUri="https://zenodo.org/record/5788267/files/figure.png" pageId="14" pageNumber="237">Fig. 16C</figureCitation>
). Since the strong laser could penetrate the somewhat opaque amber matrix better, and out-of-focus light is blocked, the confocal microscope could image this material much better than widefield ones (
<bibRefCitation id="EFEF4B45FFEEEE79FE8CFE718A916DE7" author="Fu, Y. - Z. &amp; Li, Y. - D. &amp; Su, Y. - T. &amp; Cai, C. - Y. &amp; Huang, D. - Y." box="[329,488,475,501]" pageId="14" pageNumber="237" pagination="266 - 278" refId="ref7446" refString="Fu, Y. - Z., Li, Y. - D., Su, Y. - T., Cai, C. - Y. &amp; Huang, D. - Y. (2021) Application of confocal laser scanning microscopy to the study of amber bioinclusions. Palaeoentomology, 4, 266 - 278. https: // doi. org / 10.11646 / palaeoentomology. 4.3.14" type="journal article" year="2021">
Fu
<emphasis id="B90AEAA6FFEEEE79FEABFE718ADF6DE7" box="[366,422,475,501]" italics="true" pageId="14" pageNumber="237">et al.</emphasis>
2021
</bibRefCitation>
). The medial antennomeres of
<taxonomicName id="4C7E4D37FFEEEE79FCFAFE7188BB6DE6" box="[831,962,475,500]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="species" species="pengweii">
<emphasis id="B90AEAA6FFEEEE79FCFAFE7188BB6DE6" box="[831,962,475,500]" italics="true" pageId="14" pageNumber="237">B. pengweii</emphasis>
</taxonomicName>
appears to be relative elongate (
<figureCitation id="13452A31FFEEEE79FAECFE718EF66DE4" box="[1321,1423,475,502]" captionStart="FIGURE 16" captionStartId="17.[151,250,1067,1091]" captionTargetBox="[151,1436,181,1043]" captionTargetId="figure-18@17.[151,1436,181,1043]" captionTargetPageId="17" captionText="FIGURE 16. Details of Basinosa pengweii Tihelka et al., holotype, NIGP173119. A, Weighted average stacking of micro-CT slices, showing the left antenna and antennal groove on head. BF, confocal images. B, C, Antenna, ventral view. D, Mandibles, dorsal view. E, Prosternal process, ventral view. F, Metatarsus.Abbreviations: ag, antennal groove; an, antenna; an311, antennomeres 311; md, mandible; mtt14, metatarsomeres 14; mttb, metatibia; ps, prosternum. Scale bars: 1500 μm in A, 400 μm in BF." figureDoi="http://doi.org/10.5281/zenodo.5788267" httpUri="https://zenodo.org/record/5788267/files/figure.png" pageId="14" pageNumber="237">Fig. 16B</figureCitation>
), while in extant
<taxonomicName id="4C7E4D37FFEEEE79FE82FE558AA46E0B" authorityName="Latreille" authorityYear="1804" box="[327,477,511,537]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAA6FFEEEE79FE82FE558AA46E0B" box="[327,477,511,537]" italics="true" pageId="14" pageNumber="237">Nosodendron</emphasis>
</taxonomicName>
the medial antennomeres are almost as long as wide. Confocal microscopy additionally reveals that the prosternal process (
<figureCitation id="13452A31FFEEEE79FD98FD8989BB6E2C" box="[605,706,547,574]" captionStart="FIGURE 16" captionStartId="17.[151,250,1067,1091]" captionTargetBox="[151,1436,181,1043]" captionTargetId="figure-18@17.[151,1436,181,1043]" captionTargetPageId="17" captionText="FIGURE 16. Details of Basinosa pengweii Tihelka et al., holotype, NIGP173119. A, Weighted average stacking of micro-CT slices, showing the left antenna and antennal groove on head. BF, confocal images. B, C, Antenna, ventral view. D, Mandibles, dorsal view. E, Prosternal process, ventral view. F, Metatarsus.Abbreviations: ag, antennal groove; an, antenna; an311, antennomeres 311; md, mandible; mtt14, metatarsomeres 14; mttb, metatibia; ps, prosternum. Scale bars: 1500 μm in A, 400 μm in BF." figureDoi="http://doi.org/10.5281/zenodo.5788267" httpUri="https://zenodo.org/record/5788267/files/figure.png" pageId="14" pageNumber="237">Fig. 16E</figureCitation>
) is not as slender as shown in the CT reconstruction by
<bibRefCitation id="EFEF4B45FFEEEE79FA8DFD898A4A6E73" author="Tihelka, E. &amp; Huang, D. &amp; Cai, C." pageId="14" pageNumber="237" pagination="104701" refId="ref8229" refString="Tihelka, E., Huang, D. &amp; Cai, C. (2021 a) A new genus of wounded-tree beetles from mid-Cretaceous northern Myanmar amber (Coleoptera: Nosodendridae). Cretaceous Research, 119, 104701. https: // doi. org / 10.1016 / j. cretres. 2020.104701" type="journal article" year="2021">
Tihelka
<emphasis id="B90AEAA6FFEEEE79FF52FDED8BA86E73" box="[151,209,583,609]" italics="true" pageId="14" pageNumber="237">et al.</emphasis>
(2021a)
</bibRefCitation>
, which was subsequently adopted as one of the characters differentiating
<taxonomicName id="4C7E4D37FFEEEE79FBB9FDED8F9A6E72" authorityName="Tihelka, Huang &amp; Cai" authorityYear="2020" box="[1148,1251,583,608]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAA6FFEEEE79FBB9FDED8F9A6E72" box="[1148,1251,583,608]" italics="true" pageId="14" pageNumber="237">Basinosa</emphasis>
</taxonomicName>
from
<taxonomicName id="4C7E4D37FFEEEE79FAEEFDED8AE86E94" authority="(Tihelka et al. 2020 b)" baseAuthorityName="Tihelka" baseAuthorityYear="2020" class="Insecta" family="Nosodendridae" genus="Mesonosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAA6FFEEEE79FAEEFDED8EE46E72" box="[1323,1437,583,608]" italics="true" pageId="14" pageNumber="237">Mesonosa</emphasis>
(Tihelka
<emphasis id="B90AEAA6FFEEEE79FF38FDC18A4F6E97" box="[253,310,619,645]" italics="true" pageId="14" pageNumber="237">et al.</emphasis>
2020b)
</taxonomicName>
. Detecting the boundaries of structures under CT is a challenging task (
<bibRefCitation id="EFEF4B45FFEEEE79FB79FDC18BAD6EBB" author="Coleman, M. N. &amp; Colbert, M. W." pageId="14" pageNumber="237" pagination="723 - 725" refId="ref7329" refString="Coleman, M. N. &amp; Colbert, M. W. (2007) CT thresholding protocols for taking measurements on three-dimensional models. American Journal of Physical Anthropology, 133, 723 - 725. https: // doi. org / 10.1002 / ajpa. 20583" type="journal article" year="2007">Coleman &amp; Colbert 2007</bibRefCitation>
); the boundaries between the beetle and the amber matrix are not clearly defined, but are instead represented by a continuum of transitional values. Thus, a different selection of the threshold value for extraction could sometimes greatly affect the visual appearance.
</paragraph>
<paragraph id="8BC136B4FFEEEE79FF02FD518E296FB7" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="237">
The 3-segmented antennal club, together with the previously shown subantennal groove, wide metaventrite, clearly delimited anterolateral depression of metaventrite for the reception of mesotibial apex and basal mesotarsomeres, and narrowly separated metacoxae (all present in extant
<taxonomicName id="4C7E4D37FFEEEE79FCA0FCE98F766F4F" authorityName="Erichson" authorityYear="1846" box="[869,1039,835,861]" class="Insecta" family="Nosodendridae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="family">Nosodendridae</taxonomicName>
and fossil
<taxonomicName id="4C7E4D37FFEEEE79FB4EFCE98E4C6F4F" authorityName="Erichson" authorityYear="1846" box="[1163,1333,835,861]" class="Insecta" family="Nosodendridae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="family">Nosodendridae</taxonomicName>
from the Burmese amber; fig. 23C in Ge
<emphasis id="B90AEAA6FFEEEE79FE3CFCCD89486F93" box="[505,561,871,897]" italics="true" pageId="14" pageNumber="237">et al.</emphasis>
2007), further confirm the placement of
<taxonomicName id="4C7E4D37FFEEEE79FC33FCCD8F246F92" authorityName="Tihelka, Huang &amp; Cai" authorityYear="2020" box="[1014,1117,871,896]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAA6FFEEEE79FC33FCCD8F246F92" box="[1014,1117,871,896]" italics="true" pageId="14" pageNumber="237">Basinosa</emphasis>
</taxonomicName>
in
<taxonomicName id="4C7E4D37FFEEEE79FBBAFCCD8E556F93" authorityName="Erichson" authorityYear="1846" box="[1151,1324,871,897]" class="Insecta" family="Nosodendridae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="family">Nosodendridae</taxonomicName>
.
<taxonomicName id="4C7E4D37FFEEEE79FAF3FCCD8EE46F92" authorityName="Tihelka, Huang &amp; Cai" authorityYear="2020" box="[1334,1437,871,896]" class="Insecta" family="Nosodendridae" genus="Basinosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAA6FFEEEE79FAF3FCCD8EE46F92" box="[1334,1437,871,896]" italics="true" pageId="14" pageNumber="237">Basinosa</emphasis>
</taxonomicName>
is unique in
<taxonomicName id="4C7E4D37FFEEEE79FEE4FC218AB26FB7" authorityName="Erichson" authorityYear="1846" box="[289,459,907,933]" class="Insecta" family="Nosodendridae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="237" phylum="Arthropoda" rank="family">Nosodendridae</taxonomicName>
in having a relatively large body and complete antennal grooves on prosternum.
</paragraph>
</subSubSection>
</treatment>
</document>