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<document id="87CCB93562D2C9E29FA6CD2EFC909CE9" ID-DOI="10.1649/0010-065x-69.3.507" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106135" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175653691" checkinUser="diego" docAuthor="Miller, Kelly B. &amp; Wheeler, Quentin D." docDate="2015" docId="03D687B8FFA681630688FEF3E150D235" docLanguage="en" docName="0010-065X-69.3.507.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.507" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Zimpherus Miller and Wheeler, new genus" docType="treatment" docVersion="4" lastPageNumber="508" masterDocId="FFEFFFC0FFA781620639FFE7E24FD15A" masterDocTitle="Zimpherus nancaeMiller and Wheeler (Coleoptera: Dytiscidae: Hydroporinae: Bidessini), New Genus and New Species from Venezuela" masterLastPageNumber="511" masterPageNumber="507" pageNumber="508" updateTime="1699636595428" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5EDC82FF07D5A2E901DDA589D1D0B692">Zimpherus nancaeMiller and Wheeler (Coleoptera: Dytiscidae: Hydroporinae: Bidessini), New Genus and New Species from Venezuela</mods:title>
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<mods:namePart id="19A929C75CF18BE2B0C732DE3BD4FC75">Miller, Kelly B.</mods:namePart>
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<mods:namePart id="589124439426C3F00284C9D32285318D">Wheeler, Quentin D.</mods:namePart>
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<mods:title id="7A9D0DA3B56003FECB1914FCA40DBEDD">The Coleopterists Bulletin</mods:title>
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<mods:date id="3969060ED1C99FACEA3467449B5208BC">2015</mods:date>
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<mods:number id="86AF787D9E9A0A40E0190CB9754576B3">2015-09-30</mods:number>
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<mods:identifier id="F7BA72C2BC89BE894C1DF77DDD191279" type="DOI">10.1649/0010-065x-69.3.507</mods:identifier>
<mods:identifier id="D747704447F178415ECD6BBD240F1A20" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03D687B8FFA681630688FEF3E150D235" ID-DOI="http://doi.org/10.5281/zenodo.5413267" ID-Zenodo-Dep="5413267" LSID="urn:lsid:plazi:treatment:03D687B8FFA681630688FEF3E150D235" httpUri="http://treatment.plazi.org/id/03D687B8FFA681630688FEF3E150D235" lastPageNumber="508" pageId="1" pageNumber="508">
<subSubSection id="C3656525FFA681630688FEF3E38FD01E" pageId="1" pageNumber="508" type="nomenclature">
<paragraph id="8BC036AEFFA681630688FEF3E38FD01E" blockId="1.[177,612,276,325]" pageId="1" pageNumber="508">
<emphasis id="B90BEABCFFA681630688FEF3E02BD073" bold="true" box="[177,612,276,298]" pageId="1" pageNumber="508">
<taxonomicName id="4C7F4D2DFFA681630688FEF3E3A2D073" ID-CoL="8LZ9S" authority="Miller and Wheeler" authorityName="Miller and Wheeler" box="[177,493,276,298]" class="Insecta" family="Dytiscidae" genus="Zimpherus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BEABCFFA681630688FEF3E353D070" bold="true" box="[177,284,276,298]" italics="true" pageId="1" pageNumber="508">Zimpherus</emphasis>
Miller and Wheeler
</taxonomicName>
,
<taxonomicNameLabel id="A23857C7FFA6816307C0FEF3E02BD073" box="[505,612,276,297]" pageId="1" pageNumber="508" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13442A2BFFA681630765FEC8E3F6D01E" box="[348,441,303,325]" captionStart-0="Fig" captionStart-1="Figs" captionStart-2="Fig" captionStartId-0="1.[175,207,1686,1705]" captionStartId-1="2.[202,242,1633,1652]" captionStartId-2="3.[175,207,528,547]" captionTargetBox-0="[151,637,941,1663]" captionTargetBox-1="[173,1193,183,1609]" captionTargetBox-2="[151,637,194,505]" captionTargetId-0="figure-596@1.[151,637,941,1663]" captionTargetId-1="figure-1@2.[173,1193,183,1609]" captionTargetId-2="figure-691@3.[151,637,194,505]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" captionText-1="Figs. 2–6. Zimpherus nancae. 2–4) Male genitalia: 2) Median lobe, right lateral aspect; 3) Right lateral lobe, right lateral aspect; 4) Median lobe, ventral aspect; 5) Left metatrochanter, anterior aspect. 6) Prosternal process. Scale bars: A = 0.5 mm for Fig. 5; B = 0.2 mm for Fig. 6." captionText-2="Fig. 7. Zimpherus nancae, distribution (type locality)." figureDoi-0="http://doi.org/10.5281/zenodo.10106137" figureDoi-1="http://doi.org/10.5281/zenodo.10106139" figureDoi-2="http://doi.org/10.5281/zenodo.10106141" httpUri-0="https://zenodo.org/record/10106137/files/figure.png" httpUri-1="https://zenodo.org/record/10106139/files/figure.png" httpUri-2="https://zenodo.org/record/10106141/files/figure.png" pageId="1" pageNumber="508">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656525FFA681630696FEBEE018D0D3" pageId="1" pageNumber="508" type="type_taxon">
<paragraph id="8BC036AEFFA681630696FEBEE018D0D3" blockId="1.[151,638,345,871]" pageId="1" pageNumber="508">
<emphasis id="B90BEABCFFA681630696FEBEE377D034" bold="true" box="[175,312,345,366]" pageId="1" pageNumber="508">
<typeStatus id="54C4880CFFA681630696FEBEE290D034" box="[175,223,345,366]" pageId="1" pageNumber="508">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C7F4D2DFFA68163077FFEBEE2A6D0D3" ID-CoL="8LZ7R" authority="Miller and Wheeler" authorityName="MILLER AND WHEELER" authorityYear="2014" class="Insecta" family="Dytiscidae" genus="Zimpherus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="nancae" status="sp. nov.">
<emphasis id="B90BEABCFFA68163077FFEBEE049D035" box="[326,518,345,367]" italics="true" pageId="1" pageNumber="508">Zimpherus nancae</emphasis>
Miller and Wheeler
</taxonomicName>
,
<taxonomicNameLabel id="A23857C7FFA6816306CCFE93E324D0D3" box="[245,363,372,393]" pageId="1" pageNumber="508" rank="species">new species</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C3656525FFA681630696FE68E6C5D33C" pageId="1" pageNumber="508" type="diagnosis">
<paragraph id="8BC036AEFFA681630696FE68E6C5D33C" blockId="1.[151,638,345,871]" lastBlockId="1.[685,1173,193,879]" pageId="1" pageNumber="508">
<emphasis id="B90BEABCFFA681630696FE68E357D0FE" bold="true" box="[175,280,399,420]" pageId="1" pageNumber="508">Diagnosis.</emphasis>
This species is diagnosed within Bidessini by: 1) a transverse occipital line on the head absent (
<figureCitation id="13442A2BFFA68163072CFE23E302D083" box="[277,333,452,473]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
); 2) the anterior clypeal margin unmodified (
<figureCitation id="13442A2BFFA68163072CFE39E300D0A9" box="[277,335,478,499]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
); 3) a transverse carina across the elytral epipleuron at the humeral angle absent (
<figureCitation id="13442A2BFFA6816306A6FDF4E296D372" box="[159,217,531,552]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
); 4) basal elytral stria present (
<figureCitation id="13442A2BFFA68163042FFDF4E01ED372" box="[534,593,531,552]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
); 5) elytral sutural stria absent (
<figureCitation id="13442A2BFFA68163079CFDC9E390D319" box="[421,479,558,579]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
); 6) the elytron without rows of punctures (
<figureCitation id="13442A2BFFA68163079FFDAEE390D304" box="[422,479,585,606]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
); 7) the median lobe of the male aedeagus apically extremely complex (
<figureCitation id="13442A2BFFA6816306EBFD99E379D3C9" box="[210,310,638,659]" captionStart="Figs" captionStartId="2.[202,242,1633,1652]" captionTargetBox="[173,1193,183,1609]" captionTargetId="figure-1@2.[173,1193,183,1609]" captionTargetPageId="2" captionText="Figs. 2–6. Zimpherus nancae. 2–4) Male genitalia: 2) Median lobe, right lateral aspect; 3) Right lateral lobe, right lateral aspect; 4) Median lobe, ventral aspect; 5) Left metatrochanter, anterior aspect. 6) Prosternal process. Scale bars: A = 0.5 mm for Fig. 5; B = 0.2 mm for Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.10106139" httpUri="https://zenodo.org/record/10106139/files/figure.png" pageId="1" pageNumber="508">Figs. 2, 3</figureCitation>
); and 8) the lateral lobes of the male aedeagus with two segments and apically rounded (
<figureCitation id="13442A2BFFA6816306CCFD54E37DD392" box="[245,306,691,712]" captionStart="Figs" captionStartId="2.[202,242,1633,1652]" captionTargetBox="[173,1193,183,1609]" captionTargetId="figure-1@2.[173,1193,183,1609]" captionTargetPageId="2" captionText="Figs. 2–6. Zimpherus nancae. 2–4) Male genitalia: 2) Median lobe, right lateral aspect; 3) Right lateral lobe, right lateral aspect; 4) Median lobe, ventral aspect; 5) Left metatrochanter, anterior aspect. 6) Prosternal process. Scale bars: A = 0.5 mm for Fig. 5; B = 0.2 mm for Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.10106139" httpUri="https://zenodo.org/record/10106139/files/figure.png" pageId="1" pageNumber="508">Fig. 4</figureCitation>
). The genus is similar to
<taxonomicName id="4C7F4D2DFFA68163040BFD55E037D392" authorityName="Guignot" authorityYear="1939" box="[562,632,690,712]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA68163040BFD55E037D392" box="[562,632,690,712]" italics="true" pageId="1" pageNumber="508">Uvarus</emphasis>
</taxonomicName>
, but differs in the body shape and the shape of the male aedeagus. The overall body shape of
<taxonomicName id="4C7F4D2DFFA6816306AEFCE5E2B2D242" box="[151,253,770,792]" class="Insecta" family="Dytiscidae" genus="Zimpherus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA6816306AEFCE5E2B2D242" box="[151,253,770,792]" italics="true" pageId="1" pageNumber="508">Zimpherus</emphasis>
</taxonomicName>
specimens is narrow and anteriorly narrowed with the head somewhat deflexed ventrad and the lateral outline distinctly discontinuous between the pronotum and elytron (
<figureCitation id="13442A2BFFA6816307A1FCB5E39FD23D" box="[408,464,850,871]" captionStart="Fig" captionStartId="1.[175,207,1686,1705]" captionTargetBox="[151,637,941,1663]" captionTargetId="figure-596@1.[151,637,941,1663]" captionTargetPageId="1" captionText="Fig. 1. Zimpherus nancae, dorsal habitus. Scale bar =" figureDoi="http://doi.org/10.5281/zenodo.10106137" httpUri="https://zenodo.org/record/10106137/files/figure.png" pageId="1" pageNumber="508">Fig. 1</figureCitation>
). The body shape in
<taxonomicName id="4C7F4D2DFFA6816304F0FF26E15FD18D" authorityName="Guignot" authorityYear="1939" box="[713,784,193,215]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA6816304F0FF26E15FD18D" box="[713,784,193,215]" italics="true" pageId="1" pageNumber="508">Uvarus</emphasis>
</taxonomicName>
specimens is broadly to elongate oval and the lateral outline is approximately continuous between the pronotum and elytron. The median lobe is apically multifurcated and multilobed in
<taxonomicName id="4C7F4D2DFFA681630494FECBE15FD018" box="[685,784,300,322]" class="Insecta" family="Dytiscidae" genus="Zimpherus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA681630494FECBE15FD018" box="[685,784,300,322]" italics="true" pageId="1" pageNumber="508">Zimpherus</emphasis>
</taxonomicName>
(
<figureCitation id="13442A2BFFA681630526FECBE136D01B" box="[799,889,300,321]" captionStart="Figs" captionStartId="2.[202,242,1633,1652]" captionTargetBox="[173,1193,183,1609]" captionTargetId="figure-1@2.[173,1193,183,1609]" captionTargetPageId="2" captionText="Figs. 2–6. Zimpherus nancae. 2–4) Male genitalia: 2) Median lobe, right lateral aspect; 3) Right lateral lobe, right lateral aspect; 4) Median lobe, ventral aspect; 5) Left metatrochanter, anterior aspect. 6) Prosternal process. Scale bars: A = 0.5 mm for Fig. 5; B = 0.2 mm for Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.10106139" httpUri="https://zenodo.org/record/10106139/files/figure.png" pageId="1" pageNumber="508">Figs. 2, 3</figureCitation>
), and the lateral lobe is broad with the apex rounded, without a small, curved, tooth–shaped apical lobe (
<figureCitation id="13442A2BFFA68163059CFE86E1AFD02C" box="[933,992,353,374]" captionStart="Figs" captionStartId="2.[202,242,1633,1652]" captionTargetBox="[173,1193,183,1609]" captionTargetId="figure-1@2.[173,1193,183,1609]" captionTargetPageId="2" captionText="Figs. 2–6. Zimpherus nancae. 2–4) Male genitalia: 2) Median lobe, right lateral aspect; 3) Right lateral lobe, right lateral aspect; 4) Median lobe, ventral aspect; 5) Left metatrochanter, anterior aspect. 6) Prosternal process. Scale bars: A = 0.5 mm for Fig. 5; B = 0.2 mm for Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.10106139" httpUri="https://zenodo.org/record/10106139/files/figure.png" pageId="1" pageNumber="508">Fig. 4</figureCitation>
). A simple, evenly curved male median apical lobe is characteristic of
<taxonomicName id="4C7F4D2DFFA681630494FE71E0B9D0F6" authorityName="Guignot" authorityYear="1939" box="[685,758,406,428]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA681630494FE71E0B9D0F6" box="[685,758,406,428]" italics="true" pageId="1" pageNumber="508">Uvarus</emphasis>
</taxonomicName>
. The male median lobe of members of
<taxonomicName id="4C7F4D2DFFA681630494FE56E16CD09D" authorityName="Young" authorityYear="1967" box="[685,803,433,455]" class="Insecta" family="Dytiscidae" genus="Microdessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA681630494FE56E16CD09D" box="[685,803,433,455]" italics="true" pageId="1" pageNumber="508">Microdessus</emphasis>
</taxonomicName>
is also apically divided into multiple lobes (four), but
<taxonomicName id="4C7F4D2DFFA681630575FE2CE646D0BB" box="[844,1033,459,481]" class="Insecta" family="Dytiscidae" genus="Microdessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90BEABCFFA681630575FE2CE1F0D0BB" box="[844,959,459,481]" italics="true" pageId="1" pageNumber="508">Microdessus</emphasis>
species
</taxonomicName>
are very small (
<emphasis id="B90BEABCFFA68163048CFE01E08CD0A6" box="[693,707,486,508]" italics="true" pageId="1" pageNumber="508">∼</emphasis>
<quantity id="4C879B4BFFA6816304FBFE01E141D0A1" box="[706,782,486,507]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="508" unit="mm" value="1.5">1.5 mm</quantity>
), do not have an unusually offset metatrochanter, and are not overall uniquely shaped as are
<taxonomicName id="4C7F4D2DFFA681630494FDFCE157D36B" box="[685,792,539,561]" class="Insecta" family="Dytiscidae" genus="Zimpherus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA681630494FDFCE157D36B" box="[685,792,539,561]" italics="true" pageId="1" pageNumber="508">Zimpherus</emphasis>
</taxonomicName>
specimens.
<taxonomicName id="4C7F4D2DFFA6816305A5FDFCE624D36A" box="[924,1131,539,561]" class="Insecta" family="Dytiscidae" genus="Microdessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90BEABCFFA6816305A5FDFCE656D36B" box="[924,1049,539,561]" italics="true" pageId="1" pageNumber="508">Microdessus</emphasis>
species
</taxonomicName>
are somewhat flattened dorsoventrally and elongate oval with the lateral margins continuously curved.
</paragraph>
</subSubSection>
<subSubSection id="C3656525FFA6816304FCFD8CE17BD3EF" pageId="1" pageNumber="508" type="etymology">
<paragraph id="8BC036AEFFA6816304FCFD8CE17BD3EF" blockId="1.[685,1173,193,879]" pageId="1" pageNumber="508">
<emphasis id="B90BEABCFFA6816304FCFD8CE177D3DA" bold="true" box="[709,824,619,640]" pageId="1" pageNumber="508">Etymology.</emphasis>
The genus is named in honor of Nancy Zimpher, Chancellor of the State University of
<collectingRegion id="49BBF84CFFA6816304F0FD47E17FD3EF" box="[713,816,672,693]" country="United States of America" name="New York" pageId="1" pageNumber="508">New York</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656525FFA6816304FCFD5CE150D235" pageId="1" pageNumber="508" type="discussion">
<paragraph id="8BC036AEFFA6816304FCFD5CE150D235" blockId="1.[685,1173,193,879]" pageId="1" pageNumber="508">
<emphasis id="B90BEABCFFA6816304FCFD5CE178D38A" bold="true" box="[709,823,699,720]" pageId="1" pageNumber="508">Discussion.</emphasis>
This genus bears similarity with both
<taxonomicName id="4C7F4D2DFFA6816304E7FD32E16BD3B1" authorityName="Guignot" authorityYear="1939" box="[734,804,725,747]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA6816304E7FD32E16BD3B1" box="[734,804,725,747]" italics="true" pageId="1" pageNumber="508">Uvarus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D2DFFA68163056DFD32E185D3B1" authorityName="Young" authorityYear="1967" box="[852,970,725,747]" class="Insecta" family="Dytiscidae" genus="Microdessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEABCFFA68163056DFD32E185D3B1" box="[852,970,725,747]" italics="true" pageId="1" pageNumber="508">Microdessus</emphasis>
</taxonomicName>
in its basic character combination but differs from each (see Diagnosis). Nevertheless, it is not without some misgivings that the new species described here is placed within a new genus given the expanding number of genera in the tribe.
</paragraph>
</subSubSection>
</treatment>
</document>
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