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<mods:title id="8F6FA362EC8F48A1F4327CCB887153BB">The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae)</mods:title>
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<mods:namePart id="E54DA72FE764D1FBD0540CA71C0F7822">Chiba, Hideyuki</mods:namePart>
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<mods:namePart id="F1228528229610CDCEB7E3961ED4C5D4">Tsukiyama, Hiroshi</mods:namePart>
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<mods:namePart id="C59B52BFBA24B4E5E49C2627E6569A60">Liang, Jia-Yuan</mods:namePart>
<mods:affiliation id="988A2E912888D91B78BB0EE63293A8C1">Department of Life Science, National Taiwan Normal University, Taipei, Taiwan 116, R. O. C. &amp; sesiidae 760402 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4320 - 3695</mods:affiliation>
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<mods:namePart id="63EDEFAF12150771414B617CF5F02CBF">Wang, Shou-Ming</mods:namePart>
<mods:affiliation id="251FD331739DA88DFB0FA93D4707F3CD">Conservation Section, Conservation and Recreation Division, Forestry Bureau, Council of Agriculture, Executive Yuan, Taipei, Taiwan 100, R. O. C. falcon @ forest. gov. tw; https: // orcid. org / 0000 - 0003 - 4157 - 9736</mods:affiliation>
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<mods:affiliation id="C898A1EECE302C0AB64A7F01EFA7A269">Department of Life Science, National Taiwan Normal University, Taipei, Taiwan 116, R. O. C. &amp; jeffshannnu 2014 mouse @ gmail. com; https: // orcid. org / 0000 - 0003 - 2566 - 7349</mods:affiliation>
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<paragraph id="8BC33261FFBDFF87C6D4FC0BFD3CA2F8" blockId="7.[151,641,945,1007]" box="[151,641,945,971]" pageId="7" pageNumber="287">
<heading id="D08B850DFFBDFF87C6D4FC0BFD3CA2F8" box="[151,641,945,971]" level="2" pageId="7" pageNumber="287">
<taxonomicName id="4C7C49E2FFBDFF87C6D4FC0BFD3CA2F8" ID-CoL="3JGF6" authority="Murayama &amp; Okamura, 1973" authorityName="Murayama &amp; Okamura" authorityYear="1973" box="[151,641,945,971]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="287" phylum="Arthropoda" rank="species" species="nuydai">
<emphasis id="B908EE73FFBDFF87C6D4FC0BFE92A2F8" box="[151,303,945,971]" italics="true" pageId="7" pageNumber="287">Halpe nuydai</emphasis>
<bibRefCitation id="EFED4F90FFBDFF87C775FC0BFD3CA2F8" author="Murayama, S. &amp; Okamura, H." box="[310,641,945,971]" pageId="7" pageNumber="287" pagination="10 - 25" refId="ref6180" refString="Murayama, S. &amp; Okamura, H. (1973) Butterflies of Luzon Island in Philippines, with descriptions of new species and subspecies. Tyo to Ga, 24 (1), 10 - 25." type="journal article" year="1973">Murayama &amp; Okamura, 1973</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC33261FFBDFF87C6D4FC6FFEC6A2DC" blockId="7.[151,641,945,1007]" box="[151,379,981,1007]" pageId="7" pageNumber="287">
<emphasis id="B908EE73FFBDFF87C6D4FC6FFEB8A2DC" box="[151,261,981,1007]" italics="true" pageId="7" pageNumber="287">Tyô to Ga</emphasis>
24(1): 24.
</paragraph>
</subSubSection>
<caption id="DF0362E9FFBDFF87C6D4F817FAEDA6DA" ID-DOI="http://doi.org/10.5281/zenodo.3990963" ID-Zenodo-Dep="3990963" httpUri="https://zenodo.org/record/3990963/files/figure.png" pageId="7" pageNumber="287" startId="7.[151,264,1965,1990]" targetBox="[157,1436,1029,1935]" targetPageId="7">
<paragraph id="8BC33261FFBDFF87C6D4F817FAEDA6DA" blockId="7.[151,1436,1965,2026]" pageId="7" pageNumber="287">
<emphasis id="B908EE73FFBDFF87C6D4F817FEEEA6F5" bold="true" box="[151,339,1965,1990]" pageId="7" pageNumber="287">FIGURES 3745.</emphasis>
3739.
<taxonomicName id="4C7C49E2FFBDFF87C7E0F814FC55A6F5" authority="Murayama, 1983" authorityName="Murayama" authorityYear="1983" box="[419,1000,1966,1990]" class="Insecta" family="Hesperiidae" genus="Carterocephalus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="287" phylum="Arthropoda" rank="subSpecies" species="dieckmanni" subSpecies="soyosanus">
<emphasis id="B908EE73FFBDFF87C7E0F814FC89A6F5" box="[419,820,1966,1990]" italics="true" pageId="7" pageNumber="287">Carterocephalus dieckmanni soyosanus</emphasis>
<bibRefCitation id="EFED4F90FFBDFF87C579F814FC55A6F5" author="Murayama, S." box="[826,1000,1966,1990]" pageId="7" pageNumber="287" pagination="281 - 288" refId="ref6043" refString="Murayama, S. (1983) Some new Rhopalocera from southwest and northwest China (Lepidoptera: Rhopalocera). Entomotaxonomia, 5 (4), 281 - 288." type="journal article" year="1983">Murayama, 1983</bibRefCitation>
</taxonomicName>
, holotype; 4042.
<taxonomicName id="4C7C49E2FFBDFF87C2E1F814FE09A6D9" authority="Murayama &amp; Kimura, 1990" authorityName="Murayama &amp; Kimura. Valid" authorityYear="1990" class="Insecta" family="Hesperiidae" genus="Sebastonyma" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="287" phylum="Arthropoda" rank="species" species="suthepiana">
<emphasis id="B908EE73FFBDFF87C2E1F814FA21A6F5" box="[1186,1436,1966,1990]" italics="true" pageId="7" pageNumber="287">Sebastonyma suthepiana</emphasis>
<bibRefCitation id="EFED4F90FFBDFF87C6D4F868FE09A6D9" author="Murayama, S &amp; Kimura, Y." box="[151,436,2001,2026]" pageId="7" pageNumber="287" pagination="19 - 24" refId="ref6146" refString="Murayama, S &amp; Kimura, Y. (1990) Some new butterflies from Thailand. Nature &amp; Insects, 25 (2), 19 - 24. [in Japanese]" type="journal article" year="1990">Murayama &amp; Kimura, 1990</bibRefCitation>
</taxonomicName>
, holotype; 4345.
<taxonomicName id="4C7C49E2FFBDFF87C432F868FB93A6D9" authority="Murayama &amp; Okamura, 1973" authorityName="Murayama &amp; Okamura" authorityYear="1973" box="[625,1070,2001,2026]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="287" phylum="Arthropoda" rank="species" species="nuydai">
<emphasis id="B908EE73FFBDFF87C432F868FD47A6D9" box="[625,762,2002,2026]" italics="true" pageId="7" pageNumber="287">Halpe nuydai</emphasis>
<bibRefCitation id="EFED4F90FFBDFF87C543F868FB93A6D9" author="Murayama, S. &amp; Okamura, H." box="[768,1070,2001,2026]" pageId="7" pageNumber="287" pagination="10 - 25" refId="ref6180" refString="Murayama, S. &amp; Okamura, H. (1973) Butterflies of Luzon Island in Philippines, with descriptions of new species and subspecies. Tyo to Ga, 24 (1), 10 - 25." type="journal article" year="1973">Murayama &amp; Okamura, 1973</bibRefCitation>
</taxonomicName>
, holotype. Scale bar = 1 cm.
</paragraph>
</caption>
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<paragraph id="8BC33261FFB2FF88C684FF23FB11A180" blockId="8.[151,1437,153,395]" box="[199,1196,153,179]" pageId="8" pageNumber="288">
<materialsCitation id="3B14383CFFB2FF88C684FF23FB11A180" ID-GBIF-Occurrence="2831948304" box="[199,1196,153,179]" collectingDate="1972-04-02" country="Virgin Islands" location="Labels" pageId="8" pageNumber="288" specimenCount="1" typeStatus="holotype">
<location id="8EA364BAFFB2FF88C684FF23FEABA180" LSID="urn:lsid:plazi:treatment:03D58377FFBDFF88C6D4FC0BFC71A0B8:8EA364BAFFB2FF88C684FF23FEABA180" box="[199,278,153,179]" country="Virgin Islands" name="Labels" pageId="8" pageNumber="288">Labels</location>
: “
<collectingCountry id="F36B72F1FFB2FF88C768FF23FE15A180" box="[299,424,153,179]" name="Virgin Islands" pageId="8" pageNumber="288">St Thomas</collectingCountry>
/Luzon/
<date id="FFC214A1FFB2FF88C441FF23FDC9A180" box="[514,628,153,179]" pageId="8" pageNumber="288" value="1972-04-02">
<collectingDate id="EF86ED49FFB2FF88C441FF23FDC9A180" box="[514,628,153,179]" pageId="8" pageNumber="288" value="1972-04-02">2-IV-1972</collectingDate>
</date>
/S. MURAYAMA,” “
<taxonomicName id="4C7C49E2FFB2FF88C51EFF23FC1CA180" authorityName="Moore" authorityYear="1878" box="[861,929,153,179]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="genus">Halpe</taxonomicName>
/nuydai/
<typeStatus id="54C78CC3FFB2FF88C245FF23FB1DA180" box="[1030,1184,153,179]" pageId="8" pageNumber="288" type="holotype">HOLOTYPE</typeStatus>
.”
</materialsCitation>
</paragraph>
<paragraph id="8BC33261FFB2FF88C684FF07FC54A1C8" blockId="8.[151,1437,153,395]" pageId="8" pageNumber="288">
A male specimen was designated as the
<typeStatus id="54C78CC3FFB2FF88C4C8FF07FD50A1E4" box="[651,749,189,215]" pageId="8" pageNumber="288" type="holotype">holotype</typeStatus>
by
<bibRefCitation id="EFED4F90FFB2FF88C55BFF07FBCEA1E4" author="Murayama, S. &amp; Okamura, H." box="[792,1139,189,215]" pageId="8" pageNumber="288" pagination="10 - 25" refId="ref6180" refString="Murayama, S. &amp; Okamura, H. (1973) Butterflies of Luzon Island in Philippines, with descriptions of new species and subspecies. Tyo to Ga, 24 (1), 10 - 25." type="journal article" year="1973">Murayama &amp; Okamura (1973)</bibRefCitation>
, and the specimen was retrieved in
<collectionCode id="ED6DAAA4FFB2FF88C74AFF5BFEFBA1C8" box="[265,326,225,251]" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="8" pageNumber="288">LBM</collectionCode>
(
<figureCitation id="13472EE4FFB2FF88C716FF5BFE60A1C8" box="[341,477,224,251]" captionStart="FIGURES 3745" captionStartId="7.[151,264,1965,1990]" captionTargetBox="[157,1436,1029,1935]" captionTargetId="figure@7.[151,1436,1029,1940]" captionTargetPageId="7" captionText="FIGURES 3745. 3739. Carterocephalus dieckmanni soyosanus Murayama, 1983, holotype; 4042. Sebastonyma suthepiana Murayama &amp; Kimura, 1990, holotype; 4345. Halpe nuydai Murayama &amp; Okamura, 1973, holotype. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3990963" httpUri="https://zenodo.org/record/3990963/files/figure.png" pageId="8" pageNumber="288">Figs. 4345</figureCitation>
; Dried Insect Database Reg. no. 1500022031).
</paragraph>
<paragraph id="8BC33261FFB2FF88C684FEBFFC71A0B8" blockId="8.[151,1437,153,395]" pageId="8" pageNumber="288">
Though
<bibRefCitation id="EFED4F90FFB2FF88C765FEBFFD3DA02C" author="Murayama, S. &amp; Okamura, H." box="[294,640,261,287]" pageId="8" pageNumber="288" pagination="10 - 25" refId="ref6180" refString="Murayama, S. &amp; Okamura, H. (1973) Butterflies of Luzon Island in Philippines, with descriptions of new species and subspecies. Tyo to Ga, 24 (1), 10 - 25." type="journal article" year="1973">Murayama &amp; Okamura (1973)</bibRefCitation>
claimed that the genitalia of
<taxonomicName id="4C7C49E2FFB2FF88C58FFEBFFBA5A02C" authorityName="Murayama &amp; Okamura" authorityYear="1973" box="[972,1048,261,287]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="species" species="nuydai">
<emphasis id="B908EE73FFB2FF88C58FFEBFFBA5A02C" box="[972,1048,261,287]" italics="true" pageId="8" pageNumber="288">nuydai</emphasis>
</taxonomicName>
resembled that of
<taxonomicName id="4C7C49E2FFB2FF88C2AFFEBFFE10A070" authority="(Matsumura, 1910)" baseAuthorityName="Matsumura" baseAuthorityYear="1910" class="Insecta" family="Hesperiidae" genus="Thoressa" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="species" species="horishana">
<emphasis id="B908EE73FFB2FF88C2AFFEBFFF72A070" italics="true" pageId="8" pageNumber="288">Thoressa horishana</emphasis>
(Matsumura, 1910)
</taxonomicName>
from
<collectingCountry id="F36B72F1FFB2FF88C7ACFE93FDF9A070" box="[495,580,297,323]" name="Taiwan" pageId="8" pageNumber="288">Taiwan</collectingCountry>
, they somehow classified
<taxonomicName id="4C7C49E2FFB2FF88C525FE93FC0FA070" authorityName="Murayama &amp; Okamura" authorityYear="1973" box="[870,946,297,323]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="species" species="nuydai">
<emphasis id="B908EE73FFB2FF88C525FE93FC0FA070" box="[870,946,297,323]" italics="true" pageId="8" pageNumber="288">nuydai</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="4C7C49E2FFB2FF88C206FE93FA9EA070" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[1093,1315,297,323]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B908EE73FFB2FF88C206FE93FB34A070" box="[1093,1161,297,323]" italics="true" pageId="8" pageNumber="288">Halpe</emphasis>
Moore, 1878
</taxonomicName>
. The taxon was synonymized by
<bibRefCitation id="EFED4F90FFB2FF88C7CAFEF7FDF8A054" author="Kawazoe, A." box="[393,581,333,359]" pageId="8" pageNumber="288" pagination="91 - 98" refId="ref5719" refString="Kawazoe, A. (1973) A revisional note on some Philippine Lycaenidae and Hesperiidae reported by Murayama and Okamura. Tyo to Ga, 24 (4), 91 - 98." type="journal article" year="1973">Kawazoé (1973)</bibRefCitation>
, and we follow his treatment.
<taxonomicName id="4C7C49E2FFB2FF88C5DBFEF7FA3FA054" authority="Murayama &amp; Okamura, 1973" authorityName="Murayama &amp; Okamura" authorityYear="1973" box="[920,1410,333,359]" class="Insecta" family="Hesperiidae" genus="Halpe" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="species" species="nuydai">
<emphasis id="B908EE73FFB2FF88C5DBFEF7FB8DA054" box="[920,1072,333,359]" italics="true" pageId="8" pageNumber="288">Halpe nuydai</emphasis>
<bibRefCitation id="EFED4F90FFB2FF88C274FEF7FA3FA054" author="Murayama, S. &amp; Okamura, H." box="[1079,1410,333,359]" pageId="8" pageNumber="288" pagination="10 - 25" refId="ref6180" refString="Murayama, S. &amp; Okamura, H. (1973) Butterflies of Luzon Island in Philippines, with descriptions of new species and subspecies. Tyo to Ga, 24 (1), 10 - 25." type="journal article" year="1973">Murayama &amp; Okamura, 1973</bibRefCitation>
</taxonomicName>
is a junior subjective synonym of
<taxonomicName id="4C7C49E2FFB2FF88C7BAFECBFC7AA0B8" authority="Inoue &amp; Kawazoe, 1969" authorityName="Inoue &amp; Kawazoe" authorityYear="1969" box="[505,967,369,395]" class="Insecta" family="Hesperiidae" genus="Thoressa" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="288" phylum="Arthropoda" rank="species" species="justini">
<emphasis id="B908EE73FFB2FF88C7BAFECBFD11A0B8" box="[505,684,369,395]" italics="true" pageId="8" pageNumber="288">Thoressa justini</emphasis>
<bibRefCitation id="EFED4F90FFB2FF88C4F0FECBFC7AA0B8" author="Inoue, S. &amp; Kawazoe, A." box="[691,967,369,395]" pageId="8" pageNumber="288" pagination="380 - 383" refId="ref5638" refString="Inoue, S. &amp; Kawazoe, A. (1969) A new Hesperiid butterfly found in Luzon belonging to the hitherto unrecorded genus from the Philippines: Thoressa Swinhoe (Lepidoptera). Kontyu, 37, 380 - 383." type="journal article" year="1969">Inoué &amp; Kawazoé, 1969</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>