treatments-xml/data/03/D6/87/03D687AFA128FFE86AB5FF3BFBCC8004.xml

<document id="0CB00768160B2F22ACCDA124028831F2" ID-DOI="10.37520/aemnp.2021.022" ID-ISSN="1804-6487" ID-Zenodo-Dep="5820961" ID-ZooBank="32519CC9-3658-469A-926D-6A1EBEE0FC59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641388598144" checkinUser="plazi" docAuthor="Yamada, Kazutaka &amp; Yasunaga, Tomohide" docDate="2021" docId="03D687AFA128FFE86AB5FF3BFBCC8004" docLanguage="en" docName="ActaEntMusNatPra.61.2.375-426.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Anthocoris japonicus Poppius 1909" docType="treatment" docVersion="3" lastPageNumber="399" masterDocId="FFEFFFD7A13FFFF06A4AFF85FF9D8251" masterDocTitle="The tribe Anthocorini in Japan (Hemiptera: Anthocoridae): descriptions of new species, review of distribution and bionomics" masterLastPageNumber="426" masterPageNumber="375" pageNumber="398" updateTime="1699285380998" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="F19DA8785B6C5B34567EB87964FCDA62" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D129F658E98C93815C104909456A584E">
<mods:title id="E36C66934F75EA5027891B41D5AF2C09">The tribe Anthocorini in Japan (Hemiptera: Anthocoridae): descriptions of new species, review of distribution and bionomics</mods:title>
</mods:titleInfo>
<mods:name id="295512E2FFBE1F27AB7CF8487190A324" type="personal">
<mods:role id="40691C2E8DDFC6586ECB7FD48FC6C2E5">
<mods:roleTerm id="38B52ABE33F803A5F7F1027F71D85313">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2202E967F2E15A615AD101FDEFC7082D">Yamada, Kazutaka</mods:namePart>
<mods:affiliation id="E38A7AD5469E38B22E8FEDF1A8B7E359">Tokushima Prefectural Museum, Bunka-no-Mori Park, Mukôterayama, Hachiman-chô, Tokushima 770 - 8070, Japan;</mods:affiliation>
</mods:name>
<mods:name id="57AFF152D9722EEABAF7116A15FB9D8A" type="personal">
<mods:role id="9376C299D83D370CBF696A267A515AAF">
<mods:roleTerm id="B7C90A9B8AE40C08CF6750346B22F0F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A8C3313F9BE15336E31A6999BE0DC757">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="8D84308C0A43460D648762FAF92AAE32">Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, New York, c / o Nameshi</mods:affiliation>
</mods:name>
<mods:typeOfResource id="095049C619BB9D3DB4414B8400AC1B5D">text</mods:typeOfResource>
<mods:relatedItem id="B505715DB47AB998DAC43F90A0EBE31F" type="host">
<mods:titleInfo id="7E171AD29B036A67DDE042B3BFE9DE0F">
<mods:title id="36A856D96E343C28FBC3C74475E7EB92">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="D06967621822EDD6DFA06679089B67A5">
<mods:date id="7261065D588E8619BDF7B9B6C90F7A0A">2021</mods:date>
<mods:detail id="D7BC9D50421C852428F81AA920579C23" type="pubDate">
<mods:number id="2C449523457EDB1257974D43AB8651FF">2021-10-19</mods:number>
</mods:detail>
<mods:detail id="E92AB6AB00449DF28C36BBB5239F5A67" type="volume">
<mods:number id="0B344DE5650BF5F74201FA05F27088F3">61</mods:number>
</mods:detail>
<mods:detail id="598C62E4FB2E5965CA70BBD2C18A1473" type="issue">
<mods:number id="5AAEE4DAE441FD380E75446704C36332">2</mods:number>
</mods:detail>
<mods:extent id="033E14D10DE9529C39569E377DFB45E5" unit="page">
<mods:start id="20C2D6F676EC84AC2D3D944EFC162144">375</mods:start>
<mods:end id="0085D27EB67B2D1DC3EC7D6F2352BDA6">426</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D92871930518E1CA083665D1E17C5318">journal article</mods:classification>
<mods:identifier id="0B98E3D8DEA1D030C71F09C04CA7EFCC" type="DOI">10.37520/aemnp.2021.022</mods:identifier>
<mods:identifier id="667D3F5EF552576811BD608F0F5F85DD" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="354FBD2A95D4742D8B4B2CA33188C4B2" type="Zenodo-Dep">5820961</mods:identifier>
<mods:identifier id="0AA4D79FCCADDDE3A8FF228A015E6197" type="ZooBank">32519CC9-3658-469A-926D-6A1EBEE0FC59</mods:identifier>
</mods:mods>
<treatment id="03D687AFA128FFE86AB5FF3BFBCC8004" LSID="urn:lsid:plazi:treatment:03D687AFA128FFE86AB5FF3BFBCC8004" httpUri="http://treatment.plazi.org/id/03D687AFA128FFE86AB5FF3BFBCC8004" lastPageId="24" lastPageNumber="399" pageId="23" pageNumber="398">
<subSubSection id="C3656532A128FFE76AB5FF3BFD128284" box="[255,655,190,213]" pageId="23" pageNumber="398" type="nomenclature">
<paragraph id="8BC036B9A128FFE76AB5FF3BFD128284" blockId="23.[255,655,190,213]" box="[255,655,190,213]" pageId="23" pageNumber="398">
<heading id="D08881D5A128FFE76AB5FF3BFD128284" box="[255,655,190,213]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="398" reason="3">
<taxonomicName id="4C7F4D3AA128FFE76AB5FF3BFD128284" authority="Poppius, 1909" authorityName="Poppius" authorityYear="1909" box="[255,655,190,213]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="398" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90BEAABA128FFE76AB5FF3BFD128284" bold="true" box="[255,655,190,213]" pageId="23" pageNumber="398">
<emphasis id="B90BEAABA128FFE76AB5FF3BFE778284" bold="true" box="[255,490,190,213]" italics="true" pageId="23" pageNumber="398">Anthocoris japonicus</emphasis>
<bibRefCitation id="EFEE4B48A128FFE76BBBFF3BFD128284" author="POPPIUS B." box="[497,655,190,213]" pageId="23" pageNumber="398" pagination="1 - 43" refId="ref58375" refString="POPPIUS B. 1909: Beitrage zur Kenntnis der Anthocoriden. Acta Societatis Scientiarum Fennicae 37 (9): 1 - 43." type="journal article" year="1909">Poppius, 1909</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656532A128FFE76AAAFF59FD3382BE" box="[224,686,220,239]" pageId="23" pageNumber="398" type="description">
<paragraph id="8BC036B9A128FFE76AAAFF59FD3382BE" blockId="23.[151,761,220,2075]" box="[224,686,220,239]" pageId="23" pageNumber="398">
(
<figureCitation id="13442A3CA128FFE76AACFF59FEDD82BE" box="[230,320,220,239]" captionStart="Fig" captionStartId="15.[151,182,2010,2029]" captionTargetBox="[211,1313,201,1965]" captionTargetId="figure-13@15.[179,1408,189,1996]" captionTargetPageId="15" captionText="Fig. 7. Habitus of Anthocoris spp., dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A–B – A. chibi Hiura, 1959, male; C–D – A. confusus Reuter, 1884, male; E–F – A. japonicus Poppius, 1909, male; G–H – A. kalopanacis Kerzhner, 1977, female; I–J – A. nemoralis (Fabricius, 1794), male; K–L – A. takahashii Hiura, 1959, male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820983" httpUri="https://zenodo.org/record/5820983/files/figure.png" pageId="23" pageNumber="398">Figs 7E–F</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76B03FF59FEF982BE" box="[329,356,220,239]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="23" pageNumber="398">9C</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76B26FF59FE3882BE" box="[364,421,220,239]" captionStart="Fig" captionStartId="19.[151,182,2012,2031]" captionTargetBox="[151,1433,563,1999]" captionTargetId="figure-13@19.[151,1436,562,2003]" captionTargetPageId="19" captionText="Fig. 10. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. chibi Hiura, 1959; C–D – A. confusus Reuter, 1884 (from India); E–F – A. japonicus Poppius, 1909.A, C, E – pygophore with paramere, dorsal (A, C) and laterodorsal (E) views; B, D, F – paramere, dorsal (B, D) and laterodorsal (F) views. Abbreviations: end ‒ endosoma; prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820995" httpUri="https://zenodo.org/record/5820995/files/figure.png" pageId="23" pageNumber="398">10E–F</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76BE4FF59FE4E82BE" box="[430,467,220,239]" captionStart="Fig" captionStartId="22.[151,182,1996,2015]" captionTargetBox="[236,1400,657,1963]" captionTargetId="figure-13@22.[151,1436,648,1972]" captionTargetPageId="22" captionText="Fig. 12. Abdominal sterna II–III of Anthocoris spp., male, ventral view (setae omitted). A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius, 1794); E – A. takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype. Abbreviations: ma ‒ membranous area; st2–3 ‒ abdominal sternum II to III. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5820999" httpUri="https://zenodo.org/record/5820999/files/figure.png" pageId="23" pageNumber="398">12C</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76B96FF59FD8882BE" box="[476,533,220,239]" captionStart="Fig" captionStartId="25.[151,182,2008,2027]" captionTargetBox="[203,1430,205,1962]" captionTargetId="figure-13@25.[151,1436,186,1980]" captionTargetPageId="25" captionText="Fig. 13. Male genitalia of Anthocoris spp. A–C – A. chibi Hiura, 1959; D–F – A. confusus Reuter, 1884; G–I – A. japonicus Poppius, 1909; J–L – A. nemoralis (Fabricius, 1794); M–O – A. takahashii Hiura, 1959. A, D, G, J, M – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F, H–I, K–L, N–O – paramere, two different views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821001" httpUri="https://zenodo.org/record/5821001/files/figure.png" pageId="23" pageNumber="398">13G–I</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76854FF59FDDE82BE" box="[542,579,220,239]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="23" pageNumber="398">15C</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76806FF59FD1482BE" box="[588,649,220,239]" captionStart="Fig" captionStartId="33.[151,182,1974,1993]" captionTargetBox="[151,1436,189,1804]" captionTargetId="figure-13@33.[151,1437,186,1824]" captionTargetPageId="33" captionText="Fig. 17. Habitats, overwintering individuals, and mating pair of Anthocoris spp. A–B – Field survey on hibernating anthocorids under bark-flakes of Zelkova serrata at urbanized zone of Nagasaki City (Kawaguchi Park) in early February; C – A. japonicus Poppius, 1909, overwintering adults under the bark-flake of Z. serrata; D–E – same, mating pair; F – A. miyamotoi Hiura, 1959, adult males and final instar immature (right), overwintering individuals under the bark of Z. serrata; G – same, active male adult on flower of Camellia sasanqua in winter (observed on Jan. 4, 2020)." figureDoi="http://doi.org/10.5281/zenodo.5821013" httpUri="https://zenodo.org/record/5821013/files/figure.png" pageId="23" pageNumber="398">17C–E</figureCitation>
, 
<figureCitation id="13442A3CA128FFE768D8FF59FD3582BE" box="[658,680,220,239]" captionStart="Fig" captionStartId="38.[153,184,2034,2053]" captionTargetBox="[168,1419,673,1984]" captionTargetId="figure-299@38.[153,1434,654,2003]" captionTargetPageId="38" captionText="Fig. 21. Distribution of Anthocoris takahashii Hiura, 1959 (circles), A. japonicus Poppius, 1909 (triangles), and A. kalopanacis Kerzhner, 1977 (squares) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.5821022" httpUri="https://zenodo.org/record/5821022/files/figure.png" pageId="23" pageNumber="398">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656532A128FFE76ADDFF7BFDE8854D" pageId="23" pageNumber="398" type="reference_group">
<paragraph id="8BC036B9A128FFE76ADDFF7BFEF6837D" blockId="23.[151,761,220,2075]" pageId="23" pageNumber="398">
<treatmentCitationGroup id="AB6F1197A128FFE76ADDFF7BFEF6837D" pageId="23" pageNumber="398">
<treatmentCitation id="0ADE10A8A128FFE76ADDFF7BFE698340" author="POPPIUS B." box="[151,500,254,273]" page="33" pageId="23" pageNumber="398" year="1909">
<taxonomicName id="4C7F4D3AA128FFE76ADDFF7BFE698340" authority="Poppius, 1909: 33" authorityName="Poppius" authorityPageNumber="33" authorityYear="1909" box="[151,500,254,273]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="398" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90BEAABA128FFE76ADDFF7BFED38340" box="[151,334,254,273]" italics="true" pageId="23" pageNumber="398">Anthocoris japonicus</emphasis>
<materialsCitation id="3B173CE4A128FFE76B1EFF7BFE698340" box="[340,500,254,273]" collectionCode="HNHM" country="Japan" location="Honshu" pageId="23" pageNumber="398" specimenCount="1" specimenCount-female="1" stateProvince="Kanagawa" typeStatus="holotype">
<bibRefCitation id="EFEE4B48A128FFE76B1EFF7BFE698340" author="POPPIUS B." box="[340,500,254,273]" pageId="23" pageNumber="398" pagination="1 - 43" refId="ref58375" refString="POPPIUS B. 1909: Beitrage zur Kenntnis der Anthocoriden. Acta Societatis Scientiarum Fennicae 37 (9): 1 - 43." type="journal article" year="1909">Poppius, 1909: 33</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="54C4881BA128FFE76BB7FF7BFDCD8340" box="[509,592,254,273]" pageId="23" pageNumber="398" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D79FD30A128FFE76810FF7DFDF58340" box="[602,616,248,273]" pageId="23" pageNumber="398" type="female">♀</specimenCount>
, 
<collectingCountry id="F3687629A128FFE7683EFF7BFD3A8340" box="[628,679,254,273]" name="Japan" pageId="23" pageNumber="398">Japan</collectingCountry>
, 
<location id="8EA06062A128FFE768FAFF7BFD688340" LSID="urn:lsid:plazi:treatment:03D687AFA128FFE86AB5FF3BFBCC8004:8EA06062A128FFE768FAFF7BFD688340" box="[688,757,254,273]" country="Japan" name="Honshu" pageId="23" pageNumber="398" stateProvince="Kanagawa">Honshu</location>
, 
<collectingRegion id="49BBF85BA128FFE76AFFFE9CFE92837D" box="[181,271,281,300]" country="Japan" name="Kanagawa" pageId="23" pageNumber="398">Kanagawa</collectingRegion>
(HNHM).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDFEB6FDE8854D" blockId="23.[151,761,220,2075]" pageId="23" pageNumber="398">
<taxonomicName id="4C7F4D3AA128FFE76ADDFEB6FED48317" authorityName="Poppius" authorityYear="1909" box="[151,329,307,326]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="398" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90BEAABA128FFE76ADDFEB6FED48317" box="[151,329,307,326]" italics="true" pageId="23" pageNumber="398">Anthocoris japonicus</emphasis>
</taxonomicName>
(selected references): EඌൺKං (1950): 257 (redescription, habitus, distribution); Mංඒൺආඈඍඈ (1957): 76 (ovariole number); EඌൺKං et al. (1959): 105 (description of fifth instar nymph, habitus, habitat); Hංඎඋൺ (1959): 4, pls. 3–4 (distribution, habitat, phenology, figures); TൺKൺKඎඋൺ (1959): 14 (record, habitat); SඍංർHൾඅ (1960): 357 (listed, distribution); Mංඒൺආඈඍඈ (1965): 96, pl.48 (diagnosis, habitat, prey, distribution, photo); Mංඒൺආඈඍඈ &amp; Lൾൾ (1966):375–376 (record); Lൾൾ (1971):358–359, 552 (redescription, photo, record, distribution); Hංඎඋൺ (1977): pl. 32, 122 (photo, bionomics); Nඈඓൺඐൺ (1978): 373 (record); Fඈඋൽ (1979): 56 (listed, distribution); IඐൺඌൺKං (1983): 14 (record); TൺർHංKൺඐൺ (1983):106, 255 (photo, diagnosis, bionomics); HൺඒൺඌHං (1984): 427 (record); Kංආ &amp; Nൺආ (1984): 309 (record); TൺKൺKඎඋൺ (1984): 5 (record); Lൾൾ et al. (1985): 364 (record); Sൺඌൺඃං (1985): 58 (record); OඍඌඎKൺ &amp; YඈඌHංඓൺKං (1987): 596 (record); IർHංඍൺ (1988): 132 (record); KൾඋඓHඇൾඋ (1988): 773–774 (figure, in key); Mංඒൺආඈඍඈ &amp; YൺඌඎඇൺGൺ (1989): 165 (listed, distribution); AඌൺඈKൺ &amp; IൾKං (1990): 155 (listed, distribution); Yඈඈඇ et al. (1990): 106 (record); ZHൾඇG &amp; Bඎ (1990): 24 (record); Bඎ &amp; ZHൾඇG (1991a): 94, 98 (listed); Lൾൾ &amp; Kඐඈඇ (1991): 13 (record); Lൾൾ et al. (1991): 39–40 (record); Kංආ (1993): 290 (record); Lൾൾ et al. (1993): 10 (record); Nඈඓൺඐൺ (1993): 6 (record); OඍඌඎKൺ (1993):70 (listed); IർHංඍൺ (1994): 201 (listed); Lൾൾ &amp; Kඐඈඇ (1994): 63 (listed); YඈඌHංඍඈආං (1994): 25–26 (record); Kංආ (1995): 216 (listed); OඍඌඎKൺ (1995): 45 (record); OඍඌඎKൺ (1996): 282 (record); Aඇ (1996): 28 (record); Kඐඈඇ et al. (1996a): 109 (listed); Pඣඋංർൺඋඍ (1996): 112 (catalogue, distribution); NൺKൺආඎඋൺ et al. (1997): 163 (distribution); Sඈඇඈൻൾ (1997):54 (record, habitat);HൺඒൺඌHං (1998): 165 (record);KංඌHංආඈඍඈ et al. (1998): 84 (listed); YൺඌඎඇൺGൺ (1999): 22 (listed, distribution); Hඎൺ (2000): 198 (listed, distribution); OඌൺKൺ Pඋൾൿൾർඍඎඋൾ (2000): 62 (listed, distribution); Yൺඇඈ &amp; Yൺආൺආඈඍඈ (2000): 168 (listed); Bඎ &amp; ZHൾඇG (2001): 121, 160 (in key, redescription, figures); Iඐൺඍൾ Pඋൾൿൾർඍඎඋൾ (2001): 179 (listed); Kඐඈඇ et al. (2001): 79 (catalogue, record,distribution);YൺඌඎඇൺGൺ (2001b): pl. 85, 281 (photo, diagnosis, habitat, phenology); Mൺඋඎඒൺආൺ et al. (2000): 252 (record); SඎඓඎKං (2003): 147 (distribution); KൺGൺඐൺ &amp; HංGඎർHං (2003): 205 (record, distribution); HൺඒൺඌHං &amp; OඓൺKං (2004):223 (distribution); KൺඇඒඎKඈ- ඏൺ &amp; MൺඋඎඌංK (2006): 168 (listed); TൺGඈ (2006): 29 (record); Kൾ &amp; Bඎ (2007):91–92 (figure, description of female genitalia);TൺർHංKൺඐൺ (2009): 4 (record); Mංඒൺආඈඍඈ (2008): 164, pl. 57 (diagnosis, habitat, prey, distribution, photo); IർHංඍൺ (2009): 68 (listed); VංඇඈKඎඋඈඏ et al. (2010): 56 (catalogue, distribution); TඈඋංGඈൾ (2011): 69 (record); YൺආൺඓൺKං &amp; TൺKൺKඎඋൺ (2011):79 (record); UඌHංඋඈKංඍൺ et al. (2012): 16 (record); AඎKൾආൺ et al. (2013a): 85 (catalogue, distribution); Eඇඃඎ et al. (2013): 245 (photo, distribution, habitat); Hൺඈ &amp; Mൺ (2013): 36 (listed); JඎඇG et al. (2013): 422 (catalogue, diagnosis, distribution, habitat); IඐൺඌൺKං (2014): 24 (record); SඎඓඎKං (2014a): 7 (distribution); MൺൾHൺඋൺ (2015): 28–29 (record, distribution, habitat, phenology); YൺඓൺKං (2015): 144 (record); NඈඓൺKං et al. (2016): 82 (record); Yൺආൺൽൺ et al. (2016): 422 (catalogue, distribution); JඎඇG &amp; Lൾൾ (2017): 36–37, 70 (diagnosis, redescription, associated plant, prey, distribution, photo, figure); Iඍඈ &amp; IආൺඌൺKൺ (2018): 15 (record); HൺඒൺඌHං et al. (2018): 182 (distribution); Sൺඐൺൽൺ (2018): D-36 (listed); YൺඌඎඇൺGൺ et al. (2018): 33, 58 (photo); TൺKൺං &amp; SൺKඈඎ (2019): 114 (listed, distribution); TൺඇൺKൺ (2019): 64, 111 (listed); Uඋൺඒൺආൺ et al. (2019): 79, 81–82 (record, photo, habitat, phenology); MංඒൺඓൺKං et al. (2020): 65–66 (synthetic diet); OKඎൽൺ (2020): 44 (listed); SHංඓඎඈKൺ Pඋൾൿൾർඍඎඋൾ (2020): 128 (listed).
</paragraph>
</subSubSection>
<subSubSection id="C3656532A128FFE76ADDF8B7FC47818D" pageId="23" pageNumber="398" type="materials_examined">
<paragraph id="8BC036B9A128FFE76ADDF8B7FD658514" blockId="23.[151,761,220,2075]" box="[151,760,1836,1862]" pageId="23" pageNumber="398">
<emphasis id="B90BEAABA128FFE76ADDF8B7FE748514" bold="true" box="[151,489,1842,1861]" pageId="23" pageNumber="398">
Material examined. 
<collectingCountry id="F3687629A128FFE76B02F8B7FE128514" box="[328,399,1842,1861]" name="Japan" pageId="23" pageNumber="398">JAPAN</collectingCountry>
: HඈඇඌHඎ:
</emphasis>
<collectingRegion id="49BBF85BA128FFE76BA6F8B7FDC38514" box="[492,606,1842,1861]" country="Japan" name="Aomori" pageId="23" pageNumber="398">Aomori Pref.</collectingRegion>
:2JJ 
<specimenCount id="9D79FD30A128FFE768DEF8B7FD2D8514" box="[660,688,1836,1861]" pageId="23" pageNumber="398" type="female">1♀</specimenCount>
, Mutsu-
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF8C8FD658531" blockId="23.[151,761,220,2075]" box="[151,760,1863,1888]" pageId="23" pageNumber="398">
-shi, Sekine, 
<date id="FFC11079A128FFE76B4EF8C8FEF08531" box="[260,365,1869,1888]" pageId="23" pageNumber="398" value="1986-08-26">26.viii.1986</date>
, B. Tanaka. 
<collectingRegion id="49BBF85BA128FFE76B99F8C8FDF98531" box="[467,612,1869,1888]" country="Japan" name="Fukushima" pageId="23" pageNumber="398">Fukushima Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76820F8C8FD1A8531" box="[618,647,1863,1888]" pageId="23" pageNumber="398" type="female">1♀</specimenCount>
, Inawashiro-
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF8EDFD6A852A" blockId="23.[151,761,220,2075]" box="[151,759,1889,1915]" pageId="23" pageNumber="398">
-cho, Mt. Bandai-san, 
<date id="FFC11079A128FFE76B18F8EDFE33852A" box="[338,430,1896,1915]" pageId="23" pageNumber="398" value="1999-04-31">31.iv.1999</date>
, T. Shimada. 
<collectingRegion id="49BBF85BA128FFE76854F8EDFD11852A" box="[542,652,1896,1915]" country="Japan" name="Ibaraki" pageId="23" pageNumber="398">Ibaraki Pref.</collectingRegion>
: 2JJ 
<specimenCount id="9D79FD30A128FFE7688EF8EDFD6C852B" box="[708,753,1889,1915]" pageId="23" pageNumber="398" type="female">4 ♀♀</specimenCount>
,
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF807FD6485C4" blockId="23.[151,761,220,2075]" box="[151,761,1916,1941]" pageId="23" pageNumber="398">
Tsukuba-shi, 
<date id="FFC11079A128FFE76B41F807FEC385C4" box="[267,350,1922,1941]" pageId="23" pageNumber="398" value="2000-01-12">12.i.2000</date>
, K. Takahashi; 
<specimenCount id="9D79FD30A128FFE76BACF807FD9585C4" box="[486,520,1916,1941]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Tsukuba-shi, 
<date id="FFC11079A128FFE768CDF807FD4485C4" box="[647,729,1922,1941]" pageId="23" pageNumber="398" value="2000-04-01">1.iv.2000</date>
, K.
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF818FD6585E1" blockId="23.[151,761,220,2075]" box="[151,760,1943,1969]" pageId="23" pageNumber="398">
Takahashi. 
<collectingRegion id="49BBF85BA128FFE76AB2F818FEF185E1" box="[248,364,1949,1968]" country="Japan" name="Tochigi" pageId="23" pageNumber="398">Tochigi Pref.</collectingRegion>
: 3 JJ 
<specimenCount id="9D79FD30A128FFE76BEDF818FE5A85E1" box="[423,455,1943,1968]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Nikko-shi, Shichiri, 
<date id="FFC11079A128FFE768CAF818FD4085E1" box="[640,733,1949,1968]" pageId="23" pageNumber="398" value="2020-06-21">21.vi.2020</date>
, S.
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF83DFD6585B4" blockId="23.[151,761,220,2075]" pageId="23" pageNumber="398">
Maehara; 1 J, Utsunomiya-shi, Yanagita-ryokuchi, 
<date id="FFC11079A128FFE76804F83DFD30859A" box="[590,685,1976,1995]" pageId="23" pageNumber="398" value="2018-03-23">23.iii.2018</date>
, S. Maehara; 9 JJ 
<specimenCount id="9D79FD30A128FFE76AB7F857FEB085B4" box="[253,301,1996,2021]" pageId="23" pageNumber="398" type="female">9 ♀♀</specimenCount>
, Tochigi-shi, Kashiwagura, 
<date id="FFC11079A128FFE76855F857FDE085B4" box="[543,637,2002,2021]" pageId="23" pageNumber="398" value="1991-03-18">18.iii.1991</date>
, H. Yoshitomi
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF868FD648A51" blockId="23.[151,761,220,2075]" box="[151,761,2023,2048]" pageId="23" pageNumber="398">
(
<collectionCode id="ED6EAE7CA128FFE76AD7F868FF4D8A51" box="[157,208,2029,2048]" pageId="23" pageNumber="398">EUM</collectionCode>
); 
<specimenCount id="9D79FD30A128FFE76A94F868FE948A51" box="[222,265,2023,2048]" pageId="23" pageNumber="398" type="female">5♀♀</specimenCount>
, Tochigi-shi, Oominagawa, 
<date id="FFC11079A128FFE76BBFF868FDA18A51" box="[501,572,2029,2048]" pageId="23" pageNumber="398" value="1994-01-06">6.i.1994</date>
, H.Yoshitomi (
<collectionCode id="ED6EAE7CA128FFE768F1F868FD728A51" box="[699,751,2029,2048]" pageId="23" pageNumber="398">EUM</collectionCode>
);
</paragraph>
<paragraph id="8BC036B9A128FFE76ADDF78DFD6B8A4A" blockId="23.[151,761,220,2075]" box="[151,758,2049,2075]" pageId="23" pageNumber="398">
1 J 
<specimenCount id="9D79FD30A128FFE76AF3F78DFF488A4B" box="[185,213,2049,2075]" pageId="23" pageNumber="398" type="female">1♀</specimenCount>
, Tochigi-shi, Minagawajonai, 
<date id="FFC11079A128FFE76B9EF78DFD878A4A" box="[468,538,2056,2075]" pageId="23" pageNumber="398" value="1994-01-06">6.i.1994</date>
, H.Yoshitomi (
<collectionCode id="ED6EAE7CA128FFE768D3F78DFD518A4A" box="[665,716,2056,2075]" pageId="23" pageNumber="398">EUM</collectionCode>
); 1J
</paragraph>
<paragraph id="8BC036B9A128FFE76976FF41FC47818D" blockId="23.[826,1438,189,2084]" pageId="23" pageNumber="398">
<specimenCount id="9D79FD30A128FFE76976FF41FCC68287" box="[828,859,189,215]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Tochigi-shi, Shiriuch, 
<date id="FFC11079A128FFE76E6AFF41FBF58286" box="[1056,1128,196,215]" pageId="23" pageNumber="398" value="1994-01-04">4.i.1994</date>
, H. Yoshitomi (
<collectionCode id="ED6EAE7CA128FFE76EA0FF41FA838286" box="[1258,1310,196,215]" pageId="23" pageNumber="398">EUM</collectionCode>
); 6JJ 
<specimenCount id="9D79FD30A128FFE76F14FF41FA088287" box="[1374,1429,189,215]" pageId="23" pageNumber="398" type="female">10 ♀♀</specimenCount>
, Tochigi-shi, Tochigi-koukou, 
<date id="FFC11079A128FFE76E72FF5BFB0D82A0" box="[1080,1168,222,241]" pageId="23" pageNumber="398" value="1991-02-28">28.ii.1991</date>
, H.Yoshitomi (
<collectionCode id="ED6EAE7CA128FFE76F59FF5BFADA82A0" box="[1299,1351,222,241]" pageId="23" pageNumber="398">EUM</collectionCode>
); 1 J 
<specimenCount id="9D79FD30A128FFE76F32FF5BFA0B82A0" box="[1400,1430,216,241]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Tochigi-shi,Yanagibashi-cho, 
<date id="FFC11079A128FFE76E73FF7CFB16835D" box="[1081,1163,249,268]" pageId="23" pageNumber="398" value="1994-03-09">9.iii.1994</date>
, S. Nagashima;1J, Tochigi-shi, Fujioka-machi, Fujioka, 
<date id="FFC11079A128FFE76E4DFE91FBFE8376" box="[1031,1123,276,295]" pageId="23" pageNumber="398" value="2020-11-26">26.xi.2020</date>
, S. Maehara;2JJ (one in 
<figureCitation id="13442A3CA128FFE76F0AFE91FA0B8376" box="[1344,1430,276,295]" captionStart="Fig" captionStartId="15.[151,182,2010,2029]" captionTargetBox="[211,1313,201,1965]" captionTargetId="figure-13@15.[179,1408,189,1996]" captionTargetPageId="15" captionText="Fig. 7. Habitus of Anthocoris spp., dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A–B – A. chibi Hiura, 1959, male; C–D – A. confusus Reuter, 1884, male; E–F – A. japonicus Poppius, 1909, male; G–H – A. kalopanacis Kerzhner, 1977, female; I–J – A. nemoralis (Fabricius, 1794), male; K–L – A. takahashii Hiura, 1959, male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820983" httpUri="https://zenodo.org/record/5820983/files/figure.png" pageId="23" pageNumber="398">Figs 7E–F</figureCitation>
) 
<specimenCount id="9D79FD30A128FFE76976FEABFCF18310" box="[828,876,296,321]" pageId="23" pageNumber="398" type="female">3 ♀♀</specimenCount>
, Ôyama-shi, Shimada, 
<date id="FFC11079A128FFE76E70FEABFB1C8310" box="[1082,1153,302,321]" pageId="23" pageNumber="398" value="1994-01-06">6.i.1994</date>
, S. Nagashima. 
<collectingRegion id="49BBF85BA128FFE76F5AFEABFA148310" box="[1296,1417,302,321]" country="Japan" name="Saitama" pageId="23" pageNumber="398">Saitama Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76FD8FEABFCC6830A" pageId="23" pageNumber="398" type="female">2 ♀♀</specimenCount>
, Toda-shi, Doman, 
<date id="FFC11079A128FFE76E4AFECCFBF9830D" box="[1024,1124,329,348]" pageId="23" pageNumber="398" value="1998-12-27">27.xii.1998</date>
, S. Arai. 
<collectingRegion id="49BBF85BA128FFE76EE5FECCFA89830D" box="[1199,1300,329,348]" country="Japan" name="Chiba" pageId="23" pageNumber="398">Chiba Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76F53FECCFAAA830A" box="[1305,1335,322,348]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Narita-shi, Ôtake, 
<date id="FFC11079A128FFE7693DFEE1FC4F8326" box="[887,978,356,375]" pageId="23" pageNumber="398" value="2015-04-29">29.iv.2015</date>
, R. Nakamura.Tokyo-to:6JJ (one in 
<figureCitation id="13442A3CA128FFE76F5CFEE1FAEA8326" box="[1302,1399,356,375]" captionStart="Fig" captionStartId="19.[151,182,2012,2031]" captionTargetBox="[151,1433,563,1999]" captionTargetId="figure-13@19.[151,1436,562,2003]" captionTargetPageId="19" captionText="Fig. 10. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. chibi Hiura, 1959; C–D – A. confusus Reuter, 1884 (from India); E–F – A. japonicus Poppius, 1909.A, C, E – pygophore with paramere, dorsal (A, C) and laterodorsal (E) views; B, D, F – paramere, dorsal (B, D) and laterodorsal (F) views. Abbreviations: end ‒ endosoma; prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820995" httpUri="https://zenodo.org/record/5820995/files/figure.png" pageId="23" pageNumber="398">Figs 10E–F</figureCitation>
, one in 
<figureCitation id="13442A3CA128FFE7691BFEFBFC0283C0" box="[849,927,382,401]" captionStart="Fig" captionStartId="22.[151,182,1996,2015]" captionTargetBox="[236,1400,657,1963]" captionTargetId="figure-13@22.[151,1436,648,1972]" captionTargetPageId="22" captionText="Fig. 12. Abdominal sterna II–III of Anthocoris spp., male, ventral view (setae omitted). A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius, 1794); E – A. takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype. Abbreviations: ma ‒ membranous area; st2–3 ‒ abdominal sternum II to III. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5820999" httpUri="https://zenodo.org/record/5820999/files/figure.png" pageId="23" pageNumber="398">Figs 12C</figureCitation>
, 
<figureCitation id="13442A3CA128FFE769EDFEFBFC4383C0" box="[935,990,382,401]" captionStart="Fig" captionStartId="25.[151,182,2008,2027]" captionTargetBox="[203,1430,205,1962]" captionTargetId="figure-13@25.[151,1436,186,1980]" captionTargetPageId="25" captionText="Fig. 13. Male genitalia of Anthocoris spp. A–C – A. chibi Hiura, 1959; D–F – A. confusus Reuter, 1884; G–I – A. japonicus Poppius, 1909; J–L – A. nemoralis (Fabricius, 1794); M–O – A. takahashii Hiura, 1959. A, D, G, J, M – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F, H–I, K–L, N–O – paramere, two different views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821001" httpUri="https://zenodo.org/record/5821001/files/figure.png" pageId="23" pageNumber="398">13G–I</figureCitation>
) 
<specimenCount id="9D79FD30A128FFE769A3FEFBFB8283C0" box="[1001,1055,376,401]" pageId="23" pageNumber="398" type="female">11♀♀</specimenCount>
(one in 
<figureCitation id="13442A3CA128FFE76E2EFEFBFB2D83C0" box="[1124,1200,382,401]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="23" pageNumber="398">Fig. 15C</figureCitation>
), Setagaya-ku, Kinuta Park, 
<date id="FFC11079A128FFE76976FE1CFC3983FD" box="[828,932,409,428]" pageId="23" pageNumber="398" value="2001-08-10">10.viii.2001</date>
, J. Ikuta; 1 J, Chitose-funabashi, Karasuyama-ryokudo, no date, J. Narukawa. 
<collectingRegion id="49BBF85BA128FFE76993FE31FBFE8396" box="[985,1123,436,455]" country="Japan" name="Kanagawa" pageId="23" pageNumber="398">Kanagawa Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76E22FE31FB0E8397" box="[1128,1171,429,455]" pageId="23" pageNumber="398" type="female">3♀♀</specimenCount>
, Atugi-shi, Funako, 
<date id="FFC11079A128FFE76F74FE31FA048396" box="[1342,1433,436,455]" pageId="23" pageNumber="398" value="2001-04-13">13.iv.2001</date>
, S. Nagashima; 
<specimenCount id="9D79FD30A128FFE7698AFE4BFC7F83B0" box="[960,994,456,481]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Yokosuka-shi, Tsukui, Mt. Miurafuji, 
<date id="FFC11079A128FFE76F7CFE4BFA1D83B0" box="[1334,1408,462,481]" pageId="23" pageNumber="398" value="1988-05-05">5.v.1988</date>
, S. Miyakawa. 
<collectingRegion id="49BBF85BA128FFE769D5FE6CFB9C83AD" box="[927,1025,489,508]" country="Japan" name="Aichi" pageId="23" pageNumber="398">Aichi Pref.</collectingRegion>
: 1 J 
<specimenCount id="9D79FD30A128FFE76E67FE6CFBC183AA" box="[1069,1116,482,508]" pageId="23" pageNumber="398" type="female">4 ♀♀</specimenCount>
, Kasugai-shi, no date, S. Nagashima. 
<collectingRegion id="49BBF85BA128FFE76976FD81FC3E8046" box="[828,931,516,535]" country="Japan" name="Osaka" pageId="23" pageNumber="398">Osaka Pref.</collectingRegion>
: 3 JJ 
<specimenCount id="9D79FD30A128FFE76991FD81FB9A8047" box="[987,1031,509,535]" pageId="23" pageNumber="398" type="female">6♀♀</specimenCount>
, Ibaraki-shi, Niwakubo, 
<date id="FFC11079A128FFE76E91FD81FAA38046" box="[1243,1342,516,535]" pageId="23" pageNumber="398" value="2004-07-14">14.vii.2004</date>
, T. Ueda; 
<specimenCount id="9D79FD30A128FFE76FD8FD81FCD78060" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Sakai-shi, Izumigaoka, 
<date id="FFC11079A128FFE76E56FD9BFBE78060" box="[1052,1146,542,561]" pageId="23" pageNumber="398" value="2000-11-25">25.xi.2000</date>
, M.Yoshio. 
<collectingRegion id="49BBF85BA128FFE76EA8FD9BFAA38060" box="[1250,1342,542,561]" country="Japan" name="Nara" pageId="23" pageNumber="398">Nara Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76F09FD9BFAFC8060" box="[1347,1377,536,561]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Nara- -shi, Yagyû-kaidô, 
<date id="FFC11079A128FFE76990FDBCFBB8801D" box="[986,1061,569,588]" pageId="23" pageNumber="398" value="2001-05-04">4.v.2001</date>
, T.Tsuru; 2JJ 
<specimenCount id="9D79FD30A128FFE76EE6FDBCFB45801A" box="[1196,1240,562,588]" pageId="23" pageNumber="398" type="female">3♀♀</specimenCount>
, Nara-shi, Nakamachi, 
<date id="FFC11079A128FFE76976FDD1FC138036" box="[828,910,596,615]" pageId="23" pageNumber="398" value="2000-06-06">6.vi.2000</date>
, K. Yamada; 1 J, Gojô-shi, Yoshino-gawa Riv., 
<date id="FFC11079A128FFE76F6DFDD1FA198036" box="[1319,1412,596,615]" pageId="23" pageNumber="398" value="2000-06-29">29.vi.2000</date>
, T. Ueda; 
<specimenCount id="9D79FD30A128FFE76938FDEBFC1380D0" box="[882,910,616,641]" pageId="23" pageNumber="398" type="female">1♀</specimenCount>
, Gojô-shi, Minamiada-cho, 
<date id="FFC11079A128FFE76E33FDEBFB4980D0" box="[1145,1236,622,641]" pageId="23" pageNumber="398" value="2000-11-28">28.xi.2000</date>
, K.Yamada. 
<collectingRegion id="49BBF85BA128FFE76F75FDEBFCF480CD" country="Japan" name="Wakayama" pageId="23" pageNumber="398">Wakayama Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76924FD0CFC1680CA" box="[878,907,642,668]" pageId="23" pageNumber="398" type="female">1♀</specimenCount>
, Wakayama-shi, Myo-oji, 
<date id="FFC11079A128FFE76E27FD0CFB5980CD" box="[1133,1220,649,668]" pageId="23" pageNumber="398" value="2001-07-03">3.vii.2001</date>
, Y. Miyamoto. 
<emphasis id="B90BEAABA128FFE76F0EFD0CFA0180CD" bold="true" box="[1348,1436,649,668]" pageId="23" pageNumber="398">SHංKඈKඎ:</emphasis>
<collectingRegion id="49BBF85BA128FFE76976FD21FC5380E6" box="[828,974,676,695]" country="Japan" name="Tokushima" pageId="23" pageNumber="398">Tokushima Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE7699CFD21FC6B80E7" box="[982,1014,669,695]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Tokushima-shi, Hachiman-cho, Mukôterayama, Bunka-no-Mori Park, 
<geoCoordinate id="EE4B507EA128FFE769BCFD3BFBC28080" box="[1014,1119,702,721]" degrees="34.040278" direction="north" orientation="latitude" pageId="23" pageNumber="398" precision="1" value="34.04028">34.040278N</geoCoordinate>
<geoCoordinate id="EE4B507EA128FFE76E29FD3BFB458080" box="[1123,1240,702,721]" degrees="134.527222" direction="east" orientation="longitude" pageId="23" pageNumber="398" precision="1" value="134.52722">134.527222E</geoCoordinate>
, 
<date id="FFC11079A128FFE76E95FD3BFAAF8080" box="[1247,1330,702,721]" pageId="23" pageNumber="398" value="2008-03-01">1.iii.2008</date>
, K.Yamada; 11 JJ 
<specimenCount id="9D79FD30A128FFE76933FD5CFC2F80BA" box="[889,946,722,748]" pageId="23" pageNumber="398" type="female">16 ♀♀</specimenCount>
(one in 
<figureCitation id="13442A3CA128FFE769B2FD5CFBA780BD" box="[1016,1082,729,748]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="23" pageNumber="398">Fig. 9C</figureCitation>
), same locality, 
<date id="FFC11079A128FFE76E89FD5CFABD80BD" box="[1219,1312,729,748]" pageId="23" pageNumber="398" value="2011-03-18">18.iii.2011</date>
, K. Yamada; 2 JJ 
<specimenCount id="9D79FD30A128FFE76915FD71FC178157" box="[863,906,749,775]" pageId="23" pageNumber="398" type="female">2♀♀</specimenCount>
, Tokushima-shi, Jôroku-cho,Jôroku-Higashi Park, 
<date id="FFC11079A128FFE76F7DFD71FA048156" box="[1335,1433,756,775]" pageId="23" pageNumber="398" value="2013-12-30">30.xii.2013</date>
, T. Nakanishi. 
<collectingRegion id="49BBF85BA128FFE769FEFC8BFB868170" box="[948,1051,782,801]" country="Japan" name="Kochi" pageId="23" pageNumber="398">Kochi Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76E69FC8BFBDE8170" box="[1059,1091,776,801]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Tôyô-cho, None, 
<date id="FFC11079A128FFE76EAFFC8BFADC8170" box="[1253,1345,782,801]" pageId="23" pageNumber="398" value="2000-04-22">22.iv.2000</date>
, M. Takai. 
<emphasis id="B90BEAABA128FFE76976FCACFC10816D" bold="true" box="[828,909,809,828]" pageId="23" pageNumber="398">KඒඎඌHඎ:</emphasis>
<collectingRegion id="49BBF85BA128FFE769DFFCACFBB7816D" box="[917,1066,809,828]" country="Japan" name="Kumamoto" pageId="23" pageNumber="398">Kumamoto Pref.</collectingRegion>
: 
<specimenCount id="9D79FD30A128FFE76E79FCACFBC8816A" box="[1075,1109,802,828]" pageId="23" pageNumber="398" type="female">1 ♀</specimenCount>
, Chôyô-mura, Tochinoki, 
<date id="FFC11079A128FFE76F77FCACFA04816D" box="[1341,1433,809,828]" pageId="23" pageNumber="398" value="2002-04-27">27.iv.2002</date>
, T. Ueda. 
<collectingRegion id="49BBF85BA128FFE769C6FCC1FB8D8106" box="[908,1040,836,855]" country="Japan" name="Nagasaki" pageId="23" pageNumber="398">Nagasaki Pref.</collectingRegion>
: 2 JJ 
<specimenCount id="9D79FD30A128FFE76E05FCC1FBE28107" box="[1103,1151,829,855]" pageId="23" pageNumber="398" type="female">2 ♀♀</specimenCount>
, Nagasaki-shi, Kawaguchi-Park, 
<geoCoordinate id="EE4B507EA128FFE76976FCDBFC028120" box="[828,927,862,881]" degrees="32.76684" direction="north" orientation="latitude" pageId="23" pageNumber="398" precision="1" value="32.76684">32.76684N</geoCoordinate>
<geoCoordinate id="EE4B507EA128FFE769E2FCDBFB858120" box="[936,1048,862,881]" degrees="129.86353" direction="east" orientation="longitude" pageId="23" pageNumber="398" precision="1" value="129.86353">129.86353E</geoCoordinate>
, under flakes of bark of 
<taxonomicName id="4C7F4D3AA128FFE76F4CFCDBFA0B8120" box="[1286,1430,862,881]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="398" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA128FFE76F4CFCDBFA0B8120" box="[1286,1430,862,881]" italics="true" pageId="23" pageNumber="398">Zelkova serrata</emphasis>
</taxonomicName>
, 
<date id="FFC11079A128FFE76976FCFCFC0981DD" box="[828,916,889,908]" pageId="23" pageNumber="398" value="2020-02-21">21.ii.2020</date>
, T. Yasunaga et al. (
<collectionCode id="ED6EAE7CA128FFE76E76FCFCFB1D81DD" box="[1084,1152,889,908]" pageId="23" pageNumber="398">NWHS</collectionCode>
); 3 JJ 
<specimenCount id="9D79FD30A128FFE76E88FCFCFB6D81DA" box="[1218,1264,882,908]" pageId="23" pageNumber="398" type="female">3 ♀♀</specimenCount>
, Omura-shi, Omura High School garden, 
<date id="FFC11079A128FFE769A4FC11FBDB81F6" box="[1006,1094,916,935]" pageId="23" pageNumber="398" value="2021-02-24">24.ii.2021</date>
, under flakes of bark of 
<taxonomicName id="4C7F4D3AA128FFE76F5BFC11FA0A81F6" box="[1297,1431,916,935]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="398" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA128FFE76F5BFC11FA0A81F6" box="[1297,1431,916,935]" italics="true" pageId="23" pageNumber="398">Zelkova serrata</emphasis>
</taxonomicName>
, 
<geoCoordinate id="EE4B507EA128FFE76976FC2BFC0D8190" box="[828,912,942,961]" degrees="32.8970" direction="north" orientation="latitude" pageId="23" pageNumber="398" precision="5" value="32.897">32.8970N</geoCoordinate>
<geoCoordinate id="EE4B507EA128FFE769DDFC2BFC6A8190" box="[919,1015,942,961]" degrees="129.9631" direction="east" orientation="longitude" pageId="23" pageNumber="398" precision="5" value="129.9631">129.9631E</geoCoordinate>
, T. Yasunaga (
<collectionCode id="ED6EAE7CA128FFE76E3DFC2BFB268190" box="[1143,1211,942,961]" pageId="23" pageNumber="398">NWHS</collectionCode>
). All in TKPM except for EUM and NWHS.
</paragraph>
</subSubSection>
<subSubSection id="C3656532A128FFE76976FC68FBF287D5" pageId="23" pageNumber="398" type="diagnosis">
<paragraph id="8BC036B9A128FFE76976FC68FBF287D5" blockId="23.[826,1438,189,2084]" pageId="23" pageNumber="398">
<emphasis id="B90BEAABA128FFE76976FC68FBAD8655" bold="true" box="[828,1072,1005,1028]" pageId="23" pageNumber="398">Differential diagnosis.</emphasis>
Recognized by the following combination of characters: body (
<figureCitation id="13442A3CA128FFE76E39FB88FB798675" box="[1139,1252,1037,1060]" captionStart="Fig" captionStartId="15.[151,182,2010,2029]" captionTargetBox="[211,1313,201,1965]" captionTargetId="figure-13@15.[179,1408,189,1996]" captionTargetPageId="15" captionText="Fig. 7. Habitus of Anthocoris spp., dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A–B – A. chibi Hiura, 1959, male; C–D – A. confusus Reuter, 1884, male; E–F – A. japonicus Poppius, 1909, male; G–H – A. kalopanacis Kerzhner, 1977, female; I–J – A. nemoralis (Fabricius, 1794), male; K–L – A. takahashii Hiura, 1959, male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820983" httpUri="https://zenodo.org/record/5820983/files/figure.png" pageId="23" pageNumber="398">Figs 7E–F</figureCitation>
) generally black, covered with pruinose setae on dorsal surface; hemelytra and legs (
<figureCitation id="13442A3CA128FFE769D6FBC8FB948635" box="[924,1033,1101,1124]" captionStart="Fig" captionStartId="15.[151,182,2010,2029]" captionTargetBox="[211,1313,201,1965]" captionTargetId="figure-13@15.[179,1408,189,1996]" captionTargetPageId="15" captionText="Fig. 7. Habitus of Anthocoris spp., dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A–B – A. chibi Hiura, 1959, male; C–D – A. confusus Reuter, 1884, male; E–F – A. japonicus Poppius, 1909, male; G–H – A. kalopanacis Kerzhner, 1977, female; I–J – A. nemoralis (Fabricius, 1794), male; K–L – A. takahashii Hiura, 1959, male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820983" httpUri="https://zenodo.org/record/5820983/files/figure.png" pageId="23" pageNumber="398">Figs 7E–F</figureCitation>
) sometimes tinged with reddish brown; membrane (
<figureCitation id="13442A3CA128FFE769F2FBE8FB9B86D5" box="[952,1030,1133,1156]" captionStart="Fig" captionStartId="15.[151,182,2010,2029]" captionTargetBox="[211,1313,201,1965]" captionTargetId="figure-13@15.[179,1408,189,1996]" captionTargetPageId="15" captionText="Fig. 7. Habitus of Anthocoris spp., dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A–B – A. chibi Hiura, 1959, male; C–D – A. confusus Reuter, 1884, male; E–F – A. japonicus Poppius, 1909, male; G–H – A. kalopanacis Kerzhner, 1977, female; I–J – A. nemoralis (Fabricius, 1794), male; K–L – A. takahashii Hiura, 1959, male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820983" httpUri="https://zenodo.org/record/5820983/files/figure.png" pageId="23" pageNumber="398">Fig. 7E</figureCitation>
) smoky dark brown with broad whitish band on basal part; ostiolar peritreme (
<figureCitation id="13442A3CA128FFE76EA8FB08FAAB86F5" box="[1250,1334,1165,1188]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="23" pageNumber="398">Fig. 9C</figureCitation>
) straight, curved anteriorly at apex; abdominal sternum II (
<figureCitation id="13442A3CA128FFE76F72FB28FA088695" box="[1336,1429,1197,1220]" captionStart="Fig" captionStartId="22.[151,182,1996,2015]" captionTargetBox="[236,1400,657,1963]" captionTargetId="figure-13@22.[151,1436,648,1972]" captionTargetPageId="22" captionText="Fig. 12. Abdominal sterna II–III of Anthocoris spp., male, ventral view (setae omitted). A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius, 1794); E – A. takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype. Abbreviations: ma ‒ membranous area; st2–3 ‒ abdominal sternum II to III. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5820999" httpUri="https://zenodo.org/record/5820999/files/figure.png" pageId="23" pageNumber="398">Fig. 12C</figureCitation>
) with a pair of triangle-shaped membranous areas along posterior margin. General appearance similar to that of 
<taxonomicName id="4C7F4D3AA128FFE76976FA88FC558775" authorityName="Hiura" authorityYear="1959" box="[828,968,1293,1316]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="398" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="B90BEAABA128FFE76976FA88FC558775" box="[828,968,1293,1316]" italics="true" pageId="23" pageNumber="398">A. takahashii</emphasis>
</taxonomicName>
, but readily distinguished from the latter by the rather smaller body, the coloration of the hemelytral membrane, the shape of the paramere, and the overall structure of female genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C3656532A128FFE86976FA08FEDC8764" lastPageId="24" lastPageNumber="399" pageId="23" pageNumber="398" type="description">
<paragraph id="8BC036B9A128FFE76976FA08FAD084F5" blockId="23.[826,1438,189,2084]" pageId="23" pageNumber="398">
<emphasis id="B90BEAABA128FFE76976FA08FB1B87F5" bold="true" box="[828,1158,1421,1444]" pageId="23" pageNumber="398">
Redescription. 
<emphasis id="B90BEAABA128FFE769ADFA08FB1B87F5" bold="true" box="[999,1158,1421,1444]" italics="true" pageId="23" pageNumber="398">Male genitalia</emphasis>
</emphasis>
(
<figureCitation id="13442A3CA128FFE76EDEFA08FA8C87F5" box="[1172,1297,1421,1444]" captionStart="Fig" captionStartId="19.[151,182,2012,2031]" captionTargetBox="[151,1433,563,1999]" captionTargetId="figure-13@19.[151,1436,562,2003]" captionTargetPageId="19" captionText="Fig. 10. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. chibi Hiura, 1959; C–D – A. confusus Reuter, 1884 (from India); E–F – A. japonicus Poppius, 1909.A, C, E – pygophore with paramere, dorsal (A, C) and laterodorsal (E) views; B, D, F – paramere, dorsal (B, D) and laterodorsal (F) views. Abbreviations: end ‒ endosoma; prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820995" httpUri="https://zenodo.org/record/5820995/files/figure.png" pageId="23" pageNumber="398">Figs 10E–F</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76F56FA08FAF887F5" box="[1308,1381,1421,1444]" captionStart="Fig" captionStartId="25.[151,182,2008,2027]" captionTargetBox="[203,1430,205,1962]" captionTargetId="figure-13@25.[151,1436,186,1980]" captionTargetPageId="25" captionText="Fig. 13. Male genitalia of Anthocoris spp. A–C – A. chibi Hiura, 1959; D–F – A. confusus Reuter, 1884; G–I – A. japonicus Poppius, 1909; J–L – A. nemoralis (Fabricius, 1794); M–O – A. takahashii Hiura, 1959. A, D, G, J, M – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F, H–I, K–L, N–O – paramere, two different views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821001" httpUri="https://zenodo.org/record/5821001/files/figure.png" pageId="23" pageNumber="398">13G–I</figureCitation>
): Pygophore (
<figureCitation id="13442A3CA128FFE769E9FA28FB988795" box="[931,1029,1453,1476]" captionStart="Fig" captionStartId="25.[151,182,2008,2027]" captionTargetBox="[203,1430,205,1962]" captionTargetId="figure-13@25.[151,1436,186,1980]" captionTargetPageId="25" captionText="Fig. 13. Male genitalia of Anthocoris spp. A–C – A. chibi Hiura, 1959; D–F – A. confusus Reuter, 1884; G–I – A. japonicus Poppius, 1909; J–L – A. nemoralis (Fabricius, 1794); M–O – A. takahashii Hiura, 1959. A, D, G, J, M – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F, H–I, K–L, N–O – paramere, two different views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821001" httpUri="https://zenodo.org/record/5821001/files/figure.png" pageId="23" pageNumber="398">Fig. 13G</figureCitation>
) turbinate, longer than wide, covered with 8–12 long, stout setae intermixed with short, suberect setae along outer margin and on posteroventral surface; mid-dorsal surface very hirsute with suberect setae; paramere (
<figureCitation id="13442A3CA128FFE769DDF9A8FC678415" box="[919,1018,1581,1604]" captionStart="Fig" captionStartId="19.[151,182,2012,2031]" captionTargetBox="[151,1433,563,1999]" captionTargetId="figure-13@19.[151,1436,562,2003]" captionTargetPageId="19" captionText="Fig. 10. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. chibi Hiura, 1959; C–D – A. confusus Reuter, 1884 (from India); E–F – A. japonicus Poppius, 1909.A, C, E – pygophore with paramere, dorsal (A, C) and laterodorsal (E) views; B, D, F – paramere, dorsal (B, D) and laterodorsal (F) views. Abbreviations: end ‒ endosoma; prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820995" httpUri="https://zenodo.org/record/5820995/files/figure.png" pageId="23" pageNumber="398">Figs 10F</figureCitation>
, 
<figureCitation id="13442A3CA128FFE76E4DF9A8FBD08415" box="[1031,1101,1581,1604]" captionStart="Fig" captionStartId="25.[151,182,2008,2027]" captionTargetBox="[203,1430,205,1962]" captionTargetId="figure-13@25.[151,1436,186,1980]" captionTargetPageId="25" captionText="Fig. 13. Male genitalia of Anthocoris spp. A–C – A. chibi Hiura, 1959; D–F – A. confusus Reuter, 1884; G–I – A. japonicus Poppius, 1909; J–L – A. nemoralis (Fabricius, 1794); M–O – A. takahashii Hiura, 1959. A, D, G, J, M – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F, H–I, K–L, N–O – paramere, two different views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821001" httpUri="https://zenodo.org/record/5821001/files/figure.png" pageId="23" pageNumber="398">13H–I</figureCitation>
) slender, very slightly curved, apically acute, and bent anteriad, with a few very short setae on median portion; longitudinal groove lacking, but a weak longitudinal edge present on anterior half.
</paragraph>
<paragraph id="8BC036B9A128FFE76910F928FC7685D5" blockId="23.[826,1438,189,2084]" pageId="23" pageNumber="398">
<emphasis id="B90BEAABA128FFE76910F928FB858495" bold="true" box="[858,1048,1709,1732]" italics="true" pageId="23" pageNumber="398">Female genitalia</emphasis>
(
<figureCitation id="13442A3CA128FFE76E60F928FB0C8495" box="[1066,1169,1709,1732]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="23" pageNumber="398">Fig. 15C</figureCitation>
): Copulatory tube fused on intersegmental membrane slightly left to the middle, between sterna VII and VIII, approximately 
<quantity id="4C879B5CA128FFE76F51F968FA018555" box="[1307,1436,1773,1796]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="23" pageNumber="398" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
in length, very thin and almost same width from base to apex (junction to trunk of conductive tissue), with few distinct twists; trunk of conductive tissue invisible (possibly dissolved).
</paragraph>
<paragraph id="8BC036B9A128FFE76910F808FA008A75" blockId="23.[826,1438,189,2084]" pageId="23" pageNumber="398">
<emphasis id="B90BEAABA128FFE76910F808FC6685F5" bold="true" box="[858,1019,1933,1956]" italics="true" pageId="23" pageNumber="398">Measurements</emphasis>
[mm; JJ (n = 10) / 
<specimenCount id="9D79FD30A128FFE76E84F803FB6985F4" box="[1230,1268,1926,1957]" pageId="23" pageNumber="398" type="female">♀♀</specimenCount>
(n = 10)]. Body length 3.45–4.08 / 3.43–4.25; head length (excl. neck) 0.45–0.53 / 0.45–0.55; head width across eyes 0.49–0.55 / 0.48–0.56; vertex width 0.27–0.30 / 0.26–0.32; length of antennal segments I – 0.15–0.19 / 0.15–0.20, II – 0.44–0.52
</paragraph>
<paragraph id="8BC036B9A127FFE86ADDFF3BFE8A8384" blockId="24.[151,760,190,2069]" pageId="24" pageNumber="399">/ 0.39–0.50, III – 0.26–0.32 / 0.26–0.32, and IV – 0.33–0.37 / 0.32–0.39; length of labial segments II – 0.13–0.17 / 0.14–0.19, III – 0.46–0.54 / 0.47–0.56, and IV – 0.27–0.34 / 0.29–0.35; anterior pronotal width 0.43–0.46 / 0.42–0.50; mesal pronotal length 0.49–0.63 / 0.52–0.66; basal pronotal width 1.15–1.27 / 1.05–1.35; length of embolial margin 1.08–1.26 / 1.00–1.35; length of cuneal margin 0.63–0.74 / 0.57–0.75; maximum width across hemelytra 1.20–1.38 / 1.06–1.40.</paragraph>
<paragraph id="8BC036B9A127FFE86ADDFE5BFEDC8764" blockId="24.[151,760,190,2069]" pageId="24" pageNumber="399">
<emphasis id="B90BEAABA127FFE86ADDFE5BFE8883A4" bold="true" box="[151,277,478,501]" pageId="24" pageNumber="399">Bionomics.</emphasis>
<taxonomicName id="4C7F4D3AA127FFE86B56FE5BFD9C83A4" authorityName="Poppius" authorityYear="1909" box="[284,513,478,501]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90BEAABA127FFE86B56FE5BFD9C83A4" box="[284,513,478,501]" italics="true" pageId="24" pageNumber="399">Anthocoris japonicus</emphasis>
</taxonomicName>
is frequently collected from 
<taxonomicName id="4C7F4D3AA127FFE86A85FE7BFEEF8044" box="[207,370,510,533]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA127FFE86A85FE7BFEEF8044" box="[207,370,510,533]" italics="true" pageId="24" pageNumber="399">Zelkova serrata</emphasis>
</taxonomicName>
(Mංඒൺආඈඍඈ 1965, YൺඌඎඇൺGൺ 2001b), where its prey consists of aphids, 
<taxonomicName id="4C7F4D3AA127FFE86BB7FD9BFEB38004" authority="(Baker, 1919)" baseAuthorityName="Baker" baseAuthorityYear="1919" class="Insecta" family="Aphididae" genus="Paracolopha" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="species" species="morrisoni">
<emphasis id="B90BEAABA127FFE86BB7FD9BFD6A8064" box="[509,759,542,565]" italics="true" pageId="24" pageNumber="399">Paracolopha morrisoni</emphasis>
(Baker, 1919)
</taxonomicName>
(
<taxonomicName id="4C7F4D3AA127FFE86B75FDBBFE2B8004" box="[319,438,574,597]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
: Sternorrhyncha: 
<taxonomicName id="4C7F4D3AA127FFE86833FDBBFD708004" box="[633,749,574,597]" class="Insecta" family="Aphididae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="family">Aphididae</taxonomicName>
), living on the leaves. They also prey on the Japanese pine blast scale, 
<taxonomicName id="4C7F4D3AA127FFE86B56FDFBFD6580C4" authority="(Kuwana, 1905)" baseAuthorityName="Kuwana" baseAuthorityYear="1905" box="[284,760,638,661]" class="Insecta" family="Margarodidae" genus="Matsucoccus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="species" species="matsumurae">
<emphasis id="B90BEAABA127FFE86B56FDFBFDA780C4" box="[284,570,638,661]" italics="true" pageId="24" pageNumber="399">Matsucoccus matsumurae</emphasis>
(Kuwana, 1905)
</taxonomicName>
(Coccomorpha: 
<taxonomicName id="4C7F4D3AA127FFE86B06FD1BFE6E80E4" authorityName="Morrison" authorityYear="1927" box="[332,499,670,693]" class="Insecta" family="Matsucoccidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="family">Matsucoccidae</taxonomicName>
) (Bඎ &amp; ZHൾඇG 2001). Almost nothing is known about this species’ movements once it departs 
<taxonomicName id="4C7F4D3AA127FFE86B73FD5BFE7C80A4" box="[313,481,734,757]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA127FFE86B73FD5BFE7C80A4" box="[313,481,734,757]" italics="true" pageId="24" pageNumber="399">Zelkova serrata</emphasis>
</taxonomicName>
generally before summer, but a few individuals have been collected from 
<taxonomicName id="4C7F4D3AA127FFE868CDFD7BFE688164" authority="Thunb. var. umbellata" class="Magnoliopsida" family="Elaeagnaceae" genus="Elaeagnus" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="variety" species="umbellata" variety="umbellata">
<emphasis id="B90BEAABA127FFE868CDFD7BFE9F8164" italics="true" pageId="24" pageNumber="399">Elaeagnus umbellata</emphasis>
Thunb. var. 
<emphasis id="B90BEAABA127FFE86BC0FC9BFE688164" box="[394,501,798,821]" italics="true" pageId="24" pageNumber="399">umbellata</emphasis>
</taxonomicName>
in June at Nikko, in the northern part of 
<collectingRegion id="49BBF85BA127FFE86B18FCBBFE358104" box="[338,424,830,853]" country="Japan" name="Tochigi" pageId="24" pageNumber="399">Tochigi</collectingRegion>
. Males and females hibernate under bark flakes of 
<taxonomicName id="4C7F4D3AA127FFE86B2DFCDBFE508124" box="[359,461,862,885]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA127FFE86B2DFCDBFE508124" box="[359,461,862,885]" italics="true" pageId="24" pageNumber="399">Z. serrata</emphasis>
</taxonomicName>
(
<figureCitation id="13442A3CA127FFE86B93FCDBFDAB8124" box="[473,566,862,885]" captionStart="Fig" captionStartId="33.[151,182,1974,1993]" captionTargetBox="[151,1436,189,1804]" captionTargetId="figure-13@33.[151,1437,186,1824]" captionTargetPageId="33" captionText="Fig. 17. Habitats, overwintering individuals, and mating pair of Anthocoris spp. A–B – Field survey on hibernating anthocorids under bark-flakes of Zelkova serrata at urbanized zone of Nagasaki City (Kawaguchi Park) in early February; C – A. japonicus Poppius, 1909, overwintering adults under the bark-flake of Z. serrata; D–E – same, mating pair; F – A. miyamotoi Hiura, 1959, adult males and final instar immature (right), overwintering individuals under the bark of Z. serrata; G – same, active male adult on flower of Camellia sasanqua in winter (observed on Jan. 4, 2020)." figureDoi="http://doi.org/10.5281/zenodo.5821013" httpUri="https://zenodo.org/record/5821013/files/figure.png" pageId="24" pageNumber="399">Fig. 17C</figureCitation>
), 
<taxonomicName id="4C7F4D3AA127FFE8680DFCDBFF4D81C4" authority="Jacq." authorityName="Jacq." class="Magnoliopsida" family="Ulmaceae" genus="Ulmus" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="parvifolia">
<emphasis id="B90BEAABA127FFE8680DFCDBFD658124" box="[583,760,862,885]" italics="true" pageId="24" pageNumber="399">Ulmus parvifolia</emphasis>
Jacq.
</taxonomicName>
, and 
<taxonomicName id="4C7F4D3AA127FFE86B47FCFBFD3C81C4" authority="(Thunb.) Planch." authorityName="Planch." baseAuthorityName="Thunb." box="[269,673,894,917]" class="Magnoliopsida" family="Cannabaceae" genus="Aphananthe" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="B90BEAABA127FFE86B47FCFBFE7C81C4" box="[269,481,894,917]" italics="true" pageId="24" pageNumber="399">Aphananthe aspera</emphasis>
(Thunb.) Planch.
</taxonomicName>
(Hංඎඋൺ 1959, Eඇඃඎ et al. 2013, JඎඇG et al. 2013, Uඋൺඒൺආൺ et al. 2019). The adults appear to be active in winter when temperatures are sufficiently warm, and even mating pairs have been observed during this season (
<figureCitation id="13442A3CA127FFE86873FC7BFD238644" box="[569,702,1022,1045]" captionStart="Fig" captionStartId="33.[151,182,1974,1993]" captionTargetBox="[151,1436,189,1804]" captionTargetId="figure-13@33.[151,1437,186,1824]" captionTargetPageId="33" captionText="Fig. 17. Habitats, overwintering individuals, and mating pair of Anthocoris spp. A–B – Field survey on hibernating anthocorids under bark-flakes of Zelkova serrata at urbanized zone of Nagasaki City (Kawaguchi Park) in early February; C – A. japonicus Poppius, 1909, overwintering adults under the bark-flake of Z. serrata; D–E – same, mating pair; F – A. miyamotoi Hiura, 1959, adult males and final instar immature (right), overwintering individuals under the bark of Z. serrata; G – same, active male adult on flower of Camellia sasanqua in winter (observed on Jan. 4, 2020)." figureDoi="http://doi.org/10.5281/zenodo.5821013" httpUri="https://zenodo.org/record/5821013/files/figure.png" pageId="24" pageNumber="399">Figs 17D–E</figureCitation>
). The searching male encounters a female and attempts to mount her. After mounting the female’s dorsum, the male curves his abdomen from the female’s right side and inserts his paramere to median part of ventral side between abdominal segment VII and VIII. The female does not struggle once the male has mounted her. Duration of copulation was generally at least 10 minutes, as seen in the other 
<taxonomicName id="4C7F4D3AA127FFE86ADDFB7BFEFF8744" box="[151,354,1278,1301]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90BEAABA127FFE86ADDFB7BFE918744" box="[151,268,1278,1301]" italics="true" pageId="24" pageNumber="399">Anthocoris</emphasis>
species
</taxonomicName>
(Hඈඋඍඈඇ et al. 2000, 2002; Hඈඋඍඈඇ &amp; Lൾඐංඌ 2005).
</paragraph>
</subSubSection>
<subSubSection id="C3656532A127FFE86ADDFABBFBCC8004" pageId="24" pageNumber="399" type="distribution">
<paragraph id="8BC036B9A127FFE86ADDFABBFBCC8004" blockId="24.[151,760,190,2069]" lastBlockId="24.[828,1437,190,597]" pageId="24" pageNumber="399">
<emphasis id="B90BEAABA127FFE86ADDFABBFEB88704" bold="true" box="[151,293,1342,1365]" pageId="24" pageNumber="399">Distribution.</emphasis>
<collectingCountry id="F3687629A127FFE86B60FABBFEF78704" box="[298,362,1342,1365]" name="Japan" pageId="24" pageNumber="399">Japan</collectingCountry>
: Honshu: 
<collectingRegion id="49BBF85BA127FFE86B84FABBFDBD8704" box="[462,544,1342,1365]" country="Japan" name="Aomori" pageId="24" pageNumber="399">Aomori</collectingRegion>
(IർHංඍൺ 1988), 
<collectingRegion id="49BBF85BA127FFE868F7FABBFD6A8704" box="[701,759,1342,1365]" country="Japan" name="Iwate" pageId="24" pageNumber="399">Iwate</collectingRegion>
(Iඐൺඍൾ Pඋൾൿൾർඍඎඋൾ 2001), 
<collectingRegion id="49BBF85BA127FFE86BFFFADBFDAC8724" box="[437,561,1374,1397]" country="Japan" name="Fukushima" pageId="24" pageNumber="399">Fukushima</collectingRegion>
*, 
<collectingRegion id="49BBF85BA127FFE86809FADBFD098724" box="[579,660,1374,1397]" country="Japan" name="Ibaraki" pageId="24" pageNumber="399">Ibaraki</collectingRegion>
*, 
<collectingRegion id="49BBF85BA127FFE868EDFADBFD658724" box="[679,760,1374,1397]" country="Japan" name="Tochigi" pageId="24" pageNumber="399">Tochigi</collectingRegion>
(Sඈඇඈൻൾ 1997, MൺൾHൺඋൺ 2015), 
<collectingRegion id="49BBF85BA127FFE86BBAFAFBFDD887C4" box="[496,581,1406,1429]" country="Japan" name="Saitama" pageId="24" pageNumber="399">Saitama</collectingRegion>
(Nඈඓൺඐൺ 1978), 
<collectingRegion id="49BBF85BA127FFE86ADDFA1BFF4487E4" box="[151,217,1438,1461]" country="Japan" name="Chiba" pageId="24" pageNumber="399">Chiba</collectingRegion>
(Mൺඋඎඒൺආൺ et al. 2000, SඎඓඎKං 2003), 
<collectingRegion id="49BBF85BA127FFE868F8FA1BFD6A87E4" box="[690,759,1438,1461]" country="Japan" name="Tokyo" pageId="24" pageNumber="399">Tokyo</collectingRegion>
(Hංඎඋൺ 1959), 
<collectingRegion id="49BBF85BA127FFE86B76FA3BFE308784" box="[316,429,1470,1493]" country="Japan" name="Kanagawa" pageId="24" pageNumber="399">Kanagawa</collectingRegion>
(Pඈඉඉංඎඌ 1909, SඎඓඎKං 1981, HൺඒൺඌHං et al. 2018), 
<collectingRegion id="49BBF85BA127FFE86B30FA5BFE2587A4" box="[378,440,1502,1525]" country="Japan" name="Fukui" pageId="24" pageNumber="399">Fukui</collectingRegion>
(Sൺඌൺඃං 1985), 
<collectingRegion id="49BBF85BA127FFE86817FA5BFD1387A4" box="[605,654,1502,1525]" country="Japan" name="Gifu" pageId="24" pageNumber="399">Gifu</collectingRegion>
(TൺKൺං &amp; SൺKඈඎ 2019), 
<collectingRegion id="49BBF85BA127FFE86B73FA7BFE018444" box="[313,412,1534,1557]" country="Japan" name="Shizuoka" pageId="24" pageNumber="399">Shizuoka</collectingRegion>
(SHංඓඎඈKൺ Pඋൾൿൾർඍඎඋൾ 2020), 
<collectingRegion id="49BBF85BA127FFE86ADDF99BFF4F8464" box="[151,210,1566,1589]" country="Japan" name="Aichi" pageId="24" pageNumber="399">Aichi</collectingRegion>
(AඌൺඈKൺ &amp; IൾKං 1990), 
<collectingRegion id="49BBF85BA127FFE86B9FF99BFD858464" box="[469,536,1566,1589]" country="Japan" name="Osaka" pageId="24" pageNumber="399">Osaka</collectingRegion>
(YൺආൺඓൺKං &amp; TൺKൺ- Kඎඋൺ 2011), 
<collectingRegion id="49BBF85BA127FFE86B6BF9BBFEF78404" box="[289,362,1598,1621]" country="Japan" name="Hyogo" pageId="24" pageNumber="399">Hyogo</collectingRegion>
(UඌHංඋඈKංඍൺ et al. 2012), 
<collectingRegion id="49BBF85BA127FFE868C9F9BBFD2A8404" box="[643,695,1598,1621]" country="Japan" name="Nara" pageId="24" pageNumber="399">Nara</collectingRegion>
(Hංඎ- උൺ 1959), 
<collectingRegion id="49BBF85BA127FFE86B43F9DBFE188424" box="[265,389,1630,1653]" country="Japan" name="Wakayama" pageId="24" pageNumber="399">Wakayama</collectingRegion>
*, 
<collectingRegion id="49BBF85BA127FFE86BD6F9DBFE798424" box="[412,484,1630,1653]" country="Japan" name="Tottori" pageId="24" pageNumber="399">Tottori</collectingRegion>
(Hංඎඋൺ 1959), 
<collectingRegion id="49BBF85BA127FFE868DBF9DBFD6A8424" box="[657,759,1630,1653]" country="Japan" name="Okayama" pageId="24" pageNumber="399">Okayama</collectingRegion>
(OKൺඒൺආൺ Pඋൾൿൾർඍඎඋൾ 2020), 
<collectingRegion id="49BBF85BA127FFE86BA9F9FBFDC984C4" box="[483,596,1662,1685]" country="Japan" name="Hiroshima" pageId="24" pageNumber="399">Hiroshima</collectingRegion>
(NൺKൺආඎඋൺ et al. 1997), 
<collectingRegion id="49BBF85BA127FFE86AB6F91BFEEF84E4" box="[252,370,1694,1717]" country="Japan" name="Yamaguchi" pageId="24" pageNumber="399">Yamaguchi</collectingRegion>
(Hංඎඋൺ 1959); Shikoku: 
<collectingRegion id="49BBF85BA127FFE8683BF91BFD7684E4" box="[625,747,1694,1717]" country="Japan" name="Tokushima" pageId="24" pageNumber="399">Tokushima</collectingRegion>
*, 
<collectingRegion id="49BBF85BA127FFE86ADDF93BFF408484" box="[151,221,1726,1749]" country="Japan" name="Ehime" pageId="24" pageNumber="399">Ehime</collectingRegion>
(Yൺඇඈ &amp; Yൺආൺආඈඍඈ 2000), 
<collectingRegion id="49BBF85BA127FFE8685FF93BFDC18484" box="[533,604,1726,1749]" country="Japan" name="Kochi" pageId="24" pageNumber="399">Kochi</collectingRegion>
*; Kyushu: 
<collectingRegion id="49BBF85BA127FFE86899F93BFF4584A4" country="Japan" name="Fukuoka" pageId="24" pageNumber="399">Fukuoka</collectingRegion>
(TൺKൺKඎඋൺ 1959), 
<collectingRegion id="49BBF85BA127FFE86BEFF95BFD9584A4" box="[421,520,1758,1781]" country="Japan" name="Nagasaki" pageId="24" pageNumber="399">Nagasaki</collectingRegion>
(Uඋൺඒൺආൺ et al. 2019), 
<collectingRegion id="49BBF85BA127FFE86ADDF97BFE938544" box="[151,270,1790,1813]" country="Japan" name="Kumamoto" pageId="24" pageNumber="399">Kumamoto</collectingRegion>
(OඍඌඎKൺ &amp; YඈඌHංඓൺKං 1987), 
<collectingRegion id="49BBF85BA127FFE86806F97BFDE48544" box="[588,633,1790,1813]" country="Japan" name="Oita" pageId="24" pageNumber="399">Oita</collectingRegion>
(TൺKൺKඎඋൺ 1984); Tsushima Island (YൺඌඎඇൺGൺ 1999). Chishima (Kuril) Islands: Kunashir Is. (KൾඋඓHඇൾඋ 1988, KൺඇඒඎKඈඏൺ &amp; MൺඋඎඌංK 2006). 
<collectingCountry id="F3687629A127FFE86B09F8DBFE118524" box="[323,396,1886,1909]" name="Russia" pageId="24" pageNumber="399">Russia</collectingCountry>
: Far East: 
<collectingRegion id="49BBF85BA127FFE86BBEF8DBFD048524" box="[500,665,1886,1909]" country="Russia" name="Primorskiy" pageId="24" pageNumber="399">Primorsky Kray</collectingRegion>
(KൾඋඓH- ඇൾඋ 1988, VංඇඈKඎඋඈඏ et al. 2010). 
<collectingCountry id="F3687629A127FFE8685DF8FBFDC185C4" box="[535,604,1918,1941]" name="South Korea" pageId="24" pageNumber="399">Korea</collectingCountry>
: Gangwon-do, 
<collectingRegion id="49BBF85BA127FFE86ADDF81BFEBE85E4" box="[151,291,1950,1973]" country="South Korea" name="Gyeonggi" pageId="24" pageNumber="399">Gyeonggi-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE86B61F81BFD9585E4" box="[299,520,1950,1973]" country="South Korea" name="Chungbuk" pageId="24" pageNumber="399">Chungcheongbuk-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE8685BF81BFD6985E4" box="[529,756,1950,1973]" country="South Korea" name="Chungnam" pageId="24" pageNumber="399">Chungcheongnam-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE86ADDF83BFEF08584" box="[151,365,1982,2005]" country="South Korea" name="Gyeongbuk" pageId="24" pageNumber="399">Gyeongsangbuk-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE86B31F83BFDCA8584" box="[379,599,1982,2005]" country="South Korea" name="Gyeongnam" pageId="24" pageNumber="399">Gyeongsangnam-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE8682FF83BFD6E8584" box="[613,755,1982,2005]" country="South Korea" name="Jeonbuk" pageId="24" pageNumber="399">Jeollabuk-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE86ADDF85BFEB385A4" box="[151,302,2014,2037]" country="South Korea" name="Jeonnam" pageId="24" pageNumber="399">Jeollanam-do</collectingRegion>
, 
<collectingRegion id="49BBF85BA127FFE86B77F85BFE1385A4" box="[317,398,2014,2037]" country="South Korea" name="Jeju" pageId="24" pageNumber="399">Jeju-do</collectingRegion>
(KൾඋඓHඇൾඋ 1988, Kඐඈඇ et al. 2001, JඎඇG &amp; Lൾൾ 2017). 
<collectingCountry id="F3687629A127FFE86B8FF87BFD978A44" box="[453,522,2046,2069]" name="China" pageId="24" pageNumber="399">China</collectingCountry>
: 
<collectingRegion id="49BBF85BA127FFE86852F87BFDE48A44" box="[536,633,2046,2069]" country="China" name="Zhejiang" pageId="24" pageNumber="399">Zhejiang</collectingRegion>
(ZHൾඇG &amp; Bඎ 1990, Bඎ &amp; ZHൾඇG 2001). This species is one of the more common species of Japanese 
<taxonomicName id="4C7F4D3AA127FFE86E8AFF5BFACC82A4" box="[1216,1361,222,245]" class="Insecta" family="Anthocoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="399" phylum="Arthropoda" rank="family">Anthocoridae</taxonomicName>
and is widespread in the lowland areas of Honshu, Shikoku, and Kyushu, 
<collectingCountry id="F3687629A127FFE86982FE9BFB988364" box="[968,1029,286,309]" name="Japan" pageId="24" pageNumber="399">Japan</collectingCountry>
(
<figureCitation id="13442A3CA127FFE86E59FE9BFBFF8364" box="[1043,1122,286,309]" captionStart="Fig" captionStartId="38.[153,184,2034,2053]" captionTargetBox="[168,1419,673,1984]" captionTargetId="figure-299@38.[153,1434,654,2003]" captionTargetPageId="38" captionText="Fig. 21. Distribution of Anthocoris takahashii Hiura, 1959 (circles), A. japonicus Poppius, 1909 (triangles), and A. kalopanacis Kerzhner, 1977 (squares) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.5821022" httpUri="https://zenodo.org/record/5821022/files/figure.png" pageId="24" pageNumber="399">Fig. 21</figureCitation>
). In addition, it extends to the Korean Peninsula and to a few localities in the Russian Far East and east 
<collectingCountry id="F3687629A127FFE86981FEDBFB938324" box="[971,1038,350,373]" name="China" pageId="24" pageNumber="399">China</collectingCountry>
. Distribution records of the species in 
<collectingCountry id="F3687629A127FFE86976FEFBFCE483C4" box="[828,889,382,405]" name="Japan" pageId="24" pageNumber="399">Japan</collectingCountry>
seem to be associated with the distribution pattern of 
<taxonomicName id="4C7F4D3AA127FFE86910FE1BFB9E83E4" box="[858,1027,414,437]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA127FFE86910FE1BFB9E83E4" box="[858,1027,414,437]" italics="true" pageId="24" pageNumber="399">Zelkova serrata</emphasis>
</taxonomicName>
, a well-known tree planted in various urbanized areas such as parks and roadsides. In urbanized zones in 
<collectingRegion id="49BBF85BA127FFE869D3FE5BFBEF83A4" box="[921,1138,478,501]" country="Japan" name="Nagasaki" pageId="24" pageNumber="399">Nagasaki Prefecture</collectingRegion>
, this anthocorid is evidently expanding its habitat, utilizing 
<taxonomicName id="4C7F4D3AA127FFE86ECFFE7BFAB18044" box="[1157,1324,510,533]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="399" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA127FFE86ECFFE7BFAB18044" box="[1157,1324,510,533]" italics="true" pageId="24" pageNumber="399">Zelkova serrata</emphasis>
</taxonomicName>
planted in parks and school campuses for landscaping (
<figureCitation id="13442A3CA127FFE86F5BFD9BFA098064" box="[1297,1428,542,565]" captionStart="Fig" captionStartId="33.[151,182,1974,1993]" captionTargetBox="[151,1436,189,1804]" captionTargetId="figure-13@33.[151,1437,186,1824]" captionTargetPageId="33" captionText="Fig. 17. Habitats, overwintering individuals, and mating pair of Anthocoris spp. A–B – Field survey on hibernating anthocorids under bark-flakes of Zelkova serrata at urbanized zone of Nagasaki City (Kawaguchi Park) in early February; C – A. japonicus Poppius, 1909, overwintering adults under the bark-flake of Z. serrata; D–E – same, mating pair; F – A. miyamotoi Hiura, 1959, adult males and final instar immature (right), overwintering individuals under the bark of Z. serrata; G – same, active male adult on flower of Camellia sasanqua in winter (observed on Jan. 4, 2020)." figureDoi="http://doi.org/10.5281/zenodo.5821013" httpUri="https://zenodo.org/record/5821013/files/figure.png" pageId="24" pageNumber="399">Figs 17A–B</figureCitation>
) (cf. Uඋൺඒൺආൺ et al. 2019).
</paragraph>
</subSubSection>
</treatment>
</document>