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<document id="962D07A6929471529FFC8065847E8D37" ID-CLB-Dataset="27596" ID-DOI="10.11646/zootaxa.4551.3.5" ID-GBIF-Dataset="2a0a0531-766d-4054-8638-1134f95fe1bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2622967" ID-ZooBank="74227122-3FDB-4375-99A0-E1E8310365C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270284525" checkinUser="plazi" docAuthor="Marceniuk, Alexandre Pires, Marchena, Jose, Oliveira, Claudio &amp; Betancur-R, Ricardo" docDate="2019" docId="03D48796EC77FFADF8909951FBF3F83E" docLanguage="en" docName="zootaxa.4551.3.5.pdf" docOrigin="Zootaxa 4551 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chinchaysuyoa labiata" docType="treatment" docVersion="12" lastPageNumber="367" masterDocId="FFEDFFEEEC73FFABF8079835FFAFFFE5" masterDocTitle="Chinchaysuyoa, a new genus of the fish family Ariidae (Siluriformes), with a redescription of Chinchaysuyoa labiata from Ecuador and a new species description from Peru" masterLastPageNumber="378" masterPageNumber="361" pageNumber="365" updateTime="1698716110118" updateUser="ExternalLinkService">
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<mods:title id="025E0ED1EC489939F029AD9E0AD1D574">Chinchaysuyoa, a new genus of the fish family Ariidae (Siluriformes), with a redescription of Chinchaysuyoa labiata from Ecuador and a new species description from Peru</mods:title>
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<mods:namePart id="3D7D660368CDD8B74F22680CD03F8402">Marceniuk, Alexandre Pires</mods:namePart>
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<mods:namePart id="6B2DD6A6033B80BCF19E2177C6176E56">Marchena, Jose</mods:namePart>
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<mods:namePart id="261ED5F1557DCC936E036568C3F946E4">Oliveira, Claudio</mods:namePart>
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<mods:namePart id="E6BF6289BB4D18B67C738FFF3F8AE1C6">Betancur-R, Ricardo</mods:namePart>
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<mods:date id="AFD09C29597B9DAADAA3F0AD51038DC8">2019</mods:date>
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<treatment id="03D48796EC77FFADF8909951FBF3F83E" ID-DOI="http://doi.org/10.5281/zenodo.3510516" ID-GBIF-Taxon="154533748" ID-Zenodo-Dep="3510516" LSID="urn:lsid:plazi:treatment:03D48796EC77FFADF8909951FBF3F83E" httpUri="http://treatment.plazi.org/id/03D48796EC77FFADF8909951FBF3F83E" lastPageId="6" lastPageNumber="367" pageId="4" pageNumber="365">
<subSubSection id="C367650BEC77FFAFF8909951FD26FE9A" box="[151,649,356,383]" pageId="4" pageNumber="365" type="nomenclature">
<paragraph id="8BC23680EC77FFAFF8909951FD26FE9A" blockId="4.[151,649,356,416]" box="[151,649,356,383]" pageId="4" pageNumber="365">
<heading id="D08A81ECEC77FFAFF8909951FD26FE9A" bold="true" box="[151,649,356,383]" fontSize="11" level="1" pageId="4" pageNumber="365" reason="1">
<taxonomicName id="4C7D4D03EC77FFAFF8909951FD26FE9A" authority="(Boulenger 1898)" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[151,649,356,383]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B909EA92EC77FFAFF8909951FD26FE9A" bold="true" box="[151,649,356,383]" pageId="4" pageNumber="365">
<emphasis id="B909EA92EC77FFAFF8909951FE03FE9B" bold="true" box="[151,428,356,382]" italics="true" pageId="4" pageNumber="365">Chinchaysuyoa labiata</emphasis>
(
<bibRefCitation id="EFEC4B71EC77FFAFF9BC9950FD2EFE9A" author="Boulenger, G. A." box="[443,641,357,383]" pageId="4" pageNumber="365" pagination="1 - 3" refId="ref7112" refString="Boulenger, G. A. (1898) Viaggio del Dr. Enrico Festa nell' Ecuador e regioni vicine. Poissons de l'Equateur. [Part I]. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 13, 1 - 3." type="journal article" year="1898">Boulenger 1898</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC77FFAFF89099B2FE07FE45" box="[151,424,391,416]" pageId="4" pageNumber="365" type="description">
<paragraph id="8BC23680EC77FFAFF89099B2FE07FE45" blockId="4.[151,649,356,416]" box="[151,424,391,416]" pageId="4" pageNumber="365">
<figureCitation id="13462A05EC77FFAFF89099B2FF58FE45" box="[151,247,391,416]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="4" pageNumber="365">Figure 4</figureCitation>
,
<figureCitation id="13462A05EC77FFAFF8FB99B2FEA5FE45" box="[252,266,391,416]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="4" pageNumber="365">5</figureCitation>
,
<figureCitation id="13462A05EC77FFAFF90899B2FEB1FE45" box="[271,286,391,416]" captionStart="FIGURE 6" captionStartId="10.[151,250,1977,1999]" captionTargetBox="[150,1436,589,1955]" captionTargetId="figure@10.[150,1436,589,1955]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal view of the neurocranium and the nuchal plate of Chinchaysuyoa labiata, (A) paratype 113 mm SL (CAS 60621), and Chinchaysuyoa ortegai, (B) paratype 155 mm SL (MPEG 35976)." httpUri="https://zenodo.org/record/2622980/files/figure.png" pageId="4" pageNumber="365">6</figureCitation>
and
<tableCitation id="C6FF033BEC77FFAFF95299B2FE07FE45" box="[341,424,391,416]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="4" pageNumber="365">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC77FFAFF89099E4FEE2FD59" pageId="4" pageNumber="365" type="reference_group">
<paragraph id="8BC23680EC77FFAFF89099E4FA34FDC6" blockId="4.[151,1436,464,701]" pageId="4" pageNumber="365">
<treatmentCitationGroup id="AB6D11AEEC77FFAFF89099E4FA34FDC6" pageId="4" pageNumber="365">
<taxonomicName id="4C7D4D03EC77FFAFF89099E4FE4AFE03" authority="Boulenger 1898: 6" authorityName="Boulenger" authorityPageNumber="6" authorityYear="1898" box="[151,485,464,486]" class="Actinopterygii" family="Ariidae" genus="Arius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="species" species="labiatus">
<emphasis id="B909EA92EC77FFAFF89099E4FE88FE03" box="[151,295,464,486]" italics="true" pageId="4" pageNumber="365">Arius labiatus</emphasis>
<treatmentCitation id="0ADC1091EC77FFAFF92999E5FE4AFE03" author="Boulenger, G. A." box="[302,485,464,486]" page="6" pageId="4" pageNumber="365" year="1898">
<bibRefCitation id="EFEC4B71EC77FFAFF92999E5FE7AFE03" author="Boulenger, G. A." box="[302,469,464,486]" pageId="4" pageNumber="365" pagination="1 - 3" refId="ref7112" refString="Boulenger, G. A. (1898) Viaggio del Dr. Enrico Festa nell' Ecuador e regioni vicine. Poissons de l'Equateur. [Part I]. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 13, 1 - 3." type="journal article" year="1898">Boulenger 1898</bibRefCitation>
:6
</treatmentCitation>
</taxonomicName>
(original description; Río Peripa,
<collectingCountry id="F36A7610EC77FFAFFB4399E5FC34FE03" box="[836,923,464,486]" name="Ecuador" pageId="4" pageNumber="365">Ecuador</collectingCountry>
).—
<treatmentCitation id="0ADC1091EC77FFAFFBB999E5FB21FE03" author="Eigenmann, C. H." box="[958,1166,464,486]" page="31" pageId="4" pageNumber="365" year="1922">
<bibRefCitation id="EFEC4B71EC77FFAFFBB999E5FBDDFE03" author="Eigenmann, C. H." box="[958,1138,464,486]" pageId="4" pageNumber="365" pagination="1 - 346" refId="ref7403" refString="Eigenmann, C. H. (1922) The fishes of western South America, Part I. The fresh-water fishes of northwestern South America, including Colombia, Panama, and the Pacific slopes of Ecuador and Peru, together with an appendix upon the fishes of the Rio Meta in Colombia. Memoirs of the Carnegie Museum, 9, 1 - 346." type="journal article" year="1922">Eigenmann 1922</bibRefCitation>
:31
</treatmentCitation>
(description).—
<treatmentCitation id="0ADC1091EC77FFAFFD3299E5FED1FDE1" author="Kailola, P. J. &amp; Bussing, W. A." page="866" pageId="4" pageNumber="365" year="1995">
<bibRefCitation id="EFEC4B71EC77FFAFFD3299E5FEFDFDE1" author="Kailola, P. J. &amp; Bussing, W. A." pageId="4" pageNumber="365" pagination="860 - 886" refId="ref7919" refString="Kailola, P. J. &amp; Bussing, W. A. (1995) Ariidae. In: Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. &amp; Niem, V. H. (Eds.), Guia FAO para la identificacion para loses de la pesca. Pacifico Centro-oriental. Fol. II. Fertebrados-Parte 1. FAO, Rome, pp. 860 - 886." type="book chapter" year="1995">Kailola &amp; Bussing 1995</bibRefCitation>
:866
</treatmentCitation>
(as status uncertain).—
<treatmentCitation id="0ADC1091EC77FFAFFA7399DBFC1AFDE1" author="Marceniuk, P. A. &amp; Ferraris, C. J. Jr." box="[628,949,494,516]" page="453" pageId="4" pageNumber="365" year="2003">
<bibRefCitation id="EFEC4B71EC77FFAFFA7399DBFC25FDE1" author="Marceniuk, P. A. &amp; Ferraris, C. J. Jr." box="[628,906,494,516]" pageId="4" pageNumber="365" refId="ref8265" refString="Marceniuk, P. A. &amp; Ferraris, C. J. Jr. (2003) Check list of the freshwater fishes of South and Central America. CLOFFSCA. EDIPUCRS, Porto Alegre, xi + 729 pp. [Reis, R. E., Kullander, S. O. &amp; Ferraris, C. J. Jr. (Eds.)]" type="book" year="2003">Marceniuk &amp; Ferraris 2003</bibRefCitation>
:453
</treatmentCitation>
(as
<emphasis id="B909EA92EC77FFAFFBD999C5FB33FDE1" box="[990,1180,494,517]" italics="true" pageId="4" pageNumber="365">species inquirenda</emphasis>
).—
<treatmentCitation id="0ADC1091EC77FFAFFCC699DAFE9DFDC6" ID-DOI="http://doi.org/10.5281/zenodo.3501149" ID-Zenodo-Dep="3501149" author="Betancur-R., R. &amp; Acero, P. A." httpUri="http://treatment.plazi.org/id/D67609330691A85BB58C89468F980575" page="13" pageId="4" pageNumber="365" year="2004">
<bibRefCitation id="EFEC4B71EC77FFAFFCC699DAFEB8FDC6" author="Betancur-R., R. &amp; Acero, P. A." pageId="4" pageNumber="365" pagination="1 - 20" refId="ref6612" refString="Betancur-R., R. &amp; Acero, P. A. (2004) Description of Notarius biffi n. sp. and redescription of N. insculptus (Jordan and Gilbert) (Siluriformes: Ariidae) from the eastern Pacific, with evidence of monophyly and limits of Notarius. Zootaxa, 703 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 703.1.1" type="journal article" year="2004">Betancur-R. &amp; Acero P. 2004</bibRefCitation>
:13
</treatmentCitation>
(apparently a species of
<taxonomicName id="4C7D4D03EC77FFAFFA299A3BFD0BFDC6" ID-CoL="6V59" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[558,676,526,547]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC77FFAFFA299A3BFD0BFDC6" box="[558,676,526,547]" italics="true" pageId="4" pageNumber="365">Potamarius</emphasis>
</taxonomicName>
).—
<treatmentCitation id="0ADC1091EC77FFAFFACF9A38FBA1FDC6" ID-DOI="http://doi.org/10.5281/zenodo.3508661" ID-Zenodo-Dep="3508661" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[712,1038,525,547]" httpUri="http://treatment.plazi.org/id/2FEFE315D9E582DB63DE4068BFA422D1" page="98" pageId="4" pageNumber="365" year="2007">
<bibRefCitation id="EFEC4B71EC77FFAFFACF9A38FC5FFDC6" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[712,1008,525,547]" pageId="4" pageNumber="365" pagination="1 - 26" refId="ref8465" refString="Marceniuk, A. P. &amp; Menezes, N. A. (2007) Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa, 1416 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1416.1.1" type="journal article" year="2007">Marceniuk &amp; Menezes 2007</bibRefCitation>
:98
</treatmentCitation>
(as
<emphasis id="B909EA92EC77FFAFFC3F9A3BFB5AFDC6" box="[1080,1269,525,547]" italics="true" pageId="4" pageNumber="365">species inquirenda</emphasis>
in
<taxonomicName id="4C7D4D03EC77FFAFFD129A3BFA24FDC6" ID-CoL="6V59" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[1301,1419,526,547]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC77FFAFFD129A3BFA24FDC6" box="[1301,1419,526,547]" italics="true" pageId="4" pageNumber="365">Potamarius</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC23680EC77FFAFF8909A19FEE2FD59" blockId="4.[151,1436,464,701]" pageId="4" pageNumber="365">
<treatmentCitationGroup id="AB6D11AEEC77FFAFF8909A19FEE2FD59" pageId="4" pageNumber="365">
<taxonomicName id="4C7D4D03EC77FFAFF8909A19FD79FDA7" ID-CoL="3LGKM" authority="Fisher &amp; Eigenmann 1922: 30" authorityName="Fisher &amp; Eigenmann" authorityPageNumber="30" authorityYear="1922" box="[151,726,556,578]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC77FFAFF8909A19FE3BFDA7" box="[151,404,556,578]" italics="true" pageId="4" pageNumber="365">Hexanematichthys henni</emphasis>
Fisher &amp;
<treatmentCitation id="0ADC1091EC77FFAFFA029A19FD79FDA7" author="Eigenmann, C. H." box="[517,726,556,578]" page="30" pageId="4" pageNumber="365" year="1922">
<bibRefCitation id="EFEC4B71EC77FFAFFA029A19FD18FDA7" author="Eigenmann, C. H." box="[517,695,556,578]" pageId="4" pageNumber="365" pagination="1 - 346" refId="ref7403" refString="Eigenmann, C. H. (1922) The fishes of western South America, Part I. The fresh-water fishes of northwestern South America, including Colombia, Panama, and the Pacific slopes of Ecuador and Peru, together with an appendix upon the fishes of the Rio Meta in Colombia. Memoirs of the Carnegie Museum, 9, 1 - 346." type="journal article" year="1922">Eigenmann 1922</bibRefCitation>
:30
</treatmentCitation>
</taxonomicName>
(original description; Río Daule, Colimes).—
<treatmentCitation id="0ADC1091EC77FFAFFCBF9A19FADAFDA7" author="Burgess, W. E." box="[1208,1397,556,578]" page="168" pageId="4" pageNumber="365" year="1989">
<bibRefCitation id="EFEC4B71EC77FFAFFCBF9A19FAE4FDA7" author="Burgess, W. E." box="[1208,1355,556,578]" pageId="4" pageNumber="365" refId="ref7205" refString="Burgess, W. E. (1989) An atlas of freshwater and marine catfishes. A preliminary survey of the Siluriformes. T. F. H. Publications, Neptune City, New Jersey, 784 pp., 285 pls." type="book" year="1989">Burgess 1989</bibRefCitation>
:168
</treatmentCitation>
(as status uncertain).—Kailola &amp; Bussing in Fischer
<emphasis id="B909EA92EC77FFAFFAC39A7EFD5DFD85" box="[708,754,586,608]" italics="true" pageId="4" pageNumber="365">et al</emphasis>
. 1995:866 (as status uncertain).—
<treatmentCitation id="0ADC1091EC77FFAFFC559A7FFA33FD85" author="Marceniuk, P. A. &amp; Ferraris, C. J. Jr." box="[1106,1436,586,608]" page="453" pageId="4" pageNumber="365" year="2003">
<bibRefCitation id="EFEC4B71EC77FFAFFC559A7FFADEFD85" author="Marceniuk, P. A. &amp; Ferraris, C. J. Jr." box="[1106,1393,586,608]" pageId="4" pageNumber="365" refId="ref8265" refString="Marceniuk, P. A. &amp; Ferraris, C. J. Jr. (2003) Check list of the freshwater fishes of South and Central America. CLOFFSCA. EDIPUCRS, Porto Alegre, xi + 729 pp. [Reis, R. E., Kullander, S. O. &amp; Ferraris, C. J. Jr. (Eds.)]" type="book" year="2003">Marceniuk &amp; Ferraris 2003</bibRefCitation>
:453
</treatmentCitation>
(as
<emphasis id="B909EA92EC77FFAFF8ED9A5FFE07FD9A" box="[234,424,617,639]" italics="true" pageId="4" pageNumber="365">species inquirenda</emphasis>
).—
<treatmentCitation id="0ADC1091EC77FFAFF9CA9A5FFCB6FD9A" ID-DOI="http://doi.org/10.5281/zenodo.3501639" ID-Zenodo-Dep="3501639" author="Betancur-R., R. &amp; Acero, P. A." box="[461,793,617,639]" httpUri="http://treatment.plazi.org/id/7090794A063859BBE4B521689117F352" page="13" pageId="4" pageNumber="365" year="2004">
<bibRefCitation id="EFEC4B71EC77FFAFF9CA9A5FFD51FD9A" author="Betancur-R., R. &amp; Acero, P. A." box="[461,766,617,639]" pageId="4" pageNumber="365" pagination="1 - 20" refId="ref6612" refString="Betancur-R., R. &amp; Acero, P. A. (2004) Description of Notarius biffi n. sp. and redescription of N. insculptus (Jordan and Gilbert) (Siluriformes: Ariidae) from the eastern Pacific, with evidence of monophyly and limits of Notarius. Zootaxa, 703 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 703.1.1" type="journal article" year="2004">Betancur-R. &amp; Acero P. 2004</bibRefCitation>
:13
</treatmentCitation>
(as apparently a species of
<taxonomicName id="4C7D4D03EC77FFAFFC289A5FFB0AFD9A" ID-CoL="6V59" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[1071,1189,618,639]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC77FFAFFC289A5FFB0AFD9A" box="[1071,1189,618,639]" italics="true" pageId="4" pageNumber="365">Potamarius</emphasis>
</taxonomicName>
).—
<treatmentCitation id="0ADC1091EC77FFAFFCCE9A5CFE89FD7B" author="Jimenez-Prado, P. &amp; Aguirre, W. &amp; Laaz-Moncayo, E. &amp; Navarrete-Amaya, R. &amp; Nugra-Salazar, F. &amp; Rebolledo-Monsalve, E. &amp; Hugo, E. &amp; Torres-Noboa, A. &amp; Valdiviezo-Rivera, J." page="398" pageId="4" pageNumber="365" year="2015">
<bibRefCitation id="EFEC4B71EC77FFAFFCCE9A5CFF53FD7B" author="Jimenez-Prado, P. &amp; Aguirre, W. &amp; Laaz-Moncayo, E. &amp; Navarrete-Amaya, R. &amp; Nugra-Salazar, F. &amp; Rebolledo-Monsalve, E. &amp; Hugo, E. &amp; Torres-Noboa, A. &amp; Valdiviezo-Rivera, J." pageId="4" pageNumber="365" refId="ref7784" refString="Jimenez-Prado, P., Aguirre, W., Laaz-Moncayo, E., Navarrete-Amaya, R., Nugra-Salazar, F., Rebolledo-Monsalve, E., Zarate- Hugo, E., Torres-Noboa, A. &amp; Valdiviezo-Rivera, J. (2015) Guia de peces para aguas continentales en la vertiente occidental del Ecuador. Pontificia Universidad Catolica del Ecuador Sede Esmeraldas. Universidad del Azuay y Museo Ecuatoriano de Ciencias Naturales del Instituto Nacional de Biodiversidad, Esmeraldas, Ecuador, 416 pp." type="book" year="2015">
Jiménez-Prado
<emphasis id="B909EA92EC77FFAFFD6D9A5FFA39FD9A" box="[1386,1430,617,639]" italics="true" pageId="4" pageNumber="365">et al</emphasis>
. 2015
</bibRefCitation>
:398
</treatmentCitation>
(as apparently a species of
<taxonomicName id="4C7D4D03EC77FFAFFA7F9ABCFD41FD7B" ID-CoL="6V59" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[632,750,649,670]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC77FFAFFA7F9ABCFD41FD7B" box="[632,750,649,670]" italics="true" pageId="4" pageNumber="365">Potamarius</emphasis>
</taxonomicName>
).—
<treatmentCitation id="0ADC1091EC77FFAFFB159ABDFBDBFD7B" ID-DOI="http://doi.org/10.5281/zenodo.3508369" ID-Zenodo-Dep="3508369" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[786,1140,648,670]" httpUri="http://treatment.plazi.org/id/F00488A90945D9174C26C4E57D6DE097" page="98" pageId="4" pageNumber="365" year="2007">
<bibRefCitation id="EFEC4B71EC77FFAFFB159ABDFBF8FD7B" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[786,1111,648,670]" pageId="4" pageNumber="365" pagination="1 - 26" refId="ref8465" refString="Marceniuk, A. P. &amp; Menezes, N. A. (2007) Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa, 1416 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1416.1.1" type="journal article" year="2007">Marceniuk &amp; Menezes 2007</bibRefCitation>
:98
</treatmentCitation>
(as
<emphasis id="B909EA92EC77FFAFFCB69ABCFAD7FD7B" box="[1201,1400,648,670]" italics="true" pageId="4" pageNumber="365">species inquirenda</emphasis>
in
<taxonomicName id="4C7D4D03EC77FFAFF8C09A9DFE92FD58" ID-CoL="6V59" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[199,317,680,701]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC77FFAFF8C09A9DFE92FD58" box="[199,317,680,701]" italics="true" pageId="4" pageNumber="365">Potamarius</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC77FFAFF8909ADCFD28FC53" pageId="4" pageNumber="365" type="diagnosis">
<paragraph id="8BC23680EC77FFAFF8909ADCFD28FC53" blockId="4.[151,1437,745,1994]" pageId="4" pageNumber="365">
<emphasis id="B909EA92EC77FFAFF8909ADCFEBEFCE7" bold="true" box="[151,273,745,770]" pageId="4" pageNumber="365">Diagnosis.</emphasis>
<taxonomicName id="4C7D4D03EC77FFAFF91D9ADCFDB1FCE4" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[282,542,745,769]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B909EA92EC77FFAFF91D9ADCFDB1FCE4" box="[282,542,745,769]" italics="true" pageId="4" pageNumber="365">Chinchaysuyoa labiata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7D4D03EC77FFAFFABF9ADFFC84FCE4" authorityName="Marceniuk &amp; Marchena &amp; Oliveira &amp; Betancur-R" authorityYear="2019" box="[696,811,746,769]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="365" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="B909EA92EC77FFAFFABF9ADFFC84FCE4" box="[696,811,746,769]" italics="true" pageId="4" pageNumber="365">C. ortegai</emphasis>
</taxonomicName>
by finer granulation homogeneously distributed in the cephalic shield (
<emphasis id="B909EA92EC77FFAFF9549B3BFEC5FCC0" box="[339,362,782,805]" italics="true" pageId="4" pageNumber="365">vs</emphasis>
. thicker and irregularly distributed in the cephalic shield, especially in the sphenotic, pterotic and parieto-supraoccipital,
<figureCitation id="13462A05EC77FFAFF9D39B04FD8AFCAF" box="[468,549,817,842]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="4" pageNumber="365">Figs. 5</figureCitation>
and
<figureCitation id="13462A05EC77FFAFFA609B04FDD8FCAF" box="[615,631,817,842]" captionStart="FIGURE 6" captionStartId="10.[151,250,1977,1999]" captionTargetBox="[150,1436,589,1955]" captionTargetId="figure@10.[150,1436,589,1955]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal view of the neurocranium and the nuchal plate of Chinchaysuyoa labiata, (A) paratype 113 mm SL (CAS 60621), and Chinchaysuyoa ortegai, (B) paratype 155 mm SL (MPEG 35976)." httpUri="https://zenodo.org/record/2622980/files/figure.png" pageId="4" pageNumber="365">6</figureCitation>
), snout pointed anteriorly (
<emphasis id="B909EA92EC77FFAFFBB99B07FC7AFCAC" box="[958,981,818,841]" italics="true" pageId="4" pageNumber="365">vs</emphasis>
. truncated anteriorly,
<figureCitation id="13462A05EC77FFAFFCDA9B04FA8BFCAF" box="[1245,1316,817,842]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="4" pageNumber="365">Fig. 5</figureCitation>
), distance between the anterior nostrils 5.1–5.9% SL (
<emphasis id="B909EA92EC77FFAFFA859B63FD36FC88" box="[642,665,854,877]" italics="true" pageId="4" pageNumber="365">vs</emphasis>
. 6.3–7.6% SL,
<tableCitation id="C6FF033BEC77FFAFFB4E9B60FC33FC8B" box="[841,924,853,878]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="4" pageNumber="365">Table 1</tableCitation>
), distance between the posterior nostrils 5.2– 6.2% SL (
<emphasis id="B909EA92EC77FFAFF90C9B4FFE8DFC74" box="[267,290,890,913]" italics="true" pageId="4" pageNumber="365">vs</emphasis>
. 6.4–7.5% SL, rarely less than 6.4,
<tableCitation id="C6FF033BEC77FFAFFABF9B4CFCA4FC77" box="[696,779,889,914]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="4" pageNumber="365">Table 1</tableCitation>
), caudal–fin lobes relatively long, pointed posteriorly (
<emphasis id="B909EA92EC77FFAFFD7A9B4FFA3BFC74" box="[1405,1428,890,913]" italics="true" pageId="4" pageNumber="365">vs</emphasis>
. relatively short, rounded posteriorly,
<figureCitation id="13462A05EC77FFAFFA319BA8FDD5FC53" box="[566,634,925,950]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="4" pageNumber="365">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC77FFADF8C09BF4FB1CFB26" lastPageId="6" lastPageNumber="367" pageId="4" pageNumber="365" type="description">
<paragraph id="8BC23680EC77FFAFF8C09BF4FD6FFAA7" blockId="4.[151,1437,745,1994]" pageId="4" pageNumber="365">
<emphasis id="B909EA92EC77FFAFF8C09BF4FEFEFC3F" bold="true" box="[199,337,961,986]" pageId="4" pageNumber="365">Description</emphasis>
. Morphometric data in
<tableCitation id="C6FF033BEC77FFAFFA5F9BF4FD04FC3F" box="[600,683,961,986]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="4" pageNumber="365">Table 1</tableCitation>
. Head depressed, profile elevated posteriorly, straight at frontal and parieto-supraoccipital area. Snout long and anteriorly pointed. Anterior nostril round, with fleshy edge, posterior nostril covered by flap of skin; nostrils relatively close to one another and distant to orbit, not connected by fleshy furrow. Eye lateral and large; eyes relatively close to one another. Three pairs of moderately long teretiform barbels; maxillary barbel usually reaching gill membrane edge, mental barbels not reaching gill membrane edge. Osseous bridge formed by lateral ethmoid and frontal relatively long and slender, little evident under skin. Cephalic shield exposed, gently granulated; long and moderately large on lateral ethmoid, frontal, and supracleithral areas. Anterior portion of dorsomedial groove of neurocranium fleshy, inconspicuous and not continuous to level of posterior nares; posterior portion of osseous groove shallow, with straight margins, narrowing posteriorly. Parietosupraoccipital process funnel-shaped, long and moderately wide on posterior portion, and profile straight. Nuchal plate crescent-shaped, long and moderately wide.
</paragraph>
<paragraph id="8BC23680EC77FFAFF8C09D78FAA5F9FF" blockId="4.[151,1437,745,1994]" pageId="4" pageNumber="365">Mouth subterminal to terminal, moderately large; lower jaw arched. Lips thick, upper lip more thick than lower lip. Vomerine tooth plates absent. One pair of oval shaped accessory tooth plates, very small and transversely elongate and narrow, distant one another, with sharp teeth. Premaxilla with sharp teeth. Dentary separated at midline, with sharp teeth. Gill membranes fused, attached to isthmus. Nineteen acicular gill rakers on first arch, 21–23 spike-shaped gill rakers on second arch. Mesial surfaces of all gill arches with developed gill rakers, lateral and mesial surfaces of first and second gill arches lacking fleshy papillae intercalated with gill rakers.</paragraph>
<paragraph id="8BC23680EC77FFAFF8C09E10FE06F82F" blockId="4.[151,1437,745,1994]" pageId="4" pageNumber="365">Body width little wider than depth in pectoral-girdle area, progressively more compressed from pectoral fin to caudal peduncle. Lateral line sloping ventrally on anterior one-third, extending posteriorly to caudal peduncle, bending abruptly onto dorsal lobe of caudal fin. Dorsal-fin spine moderately long and thick; anterior margin with granules on basal two-thirds and distal one-third with serrations; posterior margin smooth on basal one-third and distal two-thirds with serrations. Seven soft dorsal-fin rays. Pectoral-fin spine moderately long, shorter than dorsalfin spine; anterior margin with granules on basal two-thirds and distal one-third with short serrations; posterior margin straight on basal one-fourth, distal three-fourths with serrations. Eleven or twelve soft pectoral-fin rays. Posterior cleithral process exposed, rough and triangular shaped, very long and pointed posteriorly. Pelvic fin deep and moderately long at base, with six rays. Adipose-fin base moderately long, its base about half as long as the anal-fin base. Anal fin deep and moderately long at base, with 17–9 rays and distal margin slightly concave. Caudal peduncle relatively low. Caudal fin forked, dorsal and ventral lobes long, wide and pointed posteriorly; dorsal lobe longer than ventral lobe.</paragraph>
<caption id="DF026608EC76FFAEF8909FA9FC66F815" httpUri="https://zenodo.org/record/2622969/files/figure.png" pageId="5" pageNumber="366" startId="5.[151,250,1948,1970]" targetBox="[208,1344,214,1925]" targetPageId="5">
<paragraph id="8BC23680EC76FFAEF8909FA9FC66F815" blockId="5.[151,1436,1948,2032]" pageId="5" pageNumber="366">
<emphasis id="B909EA92EC76FFAEF8909FA9FEA1F854" bold="true" box="[151,270,1948,1970]" pageId="5" pageNumber="366">FIGURE 1</emphasis>
. Phylogenetic tree inferred from combined evidence from molecular and morphological data based on Parsimony Analysis (A), Bayesian inference (B) and Maximum Likelihood (C), with bootstrap values greater than 50%. In red, neotropical freshwater species, included in
<taxonomicName id="4C7D4D03EC76FFAEF9D29FEEFC6AF815" authority="sensu Marceniuk &amp; Menezes (2007)" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[469,965,2010,2032]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="366" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC76FFAEF9D29FEEFDE4F815" box="[469,587,2011,2032]" italics="true" pageId="5" pageNumber="366">Potamarius</emphasis>
sensu Marceniuk &amp; Menezes (2007)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF026608EC75FFADF8909C33FD24FBBF" httpUri="https://zenodo.org/record/2622971/files/figure.png" pageId="6" pageNumber="367" startId="6.[151,250,1030,1052]" targetBox="[153,1427,204,1002]" targetPageId="6">
<paragraph id="8BC23680EC75FFADF8909C33FD24FBBF" blockId="6.[151,1436,1030,1114]" pageId="6" pageNumber="367">
<emphasis id="B909EA92EC75FFADF8909C33FEBDFBFE" bold="true" box="[151,274,1030,1052]" pageId="6" pageNumber="367">FIGURE 2.</emphasis>
Phylogenetic tree inferred from morphological data (Marceniuk
<emphasis id="B909EA92EC75FFADFBA79C32FC63FBF9" box="[928,972,1030,1052]" italics="true" pageId="6" pageNumber="367">et al</emphasis>
., 2012) based on Parsimony Analysis (A) and molecular data (Betancur–R. 2009) based on Bayesian inference (B). In red, neotropical freshwater species, included in
<taxonomicName id="4C7D4D03EC75FFADF8909C70FD29FBBF" authority="sensu Marceniuk &amp; Menezes (2007)" authorityName="sensu Marceniuk &amp; Menezes" authorityYear="2007" box="[151,646,1092,1114]" class="Actinopterygii" family="Ariidae" genus="Potamarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="genus">
<emphasis id="B909EA92EC75FFADF8909C70FEA2FBBF" box="[151,269,1093,1114]" italics="true" pageId="6" pageNumber="367">Potamarius</emphasis>
sensu Marceniuk &amp; Menezes (2007)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC23680EC75FFADF8C09CB3FB1CFB26" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="367">
<emphasis id="B909EA92EC75FFADF8C09CB3FEE7FB7A" bold="true" box="[199,328,1158,1183]" pageId="6" pageNumber="367">Coloration</emphasis>
. In alcohol, brown or dark brown on dorsum, flanks brownish to silvery and white on venter; fins brownish or dark brownish. Maxillary barbel light brownish, mental barbel lighter (
<figureCitation id="13462A05EC75FFADFC389C9FFB09FB26" box="[1087,1190,1194,1219]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="6" pageNumber="367">Fig. 4a,b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC75FFADF8C09CFBFADFFAB6" pageId="6" pageNumber="367" type="distribution">
<paragraph id="8BC23680EC75FFADF8C09CFBFADFFAB6" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="367">
<emphasis id="B909EA92EC75FFADF8C09CFBFDADFB02" bold="true" box="[199,514,1230,1255]" pageId="6" pageNumber="367">Distribution and habitat</emphasis>
.
<taxonomicName id="4C7D4D03EC75FFADFA1B9CFBFC83FB03" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[540,812,1230,1254]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B909EA92EC75FFADFA1B9CFBFC83FB03" box="[540,812,1230,1254]" italics="true" pageId="6" pageNumber="367">Chinchaysuyoa labiata</emphasis>
</taxonomicName>
is known from middle Río Peripa (
<typeStatus id="54C68822EC75FFADFD099CFBFADDFB02" box="[1294,1394,1230,1255]" pageId="6" pageNumber="367" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7D4D03EC75FFADF8909CC7FE35FAEF" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[151,410,1266,1290]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B909EA92EC75FFADF8909CC7FE35FAEF" box="[151,410,1266,1290]" italics="true" pageId="6" pageNumber="367">Chinchaysuyoa labiata</emphasis>
</taxonomicName>
), middle Río Daule (
<typeStatus id="54C68822EC75FFADFA8F9CC7FD43FAEE" box="[648,748,1266,1291]" pageId="6" pageNumber="367" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7D4D03EC75FFADFB159CC6FCDCFAEF" authorityName="Fisher &amp; Eigenmann" authorityYear="1922" box="[786,883,1266,1290]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC75FFADFB159CC6FCDCFAEF" box="[786,883,1266,1290]" italics="true" pageId="6" pageNumber="367">H. henni</emphasis>
</taxonomicName>
), middle Rio
<collectingRegion id="49B9F862EC75FFADFC179CC6FBCBFAEE" box="[1040,1124,1267,1291]" country="Ecuador" name="Guayas" pageId="6" pageNumber="367">Guayas</collectingRegion>
(
<typeStatus id="54C68822EC75FFADFC749CC6FB7AFAEE" box="[1139,1237,1267,1291]" pageId="6" pageNumber="367" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C7D4D03EC75FFADFCFC9CC6FAF4FAEF" authorityName="Fisher &amp; Eigenmann" authorityYear="1922" box="[1275,1371,1266,1290]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC75FFADFCFC9CC6FAF4FAEF" box="[1275,1371,1266,1290]" italics="true" pageId="6" pageNumber="367">H. henni</emphasis>
</taxonomicName>
), and middle Rio Vinces (non-type specimens) all from
<collectingRegion id="49B9F862EC75FFADFAC09D22FCB4FACA" box="[711,795,1303,1327]" country="Ecuador" name="Guayas" pageId="6" pageNumber="367">Guayas</collectingRegion>
Basin. Based on the know distribution of the species, we infer that the distribution of
<taxonomicName id="4C7D4D03EC75FFADF9D49D0FFD79FAB7" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[467,726,1338,1362]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B909EA92EC75FFADF9D49D0FFD79FAB7" box="[467,726,1338,1362]" italics="true" pageId="6" pageNumber="367">Chinchaysuyoa labiata</emphasis>
</taxonomicName>
is restrict to freshwater habitat from west
<collectingCountry id="F36A7610EC75FFADFCB69D0FFAA1FAB6" box="[1201,1294,1338,1363]" name="Ecuador" pageId="6" pageNumber="367">Ecuador</collectingCountry>
(
<figureCitation id="13462A05EC75FFADFD1A9D0FFACDFAB6" box="[1309,1378,1338,1363]" captionStart="FIGURE 7" captionStartId="11.[151,250,1957,1979]" captionTargetBox="[181,1407,375,1942]" captionTargetId="figure@11.[181,1407,375,1942]" captionTargetPageId="11" captionText="FIGURE 7. Premaxillary and accessory tooth plates of Chinchaysuyoa ortegai, (A) non type specimen, not cataloged, 315 mm SL and (B) paratype 155 mm SL (MPEG 35976)." httpUri="https://zenodo.org/record/2622982/files/figure.png" pageId="6" pageNumber="367">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC75FFADF8C09D6BFD98F8AF" pageId="6" pageNumber="367" type="discussion">
<paragraph id="8BC23680EC75FFADF8C09D6BFD98F8AF" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="367">
<emphasis id="B909EA92EC75FFADF8C09D6BFE9BFA92" bold="true" box="[199,308,1374,1399]" pageId="6" pageNumber="367">Remarks</emphasis>
.
<taxonomicName id="4C7D4D03EC75FFADF9449D6AFE4EFA93" authorityName="Boulenger" authorityYear="1898" box="[323,481,1374,1398]" class="Actinopterygii" family="Ariidae" genus="Arius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiatus">
<emphasis id="B909EA92EC75FFADF9449D6AFE4EFA93" box="[323,481,1374,1398]" italics="true" pageId="6" pageNumber="367">Arius labiatus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFEC4B71EC75FFADFAB39D6BFCD4FA92" author="Boulenger, G. A." box="[692,891,1374,1399]" pageId="6" pageNumber="367" pagination="1 - 3" refId="ref7112" refString="Boulenger, G. A. (1898) Viaggio del Dr. Enrico Festa nell' Ecuador e regioni vicine. Poissons de l'Equateur. [Part I]. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 13, 1 - 3." type="journal article" year="1898">Boulenger (1898)</bibRefCitation>
based on a single specimen deposited at Museo di Zoologia, Instituto di Zoologia e Anatomia Comparata Universitá di Torino,
<collectingCountry id="F36A7610EC75FFADFC1C9DB7FBE1FA7E" box="[1051,1102,1410,1435]" name="Italy" pageId="6" pageNumber="367">Italy</collectingCountry>
(
<typeStatus id="54C68822EC75FFADFC589DB7FB6AFA7E" box="[1119,1221,1410,1435]" pageId="6" pageNumber="367" type="holotype">holotype</typeStatus>
, MZUT 1540,
<figureCitation id="13462A05EC75FFADFD779DB7FF1AFA5A" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="6" pageNumber="367">Fig. 4a</figureCitation>
). Subsequently,
<bibRefCitation id="EFEC4B71EC75FFADF9719D93FDE2FA5A" author="Eigenmann, C. H." box="[374,589,1446,1471]" pageId="6" pageNumber="367" pagination="1 - 346" refId="ref7403" refString="Eigenmann, C. H. (1922) The fishes of western South America, Part I. The fresh-water fishes of northwestern South America, including Colombia, Panama, and the Pacific slopes of Ecuador and Peru, together with an appendix upon the fishes of the Rio Meta in Colombia. Memoirs of the Carnegie Museum, 9, 1 - 346." type="journal article" year="1922">Eigenmann (1922)</bibRefCitation>
described
<taxonomicName id="4C7D4D03EC75FFADFAD69D93FC45FA5B" authorityName="Fisher &amp; Eigenmann" authorityYear="1922" box="[721,1002,1446,1470]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC75FFADFAD69D93FC45FA5B" box="[721,1002,1446,1470]" italics="true" pageId="6" pageNumber="367">Hexanematichthys henni</emphasis>
</taxonomicName>
based on three specimens from Río Daule,
<collectingCountry id="F36A7610EC75FFADF8EC9DFFFEE5FA06" box="[235,330,1482,1507]" name="Ecuador" pageId="6" pageNumber="367">Ecuador</collectingCountry>
, considering that “It is possible that these specimens are the young of
<taxonomicName id="4C7D4D03EC75FFADFC8D9DFEFA83FA07" authorityName="Boulenger" authorityYear="1898" box="[1162,1324,1482,1506]" class="Actinopterygii" family="Ariidae" genus="Arius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiatus">
<emphasis id="B909EA92EC75FFADFC8D9DFEFA83FA07" box="[1162,1324,1482,1506]" italics="true" pageId="6" pageNumber="367">Arius labiatus</emphasis>
</taxonomicName>
, the only known specimen of which is three times as large as our largest”. While the descriptions of both species provide good details on the accessory tooth plates, which have a unique configuration in the ariids from the eastern Pacific, both have been considered as
<emphasis id="B909EA92EC75FFADF9F69E02FD6AF9AB" box="[497,709,1590,1614]" italics="true" pageId="6" pageNumber="367">species inquirenda</emphasis>
in the recent literature (
<bibRefCitation id="EFEC4B71EC75FFADFBDC9E03FB59F9AA" author="Kailola, P. J. &amp; Bussing, W. A." box="[987,1270,1590,1615]" pageId="6" pageNumber="367" pagination="860 - 886" refId="ref7919" refString="Kailola, P. J. &amp; Bussing, W. A. (1995) Ariidae. In: Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. &amp; Niem, V. H. (Eds.), Guia FAO para la identificacion para loses de la pesca. Pacifico Centro-oriental. Fol. II. Fertebrados-Parte 1. FAO, Rome, pp. 860 - 886." type="book chapter" year="1995">Kailola &amp; Bussing 1995</bibRefCitation>
;
<bibRefCitation id="EFEC4B71EC75FFADFD039E03FE9FF996" author="Marceniuk, P. A. &amp; Ferraris, C. J. Jr." pageId="6" pageNumber="367" refId="ref8265" refString="Marceniuk, P. A. &amp; Ferraris, C. J. Jr. (2003) Check list of the freshwater fishes of South and Central America. CLOFFSCA. EDIPUCRS, Porto Alegre, xi + 729 pp. [Reis, R. E., Kullander, S. O. &amp; Ferraris, C. J. Jr. (Eds.)]" type="book" year="2003">Marceniuk &amp; Ferraris 2003</bibRefCitation>
;
<bibRefCitation id="EFEC4B71EC75FFADF93B9E6FFD2DF996" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[316,642,1626,1651]" pageId="6" pageNumber="367" pagination="1 - 26" refId="ref8465" refString="Marceniuk, A. P. &amp; Menezes, N. A. (2007) Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa, 1416 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1416.1.1" type="journal article" year="2007">Marceniuk &amp; Menezes 2007</bibRefCitation>
), due primarily to the small number of samples available in zoological collections and the lack of taxonomic studies of the ariid catfish of the west coast of South America. Here,
<taxonomicName id="4C7D4D03EC75FFADF8909E97FE35F95F" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[151,410,1698,1722]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B909EA92EC75FFADF8909E97FE35F95F" box="[151,410,1698,1722]" italics="true" pageId="6" pageNumber="367">Chinchaysuyoa labiata</emphasis>
</taxonomicName>
is recognized as a valid species and
<taxonomicName id="4C7D4D03EC75FFADFB369E97FBEBF95F" authorityName="Fisher &amp; Eigenmann" authorityYear="1922" box="[817,1092,1698,1722]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC75FFADFB369E97FBEBF95F" box="[817,1092,1698,1722]" italics="true" pageId="6" pageNumber="367">Hexanematichthys henni</emphasis>
</taxonomicName>
(
<figureCitation id="13462A05EC75FFADFC539E97FB38F95E" box="[1108,1175,1698,1723]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="6" pageNumber="367">Fig. 4</figureCitation>
) as its junior synonym, supported by the fine, homogeneous granulation of the cephalic shield (
<figureCitation id="13462A05EC75FFADFBC49EF3FBBEF93A" box="[963,1041,1734,1759]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="6" pageNumber="367">Figs. 4</figureCitation>
and
<figureCitation id="13462A05EC75FFADFC4D9EF3FBF5F93A" box="[1098,1114,1734,1759]" captionStart="FIGURE 6" captionStartId="10.[151,250,1977,1999]" captionTargetBox="[150,1436,589,1955]" captionTargetId="figure@10.[150,1436,589,1955]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal view of the neurocranium and the nuchal plate of Chinchaysuyoa labiata, (A) paratype 113 mm SL (CAS 60621), and Chinchaysuyoa ortegai, (B) paratype 155 mm SL (MPEG 35976)." httpUri="https://zenodo.org/record/2622980/files/figure.png" pageId="6" pageNumber="367">6</figureCitation>
), the narrow snout, which is pointed anteriorly (
<figureCitation id="13462A05EC75FFADF9689EDFFE1CF8E6" box="[367,435,1770,1795]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="6" pageNumber="367">Fig. 5</figureCitation>
), the reduced distance between the nostrils (
<tableCitation id="C6FF033BEC75FFADFBA09EDFFBAEF8E6" box="[935,1025,1770,1795]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="6" pageNumber="367">Table. 1</tableCitation>
), and the well-developed dorsal and ventral lobes of the caudal fin, which are pointed posteriorly, and permit the differentiation of the species from its congeners in the river basins of
<collectingCountry id="F36A7610EC75FFADF9FA9F06FD9BF8AF" box="[509,564,1843,1866]" name="Peru" pageId="6" pageNumber="367">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C367650BEC75FFADF8C09F63FBF3F83E" pageId="6" pageNumber="367" type="materials_examined">
<paragraph id="8BC23680EC75FFADF8C09F63FBF3F83E" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="367">
<emphasis id="B909EA92EC75FFADF8C09F63FE15F88A" bold="true" box="[199,442,1878,1903]" pageId="6" pageNumber="367">Material Examined.</emphasis>
Type-specimens:
<typeStatus id="54C68822EC75FFADFA8C9F63FD57F88A" box="[651,760,1878,1903]" pageId="6" pageNumber="367" type="holotype">Holotype</typeStatus>
:
<taxonomicName id="4C7D4D03EC75FFADFB029F62FC09F88B" authorityName="Boulenger" authorityYear="1898" box="[773,934,1878,1902]" class="Actinopterygii" family="Ariidae" genus="Arius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="labiatus">
<emphasis id="B909EA92EC75FFADFB029F62FC09F88B" box="[773,934,1878,1902]" italics="true" pageId="6" pageNumber="367">Arius labiatus</emphasis>
</taxonomicName>
MZUT 1540 (
<quantity id="4C859B65EC75FFADFC5F9F62FB79F88B" box="[1112,1238,1878,1903]" metricMagnitude="0" metricUnit="m" metricValue="1.475" pageId="6" pageNumber="367" unit="mm" value="1475.0">1, 475 mm</quantity>
SL), Río Peripa,
<collectingCountry id="F36A7610EC75FFADF8909F4FFF58F876" box="[151,247,1914,1939]" name="Ecuador" pageId="6" pageNumber="367">Ecuador</collectingCountry>
.
<typeStatus id="54C68822EC75FFADF9079F4FFEC2F876" box="[256,365,1914,1939]" pageId="6" pageNumber="367" type="holotype">Holotype</typeStatus>
:
<taxonomicName id="4C7D4D03EC75FFADF9709F4FFD24F877" authorityName="Fisher &amp; Eigenmann" authorityYear="1922" box="[375,651,1914,1938]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC75FFADF9709F4FFD24F877" box="[375,651,1914,1938]" italics="true" pageId="6" pageNumber="367">Hexanematichthys henni</emphasis>
</taxonomicName>
CAS 60620 (
<quantity id="4C859B65EC75FFADFB209F4EFC1DF877" box="[807,946,1915,1939]" metricMagnitude="0" metricUnit="m" metricValue="1.1393" pageId="6" pageNumber="367" unit="mm" value="1139.3">1, 139.3 mm</quantity>
SL), Río Daule, Colimes, western
<collectingCountry id="F36A7610EC75FFADFD3E9F4FFA37F876" box="[1337,1432,1914,1939]" name="Ecuador" pageId="6" pageNumber="367">Ecuador</collectingCountry>
.
<typeStatus id="54C68822EC75FFADF8909FAAFF51F852" box="[151,254,1951,1975]" pageId="6" pageNumber="367" type="paratype">Paratype</typeStatus>
:
<taxonomicName id="4C7D4D03EC75FFADF90B9FABFD8BF853" authorityName="Fisher &amp; Eigenmann" authorityYear="1922" box="[268,548,1950,1974]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="367" phylum="Chordata" rank="species" species="henni">
<emphasis id="B909EA92EC75FFADF90B9FABFD8BF853" box="[268,548,1950,1974]" italics="true" pageId="6" pageNumber="367">Hexanematichthys henni</emphasis>
</taxonomicName>
CAS 60621 (2, 113–
<quantity id="4C859B65EC75FFADFB229FABFCD6F853" box="[805,889,1950,1975]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="367" unit="mm" value="25.0">25 mm</quantity>
SL), Rio
<collectingRegion id="49B9F862EC75FFADFBF39FAAFBE7F852" box="[1012,1096,1951,1975]" country="Ecuador" name="Guayas" pageId="6" pageNumber="367">Guayas</collectingRegion>
basin, Rio Daule. Non-Type specimens: MCZ 48771(
<quantity id="4C859B65EC75FFADF9B39FF6FD89F83F" box="[436,550,1987,2011]" metricMagnitude="0" metricUnit="m" metricValue="1.183" pageId="6" pageNumber="367" unit="mm" value="1183.0">1, 183 mm</quantity>
SL),
<collectingRegion id="49B9F862EC75FFADFA629FF6FD63F83E" box="[613,716,1986,2011]" country="Ecuador" name="Los Rios" pageId="6" pageNumber="367">Los Rios</collectingRegion>
, Rio Vinces (Rio Nuevo) at Vinces.
</paragraph>
</subSubSection>
</treatment>
</document>
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