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<mods:title id="FB0376594766C9FB69EF1DA844814A78">A review of the damselfly genus Megalestes Selys, 1862 (Insecta: Odonata Zygoptera: Synlestidae) using integrative taxonomic methods</mods:title>
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<taxonomicName id="4C7D4D56FF90FFA16EE2FD8EFDFCFB27" ID-CoL="9X3J8" authority="Karube, 2014" authorityName="Karube" authorityYear="2014" box="[151,574,547,574]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B909EAC7FF90FFA16EE2FD8EFE4EFB24" bold="true" box="[151,396,547,574]" italics="true" pageId="13" pageNumber="258">Megalestes australis</emphasis>
<bibRefCitation id="EFEC4B24FF90FFA16FE1FD8EFDFCFB27" author="Karube, H." box="[404,574,547,573]" pageId="13" pageNumber="258" pagination="73 - 76" refId="ref14198" refString="Karube, H. (2014) Vietnamese Odonata collected in 1992 - 2003 surveys. IV. Synlestidae. Tombo, 56, 73 - 76." type="journal article" year="2014">Karube, 2014</bibRefCitation>
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<paragraph id="8BC236D5FF90FFA16EE2FDEAFE58FB78" blockId="13.[151,574,547,610]" box="[151,410,583,610]" pageId="13" pageNumber="258">
(
<figureCitation id="13462A50FF90FFA16EEAFDEAFF3EFB7B" box="[159,252,583,609]" captionStart="FIGURE 3" captionStartId="14.[151,250,1420,1444]" captionTargetBox="[160,1427,195,1385]" captionTargetId="figure-34@14.[151,1436,181,1396]" captionTargetPageId="14" captionText="FIGURE 3. Caudal appendages of males (dorsal view). a) australis; b) irma, paratype; c) distans, from Yaan, Sichuan; d) distans from Dashahe, Guizhou; e) heros; f) omeiensis, paratype; g) kurahashii; h) major; i) haui; j) “palaceus”; k) micans, bd: basal disk; l) “chengi”; m) riccii; n) “maai”; o) “tuska,” from Damingshan, Guangxi; p) “tuska,” from Jinxiu, Guangxi. (Scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407489" httpUri="https://zenodo.org/record/4407489/files/figure.png" pageId="13" pageNumber="258">Figs. 3a</figureCitation>
;
<figureCitation id="13462A50FF90FFA16F7DFDEAFEE4FB7B" box="[264,294,583,609]" captionStart="FIGURE 4" captionStartId="15.[151,250,1443,1467]" captionTargetBox="[175,1424,244,1412]" captionTargetId="figure-56@15.[151,1437,237,1419]" captionTargetPageId="15" captionText="FIGURE 4. Caudal appendages of males (lateral view). a) australis; b) irma, paratype; c) distans, from Yaan, Sichuan; d) distans from Qingchengshan, Chengdu, Sichuan; e) heros; f) omeiensis, paratype; g) kurahashii; h) major; i) haui; j) “palaceus”; k) micans; l) “Chengi,” paratype; m) riccii; n) maai; o) tuska, from Damingshan, Guangxi; p) tuska, from Jinxiu, Guangxi. (Scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407493" httpUri="https://zenodo.org/record/4407493/files/figure.png" pageId="13" pageNumber="258">4a</figureCitation>
;
<figureCitation id="13462A50FF90FFA16F44FDEAFEA9FB78" box="[305,363,583,610]" captionStart="FIGURE 5" captionStartId="18.[151,250,1869,1893]" captionTargetBox="[214,1369,199,1838]" captionTargetId="figure-34@18.[205,1381,181,1845]" captionTargetPageId="18" captionText="FIGURE 5. Genital ligula of males (lateral view) (am) and ventral view (nab). a) australis; b) distans; c) heros; d) omeiensis, paratype; e) major; f) irma, paratype; g) “palaceus”; h) haui; i) “Chengi”; j) maai; k) riccii; l) tuska; m) micans; n) australis, Hr: horn; o) distans, Tlb: terminal lateral lobe; p) heros; q) omeiensis, paratype; r) major; s) irma, paratype; t) kurahashii; u) “palaceus”; v) haui; w) micans, from Jinping, Yunnan; x) “Chengi”; y) maai; z) riccii, Blb: basal lateral lobe; aa) tuska, Mlb: middle lateral lobe; ab) micans, from Fengyangshan, Zhejiang. (Scale bar = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407495" httpUri="https://zenodo.org/record/4407495/files/figure.png" pageId="13" pageNumber="258">5a, n</figureCitation>
;
<figureCitation id="13462A50FF90FFA16F03FDEAFE50FB78" box="[374,402,583,610]" captionStart="FIGURE 6" captionStartId="19.[151,250,1319,1343]" captionTargetBox="[168,1412,187,1286]" captionTargetId="figure-17@19.[163,1423,181,1295]" captionTargetPageId="19" captionText="FIGURE 6. Line drawing of male paraprocts (lateral view). a) australis; b) distans; c) gyalsey; d) kurahashii; e) major; f) haui; g) heros; h) omeiensis; i) micans; j) micans; k) riccii; l) irma." figureDoi="http://doi.org/10.5281/zenodo.4407497" httpUri="https://zenodo.org/record/4407497/files/figure.png" pageId="13" pageNumber="258">6a</figureCitation>
)
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<paragraph id="8BC236D5FF90FFA16EE2FD21FB14FBD8" blockId="13.[151,1437,651,707]" pageId="13" pageNumber="258">
<treatmentCitationGroup id="AB6D11FBFF90FFA16EE2FD21FB14FBD8" pageId="13" pageNumber="258">
<treatmentCitation id="0ADC10C4FF90FFA16EE2FD21FD88FBB9" author="Karube, H." box="[151,586,651,676]" page="76" pageId="13" pageNumber="258" year="2014">
<taxonomicName id="4C7D4D56FF90FFA16EE2FD21FD88FBB9" ID-CoL="9X3J8" authority="Karube, 2014: 73 - 76" authorityName="Karube" authorityPageNumber="73 - 76" authorityYear="2014" box="[151,586,651,676]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B909EAC7FF90FFA16EE2FD21FEABFBBE" box="[151,361,652,676]" italics="true" pageId="13" pageNumber="258">Megalestes australis</emphasis>
<bibRefCitation id="EFEC4B24FF90FFA16F05FD21FD88FBB9" author="Karube, H." box="[368,586,651,676]" pageId="13" pageNumber="258" pagination="73 - 76" refId="ref14198" refString="Karube, H. (2014) Vietnamese Odonata collected in 1992 - 2003 surveys. IV. Synlestidae. Tombo, 56, 73 - 76." type="journal article" year="2014">Karube, 2014: 7376</bibRefCitation>
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, fig. 3 [Bach Ma National Park, Thua Thien Hue,
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];
<treatmentCitation id="0ADC10C4FF90FFA16ACCFD21FA55FBB9" author="Phan, Q. T. &amp; To, V. Q." box="[1209,1431,651,676]" page="1" pageId="13" pageNumber="258" year="2018">
<bibRefCitation id="EFEC4B24FF90FFA16ACCFD21FA55FBB9" author="Phan, Q. T. &amp; To, V. Q." box="[1209,1431,651,676]" pageId="13" pageNumber="258" pagination="1 - 6" refId="ref14558" refString="Phan, Q. T. &amp; To, V. Q. (2018) The genus Megalestes Selys, 1862 in Vietnam, with first description of female of Megalestes australis Karube, 2014 (Odonata: Zygoptera: Synlestidae). Journal of the International Dragonfly Fund, 111, 1 - 6." type="journal article" year="2018">Phan &amp; To 2018: 16</bibRefCitation>
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, figs. 79, 1418, 2527 [Tay Giang,
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; Bach Ma National Park, Thua Thien Hue,
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].
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</paragraph>
<paragraph id="8BC236D5FF90FFA16EE2FD5DFD23FA49" blockId="13.[151,1436,752,851]" pageId="13" pageNumber="258">
This is a relatively large
<collectingCountry id="F36A7645FF90FFA16FC4FD5DFDD2FA10" box="[433,528,752,778]" name="Vietnam" pageId="13" pageNumber="258">Vietnam</collectingCountry>
endemic species, somewhat resembling
<taxonomicName id="4C7D4D56FF90FFA16DA9FD5CFBFDFA10" ID-CoL="6R7PM" authorityName="Needham" authorityYear="1930" box="[988,1087,752,778]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="heros">
<emphasis id="B909EAC7FF90FFA16DA9FD5CFBFDFA10" box="[988,1087,752,778]" italics="true" pageId="13" pageNumber="258">M. heros</emphasis>
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and
<taxonomicName id="4C7D4D56FF90FFA16A0DFD5CFB2FFA10" ID-CoL="737PN" authorityName="Needham" authorityYear="1930" box="[1144,1261,752,778]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="distans">
<emphasis id="B909EAC7FF90FFA16A0DFD5CFB2FFA10" box="[1144,1261,752,778]" italics="true" pageId="13" pageNumber="258">M. distans</emphasis>
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in the shape of appendages but unique in the shape of the genital ligula with relatively long and thin horns (
<figureCitation id="13462A50FF90FFA16AD0FCB9FAD1FA35" box="[1189,1299,788,815]" captionStart="FIGURE 4" captionStartId="15.[151,250,1443,1467]" captionTargetBox="[175,1424,244,1412]" captionTargetId="figure-56@15.[151,1437,237,1419]" captionTargetPageId="15" captionText="FIGURE 4. Caudal appendages of males (lateral view). a) australis; b) irma, paratype; c) distans, from Yaan, Sichuan; d) distans from Qingchengshan, Chengdu, Sichuan; e) heros; f) omeiensis, paratype; g) kurahashii; h) major; i) haui; j) “palaceus”; k) micans; l) “Chengi,” paratype; m) riccii; n) maai; o) tuska, from Damingshan, Guangxi; p) tuska, from Jinxiu, Guangxi. (Scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407493" httpUri="https://zenodo.org/record/4407493/files/figure.png" pageId="13" pageNumber="258">Fig. 4a, n</figureCitation>
). Its species status was supported by molecular analyses (
<figureCitation id="13462A50FF90FFA16CFBFC95FD10FA49" box="[654,722,824,851]" captionStart="FIGURE 7" captionStartId="20.[151,250,1978,2002]" captionTargetBox="[213,1367,496,1949]" captionTargetId="figure-214@20.[211,1375,489,1954]" captionTargetPageId="20" captionText="FIGURE 7. Phylogenetic reconstruction from ITS data modified based on BI and ML trees, with both posterior probabilities (right) and bootstrap values (left) shown." figureDoi="http://doi.org/10.5281/zenodo.4407499" httpUri="https://zenodo.org/record/4407499/files/figure.png" pageId="13" pageNumber="258">Fig. 7</figureCitation>
).
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