251 lines
33 KiB
XML
251 lines
33 KiB
XML
<document id="86D590DD067FAF7265B7C92660B0110B" ID-CLB-Dataset="45587" ID-DOI="10.11646/zootaxa.3811.4.2" ID-GBIF-Dataset="dad75c9b-1930-47fb-ba79-eb519a90c45a" ID-ISSN="1175-5326" ID-Zenodo-Dep="253023" ID-ZooBank="99629EFB-F2F6-4DE8-AABA-D71069106912" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432064619" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2014" docId="03D687DBE35A0C12BBB98E90FA7F9F08" docLanguage="en" docName="zt03811p462.pdf" docOrigin="Zootaxa 3811 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterosthetops coriaceus Bilton, 2014, sp. nov." docType="treatment" docVersion="11" lastPageNumber="443" masterDocId="FFEFFFA3E35B0C17BB2E8A1EFFE99E6C" masterDocTitle="New species and new records of Pterosthetops: eumadicolous water beetles of the South African Cape (Coleoptera, Hydraenidae)" masterLastPageNumber="462" masterPageNumber="438" pageNumber="439" updateTime="1698331499332" updateUser="ExternalLinkService">
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<mods:mods id="362103DD009800D99DBD8F63E951C51B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AF906088B3899962B95D7DFF4F45F1A1">
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<mods:title id="791C063BE60EBEA4D6E75353C97A2EA5">New species and new records of Pterosthetops: eumadicolous water beetles of the South African Cape (Coleoptera, Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6485D30AA6A4B0CE12A5192C07DCEBB4" type="personal">
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<mods:role id="96A218A5D6BCD00487072819582DCF48">
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<mods:roleTerm id="FAA0E1B97DDA7DFE82A92FC353B6E559">Author</mods:roleTerm>
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<mods:namePart id="4E5617DB6845875798E1835FEA187CAA">Bilton, David T.</mods:namePart>
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<mods:typeOfResource id="BDFC4A51A0402A1C4EE5DDCF1CCD4083">text</mods:typeOfResource>
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<mods:titleInfo id="3202B13E95197962E62978B49596E97E">
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<mods:title id="38927EF2A4364C26A6474468634F35BC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CDD52BADA168FCB94326E1931A7D2F05">
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<mods:date id="BADD8CFB33D3EA6E474B422683282BCF">2014</mods:date>
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<mods:detail id="F216AA5A30F71BB4499180B9A20E1503" type="volume">
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<mods:number id="61A3D80631FEDC7C0278BF71FCED6523">3811</mods:number>
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</mods:detail>
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<mods:detail id="14FBFE2CCFD2D7F0EAD2330ED0D712F4" type="issue">
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<mods:number id="7754A7288236C76D7502CBDF8F46FDAA">4</mods:number>
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<mods:end id="4843217C40FD1EDB014B429EF581A637">462</mods:end>
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</mods:part>
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<mods:classification id="B89A60170E1DFDADD40C10F3FA5DD43B">journal article</mods:classification>
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<mods:identifier id="EC9E9A854F8C670F6CA9FC4DEEAD1B03" type="DOI">10.11646/zootaxa.3811.4.2</mods:identifier>
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<mods:identifier id="13060583CF5D67461423C119E1A24BE7" type="GBIF-Dataset">dad75c9b-1930-47fb-ba79-eb519a90c45a</mods:identifier>
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<mods:identifier id="FE79C3D70A5E3399E2261D7131540C9D" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03D687DBE35A0C12BBB98E90FA7F9F08" ID-DOI="http://doi.org/10.5281/zenodo.6129813" ID-GBIF-Taxon="119430550" ID-Zenodo-Dep="6129813" LSID="urn:lsid:plazi:treatment:03D687DBE35A0C12BBB98E90FA7F9F08" httpUri="http://treatment.plazi.org/id/03D687DBE35A0C12BBB98E90FA7F9F08" lastPageId="5" lastPageNumber="443" pageId="1" pageNumber="439">
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<subSubSection id="C3656546E35A0C16BBB98E90FEA09AA6" pageId="1" pageNumber="439" type="nomenclature">
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<paragraph id="8BC036CDE35A0C16BBB98E90FDF09AC4" blockId="1.[151,537,1166,1226]" box="[151,537,1166,1192]" pageId="1" pageNumber="439">
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<heading id="D08881A1E35A0C16BBB98E90FDF09AC4" bold="true" box="[151,537,1166,1192]" fontSize="11" level="1" pageId="1" pageNumber="439" reason="1">
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<emphasis id="B90BEADFE35A0C16BBB98E90FDF09AC4" bold="true" box="[151,537,1166,1192]" pageId="1" pageNumber="439">
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||
<taxonomicName id="4C7F4D4EE35A0C16BBB98E90FE5D9AC4" ID-CoL="6WM3Q" box="[151,436,1166,1192]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="coriaceus" status="sp. nov.">
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<emphasis id="B90BEADFE35A0C16BBB98E90FE5D9AC4" bold="true" box="[151,436,1166,1192]" italics="true" pageId="1" pageNumber="439">Pterosthetops coriaceus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857A4E35A0C16BA928E90FDF09AC4" box="[444,537,1166,1192]" pageId="1" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BC036CDE35A0C16BBB98EAFFEA09AA6" blockId="1.[151,537,1166,1226]" box="[151,329,1201,1226]" pageId="1" pageNumber="439">
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||
(
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||
<figureCitation id="13442A48E35A0C16BBB18EAFFF089AA6" box="[159,225,1201,1226]" captionStart="FIGURE 1" captionStartId="3.[151,250,1872,1894]" captionTargetBox="[181,1409,220,1827]" captionTargetId="figure@3.[151,1436,193,1851]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pterosthetops holotypes. A) P. coriaceus sp. nov.; B) P. indwei sp. nov.; C) P. swartbergensis sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/253024/files/figure.png" pageId="1" pageNumber="439">Figs 1</figureCitation>
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A & 2A)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656546E35A0C16BBB98EECFF09986E" pageId="1" pageNumber="439" type="materials_examined">
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<paragraph id="8BC036CDE35A0C16BBB98EECFAAD9B42" blockId="1.[151,1437,1266,2030]" pageId="1" pageNumber="439">
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||
<emphasis id="B90BEADFE35A0C16BBB98EECFED89B67" bold="true" box="[151,305,1266,1291]" pageId="1" pageNumber="439">
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||
<typeStatus id="54C4886FE35A0C16BBB98EECFF3B9B67" box="[151,210,1266,1291]" pageId="1" pageNumber="439">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingCountry id="F368765DE35A0C16BA6D8EECFE099B67" box="[323,480,1266,1291]" name="South Africa" pageId="1" pageNumber="439">
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<emphasis id="B90BEADFE35A0C16BA6D8EECFE099B67" bold="true" box="[323,480,1266,1291]" pageId="1" pageNumber="439">South Africa</emphasis>
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</collectingCountry>
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, Western Cape, Cederberg mountains, seepages over exposed sloping rock beside stream @
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<quantity id="4C879B28E35A0C16BA278F08FE8C9B42" box="[265,357,1302,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="439" unit="m" value="1200.0">1,200 m</quantity>
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below Wolfberg Arch, beside Matjiesrivier-Wupperthal road, D.T. Bilton leg. (
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<figureCitation id="13442A48E35A0C16BFF38F0BFAF69B42" box="[1245,1311,1301,1326]" captionStart="FIGURE 5" captionStartId="17.[151,250,1843,1865]" captionTargetBox="[193,1381,200,1808]" captionTargetId="figure@17.[186,1391,193,1814]" captionTargetPageId="17" captionText="FIGURE 5. Habitats of Pterosthetops species. A) Cederberg, seepage over sloping rock beside stream @ 1,200 m below Wolfberg Arch, beside Matjiesrivier-Wupperthal road— P. coriaceus sp. nov., P. hawequas, P. impressus; B) Witzenberg, Michell’s Pass, wet rock face along R 46 road - P. baini sp. nov., P. hawequas, P. impressus (arrow indicates larva of Aspidytes niobe Ribera, Beutel, Balke & Vogler, 2002); C) Cederberg, Uitkyk Pass, seepages over rock face beside road— P. harrisoni, P. hawequas, P. impressus, P. pulcherrimus sp. nov., P. tuberculatus sp. nov., P. uitkyki sp. nov.; D) Table Mountain, seepage over rock in Echo Valley— P. equestrius, P. impressus. Photos D T & R Bilton." httpUri="https://zenodo.org/record/253028/files/figure.png" pageId="1" pageNumber="439">Fig. 5</figureCitation>
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A).
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</paragraph>
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<paragraph id="8BC036CDE35A0C16BBE98F27FE7D9BF4" blockId="1.[151,1437,1266,2030]" pageId="1" pageNumber="439">
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||
<emphasis id="B90BEADFE35A0C16BBE98F27FE869B3E" bold="true" box="[199,367,1337,1362]" pageId="1" pageNumber="439">
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||
<typeStatus id="54C4886FE35A0C16BBE98F27FEEB9B3E" box="[199,258,1337,1362]" pageId="1" pageNumber="439">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="54C4886FE35A0C16BA518F27FE059B3E" box="[383,492,1337,1362]" pageId="1" pageNumber="439" type="holotype">
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<emphasis id="B90BEADFE35A0C16BA518F27FE059B3E" bold="true" box="[383,492,1337,1362]" pageId="1" pageNumber="439">Holotype</emphasis>
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</typeStatus>
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(male): “
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<date id="FFC1100DE35A0C16B9778F27FD3A9B3E" box="[601,723,1337,1362]" pageId="1" pageNumber="439" value="2011-09-23">23/ix/2011</date>
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<collectingCountry id="F368765DE35A0C16B9F28F27FC849B3E" box="[732,877,1337,1362]" name="South Africa" pageId="1" pageNumber="439">South Africa</collectingCountry>
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WC// Cederberg—stream with seeps// over rock @
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<quantity id="4C879B28E35A0C16BB928F43FEF59B18" box="[188,284,1373,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="439" unit="m" value="1200.0">1,200 m</quantity>
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below// Wolfberg Arch D T Bilton leg.” (genitalia extracted and mounted on same card) and red
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<typeStatus id="54C4886FE35A0C16BBB98F61FF109BF4" box="[151,249,1407,1432]" pageId="1" pageNumber="439" type="holotype">holotype</typeStatus>
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label (
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<collectionCode id="ED6EAE08E35A0C16BA6B8F9EFE619BF4" box="[325,392,1408,1432]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="1" pageNumber="439">SAM</collectionCode>
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||
).
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||
</paragraph>
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||
<paragraph id="8BC036CDE35A0C16BBE98FBCFF09986E" blockId="1.[151,1437,1266,2030]" pageId="1" pageNumber="439">
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||
<typeStatus id="54C4886FE35A0C16BBE98FBCFED69BD7" box="[199,319,1442,1467]" pageId="1" pageNumber="439" type="paratype">
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||
<emphasis id="B90BEADFE35A0C16BBE98FBCFED69BD7" bold="true" box="[199,319,1442,1467]" pageId="1" pageNumber="439">Paratypes</emphasis>
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||
</typeStatus>
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||
(9):
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<collectingCountry id="F368765DE35A0C16BA598FBCFDF89BD7" box="[375,529,1442,1467]" name="South Africa" pageId="1" pageNumber="439">
|
||
<emphasis id="B90BEADFE35A0C16BA598FBCFDF89BD7" bold="true" box="[375,529,1442,1467]" pageId="1" pageNumber="439">South Africa</emphasis>
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||
</collectingCountry>
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||
:
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||
<specimenCount id="9D79FD44E35A0C16B90F8FBDFDA79BD6" box="[545,590,1443,1467]" pageId="1" pageNumber="439" type="generic">4 ♂</specimenCount>
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||
, 5 ♀ “
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||
<date id="FFC1100DE35A0C16B9B78FBCFCFA9BD7" box="[665,787,1442,1467]" pageId="1" pageNumber="439" value="2011-09-23">23/ix/2011</date>
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||
<collectingCountry id="F368765DE35A0C16B8358FBCFC449BD7" box="[795,941,1442,1467]" name="South Africa" pageId="1" pageNumber="439">South Africa</collectingCountry>
|
||
WC// Cederberg—stream with seeps// over rock @
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||
<quantity id="4C879B28E35A0C16BBDA8FD8FEBB9BB2" box="[244,338,1478,1502]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="439" unit="m" value="1200.0">1,200 m</quantity>
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||
below// Wolfberg Arch D T Bilton leg.” (
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<collectionCode id="ED6EAE08E35A0C16B8188FD8FC6C9BB2" box="[822,901,1478,1502]" pageId="1" pageNumber="439">CDTB</collectionCode>
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,
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<collectionCode id="ED6EAE08E35A0C16B8BC8FD9FC339BB2" box="[914,986,1479,1502]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="439">NMW</collectionCode>
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,
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<collectionCode id="ED6EAE08E35A0C16B8C98FD8FBDD9BB2" LSID="urn:lsid:biocol.org:col:34248" box="[999,1076,1478,1502]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="1" pageNumber="439">SANC</collectionCode>
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,
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<collectionCode id="ED6EAE08E35A0C16BF6F8FD8FB7D9BB2" box="[1089,1172,1478,1502]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="1" pageNumber="439">TMSA</collectionCode>
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). All with red
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<typeStatus id="54C4886FE35A0C16BE128FD9FA759BB3" box="[1340,1436,1479,1503]" pageId="1" pageNumber="439" type="paratype">paratype</typeStatus>
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labels.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3656546E35A0C12BBE98C12FC7E9E94" lastPageId="5" lastPageNumber="443" pageId="1" pageNumber="439" type="description">
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||
<paragraph id="8BC036CDE35A0C16BBE98C12FD6098DE" blockId="1.[151,1437,1266,2030]" pageId="1" pageNumber="439">
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||
<emphasis id="B90BEADFE35A0C16BBE98C12FEB89849" bold="true" box="[199,337,1548,1573]" pageId="1" pageNumber="439">Description</emphasis>
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||
. Size:
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<typeStatus id="54C4886FE35A0C16BA8A8C12FDF89849" box="[420,529,1548,1573]" pageId="1" pageNumber="439" type="holotype">Holotype</typeStatus>
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: BL
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<quantity id="4C879B28E35A0C16B9628C12FD5F9848" box="[588,694,1548,1573]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="1" pageNumber="439" unit="mm" value="2.15">2.15 mm</quantity>
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||
; EL
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||
<quantity id="4C879B28E35A0C16B9DF8C13FCA49848" box="[753,845,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="439" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; EW
|
||
<quantity id="4C879B28E35A0C16B8BE8C13FC049848" box="[912,1005,1549,1573]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="439" unit="mm" value="0.8">0.8 mm</quantity>
|
||
.
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||
<typeStatus id="54C4886FE35A0C16B8D48C13FB859849" box="[1018,1132,1549,1573]" pageId="1" pageNumber="439" type="paratype">Paratypes</typeStatus>
|
||
: ♂s BL
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||
<quantity id="4C879B28E35A0C16BFFE8C12FA829848" box="[1232,1387,1548,1573]" metricMagnitude="-3" metricUnit="m" metricValue="2.175" metricValueMax="2.2" metricValueMin="2.15" pageId="1" pageNumber="439" unit="mm" value="2.175" valueMax="2.2" valueMin="2.15">2.15–2.2 mm</quantity>
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||
; EL
|
||
<quantity id="4C879B28E35A0C16BBB98C31FEC7982B" box="[151,302,1583,1608]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.45" metricValueMin="1.4" pageId="1" pageNumber="439" unit="mm" value="1.4249999999999998" valueMax="1.45" valueMin="1.4">1.4–1.45 mm</quantity>
|
||
; EW
|
||
<quantity id="4C879B28E35A0C16BA458C31FDEB982B" box="[363,514,1583,1608]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="1" pageNumber="439" unit="mm" value="0.825" valueMax="0.85" valueMin="0.8">0.8–0.85 mm</quantity>
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||
. ♀s BL
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||
<quantity id="4C879B28E35A0C16B9728C31FD1D982B" box="[604,756,1583,1608]" metricMagnitude="-3" metricUnit="m" metricValue="2.225" metricValueMax="2.25" metricValueMin="2.2" pageId="1" pageNumber="439" unit="mm" value="2.225" valueMax="2.25" valueMin="2.2">2.2–2.25 mm</quantity>
|
||
; EL
|
||
<quantity id="4C879B28E35A0C16B8098C31FC57982B" box="[807,958,1583,1608]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.55" metricValueMin="1.4" pageId="1" pageNumber="439" unit="mm" value="1.475" valueMax="1.55" valueMin="1.4">1.4–1.55 mm</quantity>
|
||
; EW
|
||
<quantity id="4C879B28E35A0C16B8D58C31FB7B982B" box="[1019,1170,1583,1608]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="1" pageNumber="439" unit="mm" value="0.825" valueMax="0.85" valueMin="0.8">0.8–0.85 mm</quantity>
|
||
. Dorsum (
|
||
<figureCitation id="13442A48E35A0C16BE2B8C31FAAA9824" box="[1285,1347,1583,1608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1872,1894]" captionTargetBox="[181,1409,220,1827]" captionTargetId="figure@3.[151,1436,193,1851]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pterosthetops holotypes. A) P. coriaceus sp. nov.; B) P. indwei sp. nov.; C) P. swartbergensis sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/253024/files/figure.png" pageId="1" pageNumber="439">Fig. 1</figureCitation>
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||
A) dark brown to black with a weak golden sheen, particularly around elytral margins and on head and pronotum. Maxillary palpi pitchy to black, legs pitchy to black, with tibiae somewhat lighter. Venter predominantly dark piceous, with silvery hydrofuge pubescence.
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||
</paragraph>
|
||
<paragraph id="8BC036CDE35A0C16BBE98CA2FC7F9982" blockId="1.[151,1437,1266,2030]" pageId="1" pageNumber="439">Head: Labrum transverse, rounder apicolaterally, with marked apicomedian emargination, running approx. 0.3 of length. Sides of apicomedian emargination distinctly raised. Surface of labrum dull, with strong microreticulation and short, white decumbent setae around apical and lateral margins. Entire upper surface of head dull, with strong microreticulation. Scattered white decumbent setae present, especially close to lateral margins of clypeus. Fronto-clypeal suture weak but visible; stronger laterally. Frons with a central elevation, broken by a rounded concavity between the well-marked, shining ocelli, and arcuate sulci running from just in front of each ocellus towards the lateral corners of the frontoclypeal suture, opening onto frons. Lateral margins of frons and basolateral areas of clypeus thickened, ridged beside and in front of arcuate sulci. Eyes large, occupying almost 0.5 of the lateral margins of the head; 12 convex facets in longest series.</paragraph>
|
||
<paragraph id="8BC036CDE3590C15BBE98A86FE0B9F99" blockId="2.[151,1437,151,1907]" pageId="2" pageNumber="440">
|
||
Pronotum: Distinctly cordate, with strongly explanate sides. Anterior angles obtusely rounded. Sides coarsely crenulated and weakly to sharply arcuate over anterior 2/3, then almost straight to slightly acutely rounded posterior angles (
|
||
<figureCitation id="13442A48E3590C15BA778AC1FE709E94" box="[345,409,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1872,1894]" captionTargetBox="[181,1409,220,1827]" captionTargetId="figure@3.[151,1436,193,1851]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pterosthetops holotypes. A) P. coriaceus sp. nov.; B) P. indwei sp. nov.; C) P. swartbergensis sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/253024/files/figure.png" pageId="2" pageNumber="440">Fig. 1</figureCitation>
|
||
A). Anterior margin weakly bisinuate, with very narrow hyaline border in middle, which is scarcely visible. Posterior margin weakly bisinuate to centre. Entire upper surface dull, with strong microreticulation, except on parts of pronotal reliefs where it is reduced to small punctulae. Median longitudinal sulcus foveate anteriorly and posteriorly; both foveae elongate, anterior fovea running for approx ½ of pronotal disc, posterior one for approx 1/3. Sides of sulcus bordered by ridges where reticulation is weak. Anterior and posterior admedian fovea shallow and open, with long decumbent white setae; setal sockets with anterior rim raised slightly. Adlateral fovea deeper, rounded and pit-like, particularly posteriorly, and joined by a weakly marked, shallow longitudinal channel.
|
||
</paragraph>
|
||
<paragraph id="8BC036CDE3590C15BBE98BE1FB6E9A7C" blockId="2.[151,1437,151,1907]" pageId="2" pageNumber="440">Elytra: Elongate and broadest at middle. Weakly rounded at sides from shoulder to apex. Explanate lateral margin present from shoulder to apex; widest towards apex. Apices separately rounded. Each elytron 10-seriate punctate and with two transverse depressions (saddles), one just before middle, occupying intervals 2–5, the second midway between first and apex, occupying intervals 2–4. Additional, shallow depressions present between second saddle and apex, occupying intervals 2–3, and between the two saddles, occupying intervals 5–8; deepest in intervals 6 & 7. Punctures of elytral striae each bearing a small seta. Puncture rows becoming slightly irregular in anterior saddle; random in posterior saddle, with series 2 & 3 ending in the depression between posterior saddle and apex. Intervals raised to varying degrees. Interval 1 raised along suture, particularly behind the middle. Interval 2 slightly raised in front of anterior saddle and between anterior and posterior saddles. Intervals 3 & 4 strongly raised into broad ridges, with relatively weak microsculpture, both in front of anterior saddle and between in and the posterior one. Interval 4 also raised in this manner behind posterior saddle. Interval 6 raised for some distance outside anterior saddle, then flat in region between anterior and posterior saddles, then elevated posteriorly. Interval 7 with elongate raised area adjacent to median saddle. Raised portions of elytra, and saddles, giving a somewhat tuberculate appearance. Elytal intervals with short, scattered setae. Elytral dorsal surface predominantly dull and microreticulate; less so on ridges of internal intervals, where microreticulation is weakest.</paragraph>
|
||
<paragraph id="8BC036CDE3590C15BBE98E03FD6B983F" blockId="2.[151,1437,151,1907]" pageId="2" pageNumber="440">Venter: Mentum, submentum, genae and gular region shining but strongly microreticulate, giving a coriaceous appearance. Only gular suture visible. Long, golden setae present in anteriolateral areas of mentum and smaller decumbent setae on its lateral margins. Prosternum with a broad central ridge, flanked by two semicircular depressions. Surface finely reticulate and with dense hydrofuge vestiture. Pronotal hypomeron broad, with isodiametric microreticulation and sparse, long setae just outside hypomeral carinae. Meso and metaventrites with dense hydrofuge vestiture; vestiture shorter and less dense towards lateral and hind margins. Mesoventral plaques distinct, glabrous. Median ridge splitting to form admedian ridges apically, forming an inverted Y. Border between mesoventrite and anepisternum 2 marked by a broad glabrous ridge. Metaventrite with a well-marked longitudinal depression over posterior 0.75 of sternite, with a narrow median ridge anterior to this, extending to the hind border of the mesoventrite, between mesocoxae. Elytral pseudepipleurs broad and shallowly concave; strongly microreticulate. Epipleurs narrow and ridge like; microreticulate. Abdominal ventrites 1–4 fringed with long, closely-set squamiform setae at hind margins. Abdominal ventrites 1–4 entirely covered with hydrofuge vestiture, which is denser towards anterior margins and in triangular anteriolateral patches. 5th ventrite also with hydrofuge vestiture, except for the posterior 1/5, which is glabrous and microreticulate. Entire surface of ventrite 6 also glabrous and microreticulate. Ventrites 5 & 6 with transverse row of sparse, long, adpressed hair-like setae, positioned 0.8 of the way down the sternite.</paragraph>
|
||
<paragraph id="8BC036CDE3590C15BBE98C40FDA598D3" blockId="2.[151,1437,151,1907]" pageId="2" pageNumber="440">
|
||
Aedeagus: Strongly elongate, with parameres attaching close to base and extending beyond the apex of the main piece. Main piece with asymmetrical apical tooth, and long, curved, tube-like distal lobe, extending well beyond the paramere apices. (
|
||
<figureCitation id="13442A48E3590C15BACB8CB8FDC198D3" box="[485,552,1702,1727]" captionStart="FIGURE 2" captionStartId="4.[151,250,1947,1969]" captionTargetBox="[244,1311,220,1894]" captionTargetId="figure@4.[220,1337,193,1922]" captionTargetPageId="4" captionText="FIGURE 2. Aedeagi of Pterosthetops holotypes. A) P. coriaceus sp. nov.; B) P. indwei sp. nov.; C) P. swartbergensis sp. nov. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/253025/files/figure.png" pageId="2" pageNumber="440">Fig. 2</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="8BC036CDE3590C15BBE98CD4FA83988F" blockId="2.[151,1437,151,1907]" box="[199,1386,1738,1763]" pageId="2" pageNumber="440">Female: Mesoventral plaques weaker than in males, otherwise without significant differences from males.</paragraph>
|
||
<paragraph id="8BC036CDE3590C15BBE98CF0FF0D991F" blockId="2.[151,1437,151,1907]" pageId="2" pageNumber="440">
|
||
Variation:
|
||
<typeStatus id="54C4886FE3590C15BA6F8CEEFE479964" box="[321,430,1776,1800]" pageId="2" pageNumber="440" type="paratype">Paratypes</typeStatus>
|
||
vary somewhat in the development of the golden sheen on the dorsal surface. Some appear distinctly golden; others almost matt black. There is also some variation in the depth of elytral puncture rows, and the height of intervals. Raised intervals also vary in their degree of microreticulation—from weak to strong.
|
||
</paragraph>
|
||
<caption id="DF006645E3580C14BBB98D4EFEB899E9" httpUri="https://zenodo.org/record/253024/files/figure.png" pageId="3" pageNumber="441" targetBox="[181,1409,220,1827]" targetPageId="3">
|
||
<paragraph id="8BC036CDE3580C14BBB98D4EFEB899E9" blockId="3.[151,1435,1872,1925]" pageId="3" pageNumber="441">
|
||
<emphasis id="B90BEADFE3580C14BBB98D4EFEFA9909" bold="true" box="[151,275,1872,1894]" pageId="3" pageNumber="441">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C7F4D4EE3580C14BAA58D4EFDFC9909" box="[395,533,1872,1893]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEADFE3580C14BAA58D4EFDFC9909" box="[395,533,1872,1893]" italics="true" pageId="3" pageNumber="441">Pterosthetops</emphasis>
|
||
</taxonomicName>
|
||
holotypes. A)
|
||
<taxonomicName id="4C7F4D4EE3580C14B9838D4FFCC0990A" box="[685,809,1873,1894]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="coriaceus" status="sp. nov.">
|
||
<emphasis id="B90BEADFE3580C14B9838D4FFCC0990A" box="[685,809,1873,1894]" italics="true" pageId="3" pageNumber="441">P. coriaceus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857A4E3580C14B81E8D4EFC699909" box="[816,896,1872,1893]" pageId="3" pageNumber="441" rank="species">
|
||
<emphasis id="B90BEADFE3580C14B81E8D4EFC699909" bold="true" box="[816,896,1872,1893]" pageId="3" pageNumber="441">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; B)
|
||
<taxonomicName id="4C7F4D4EE3580C14B8818D4FFBE59909" box="[943,1036,1872,1894]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="indwei" status="sp. nov.">
|
||
<emphasis id="B90BEADFE3580C14B8818D4FFBE59909" box="[943,1036,1872,1894]" italics="true" pageId="3" pageNumber="441">P. indwei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857A4E3580C14BF3D8D4EFB8A9909" box="[1043,1123,1872,1893]" pageId="3" pageNumber="441" rank="species">
|
||
<emphasis id="B90BEADFE3580C14BF3D8D4EFB8A9909" bold="true" box="[1043,1123,1872,1893]" pageId="3" pageNumber="441">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; C)
|
||
<taxonomicName id="4C7F4D4EE3580C14BFBC8D4FFAAD9909" box="[1170,1348,1872,1894]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="swartbergensis" status="sp. nov.">
|
||
<emphasis id="B90BEADFE3580C14BFBC8D4FFAAD9909" box="[1170,1348,1872,1894]" italics="true" pageId="3" pageNumber="441">P. swartbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857A4E3580C14BE658D4EFA729909" box="[1355,1435,1872,1893]" pageId="3" pageNumber="441" rank="species">
|
||
<emphasis id="B90BEADFE3580C14BE658D4EFA729909" bold="true" box="[1355,1435,1872,1893]" pageId="3" pageNumber="441">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF006645E35F0C13BBB98D85FE8D99BC" httpUri="https://zenodo.org/record/253025/files/figure.png" pageId="4" pageNumber="442" targetBox="[244,1311,220,1894]" targetPageId="4">
|
||
<paragraph id="8BC036CDE35F0C13BBB98D85FE8D99BC" blockId="4.[151,1435,1947,2000]" pageId="4" pageNumber="442">
|
||
<emphasis id="B90BEADFE35F0C13BBB98D85FEFA99DC" bold="true" box="[151,275,1947,1969]" pageId="4" pageNumber="442">FIGURE 2.</emphasis>
|
||
Aedeagi of
|
||
<taxonomicName id="4C7F4D4EE35F0C13BABE8D85FDF399DC" box="[400,538,1947,1968]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEADFE35F0C13BABE8D85FDF399DC" box="[400,538,1947,1968]" italics="true" pageId="4" pageNumber="442">Pterosthetops</emphasis>
|
||
</taxonomicName>
|
||
holotypes. A)
|
||
<taxonomicName id="4C7F4D4EE35F0C13B99F8D82FCC499DD" box="[689,813,1948,1969]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="442" phylum="Arthropoda" rank="species" species="coriaceus" status="sp. nov.">
|
||
<emphasis id="B90BEADFE35F0C13B99F8D82FCC499DD" box="[689,813,1948,1969]" italics="true" pageId="4" pageNumber="442">P. coriaceus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857A4E35F0C13B81D8D85FC6A99DC" box="[819,899,1947,1968]" pageId="4" pageNumber="442" rank="species">
|
||
<emphasis id="B90BEADFE35F0C13B81D8D85FC6A99DC" bold="true" box="[819,899,1947,1968]" pageId="4" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; B)
|
||
<taxonomicName id="4C7F4D4EE35F0C13B89F8D82FBE699DC" box="[945,1039,1947,1969]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="442" phylum="Arthropoda" rank="species" species="indwei" status="sp. nov.">
|
||
<emphasis id="B90BEADFE35F0C13B89F8D82FBE699DC" box="[945,1039,1947,1969]" italics="true" pageId="4" pageNumber="442">P. indwei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857A4E35F0C13BF3A8D85FB8C99DC" box="[1044,1125,1947,1968]" pageId="4" pageNumber="442" rank="species">
|
||
<emphasis id="B90BEADFE35F0C13BF3A8D85FB8C99DC" bold="true" box="[1044,1125,1947,1968]" pageId="4" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; C)
|
||
<taxonomicName id="4C7F4D4EE35F0C13BFBD8D82FAAC99DC" box="[1171,1349,1947,1969]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="442" phylum="Arthropoda" rank="species" species="swartbergensis" status="sp. nov.">
|
||
<emphasis id="B90BEADFE35F0C13BFBD8D82FAAC99DC" box="[1171,1349,1947,1969]" italics="true" pageId="4" pageNumber="442">P. swartbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857A4E35F0C13BE658D85FA7299DC" box="[1355,1435,1947,1968]" pageId="4" pageNumber="442" rank="species">
|
||
<emphasis id="B90BEADFE35F0C13BE658D85FA7299DC" bold="true" box="[1355,1435,1947,1968]" pageId="4" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036CDE35E0C12BBE98A89FC7E9E94" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="443">
|
||
<emphasis id="B90BEADFE35E0C12BBE98A89FE269EDC" bold="true" box="[199,463,151,176]" pageId="5" pageNumber="443">Differential diagnosis.</emphasis>
|
||
Recognized by a combination of the very strongly microreticulate dorsum (including elytra), the lack of long recumbent setae on the elytral intervals, and the two weak elytral saddles, both restricted to intervals 2–4. Male genitalia also highly characteristic (see
|
||
<figureCitation id="13442A48E35E0C12B81F8AC1FC9D9E94" box="[817,884,223,248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1947,1969]" captionTargetBox="[244,1311,220,1894]" captionTargetId="figure@4.[220,1337,193,1922]" captionTargetPageId="4" captionText="FIGURE 2. Aedeagi of Pterosthetops holotypes. A) P. coriaceus sp. nov.; B) P. indwei sp. nov.; C) P. swartbergensis sp. nov. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/253025/files/figure.png" pageId="5" pageNumber="443">Fig. 2</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656546E35E0C12BBE98B1AFA7F9F08" pageId="5" pageNumber="443" type="distribution">
|
||
<paragraph id="8BC036CDE35E0C12BBE98B1AFA7F9F08" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="443">
|
||
<emphasis id="B90BEADFE35E0C12BBE98B1AFEB19F71" bold="true" box="[199,344,260,285]" pageId="5" pageNumber="443">Distribution</emphasis>
|
||
. Known to date only the
|
||
<typeStatus id="54C4886FE35E0C12B9588B1BFD4F9F71" box="[630,678,261,285]" pageId="5" pageNumber="443">type</typeStatus>
|
||
locality, seepages on a sloping rock platform beside a stream in the southeastern part of the Cederberg range, in an area transitional between fynbos and succulent karoo biomes (
|
||
<figureCitation id="13442A48E35E0C12BE658B39FA669F2C" box="[1355,1423,295,320]" captionStart="FIGURE 9" captionStartId="21.[151,250,1909,1931]" captionTargetBox="[216,1370,978,1887]" captionTargetId="figure@21.[216,1370,978,1887]" captionTargetPageId="21" captionText="FIGURE 9. Distribution of Pterosthetops species in the Western Cape: closed white symbol P. baini sp. nov., blue symbol P. coriaceus sp. nov., green symbols P. harrisoni, open white symbols P. impressus, red triangle P. harrisoni, P. hawequas, P. impressus, P. pulcherrimus sp. nov., P. uitkyki sp. nov. & P. tuberculatus sp. nov. Basemap source: ESri, GigitalGlobe, GeoEye, i-cubed, USDA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, Swisstopo and the GIS User Community." httpUri="https://zenodo.org/record/253032/files/figure.png" pageId="5" pageNumber="443">Fig. 9</figureCitation>
|
||
).
|
||
<emphasis id="B90BEADFE35E0C12BBE98B52FEAF9F09" bold="true" box="[199,326,332,357]" pageId="5" pageNumber="443">Etymology</emphasis>
|
||
. Named in reference to the wrinkled appearance of the dorsal surface and some parts of the venter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |