675 lines
92 KiB
XML
675 lines
92 KiB
XML
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<mods:title id="6644F3D7AF417FCCBE2FA9F2ADC734B1">Taxonomic revision and phylogeny of the Ophiocoma brevipes group (Echinodermata, Ophiuroidea), with description of a new subgenus (Breviturma) and a new species</mods:title>
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<mods:namePart id="AD7F6D2C30339AF7E5E9DB9203571EDE">Stöhr, Sabine</mods:namePart>
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<mods:affiliation id="C7F02B1F4F07B98BA7511C1E261EFF30">Swedish Museum of Natural History, Department of Zoology, Box 50007, 104 05 Stockholm, Sweden Email: sabine. stohr @ nrm. se (corresponding author) & urn: lsid: zoobank. org: author: 412800 EB-AACE- 4313 - 9810 - 61 F 89 B 740405</mods:affiliation>
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<mods:namePart id="DD88BE1997C5D157B79BFED49BCDBD17">Boissin, Emilie</mods:namePart>
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<mods:affiliation id="8DCCD883C73372F407B0F9D44A374E68">Laboratoire ECOMAR, Université de La Réunion, BP 7151, 97715 Saint-Denis, La Réunion, France & USR 3278 - CRIOBE-CNRS-EPHE, Laboratoire d’Excellence « CORAIL », Université de Perpignan-CBETM, 58 rue Paul Alduy, 66860 Perpignan Cedex & Email: eboissin @ gmail. com & urn: lsid: zoobank. org: author: B 35 FF 544 - AE 44 - 43 B 2 - B 1 CA- 3769 A 281 F 386</mods:affiliation>
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<mods:nameIdentifier id="FF972BF70502977124150702789F2AC2" type="email">eboissin@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A423F63DC8712399DBE721CC55EA71C2">Hoareau, Thierry B.</mods:namePart>
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<mods:affiliation id="15D943316AAA1250AE7F960C40F12A52">Laboratoire ECOMAR, Université de La Réunion, BP 7151, 97715 Saint-Denis, La Réunion, France & Molecular Ecology and Evolution Programme, Department of Genetics, University of Pretoria, Private bag X 20, Hatfield, Pretoria 0028, South Africa & Email: thoareau @ gmail. com & urn: lsid: zoobank. org: author: 44578720 - 231 E- 4 D 99 - 8 F 38 - 2 A 94 B 7 C 4 E 4 F 2</mods:affiliation>
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<mods:title id="CA338DF7A89C032582B793BB1D1C75F6">European Journal of Taxonomy</mods:title>
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<treatment id="03D287F74532842D5F91F959FC11FD27" ID-DOI="http://doi.org/10.5281/zenodo.3846757" ID-GBIF-Taxon="164152680" ID-Zenodo-Dep="3846757" LSID="urn:lsid:plazi:treatment:03D287F74532842D5F91F959FC11FD27" httpUri="http://treatment.plazi.org/id/03D287F74532842D5F91F959FC11FD27" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
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<taxonomicName id="4C7B4D62453284315F91F959FBE0F900" authority="Muller & Troschel, 1842" authorityName="Muller & Troschel" authorityYear="1842" box="[443,1145,1726,1753]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="dentata" subGenus="Breviturma">
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<emphasis id="B90FEAF3453284315F91F959FDDEF900" bold="true" box="[443,583,1726,1753]" italics="true" pageId="12" pageNumber="13">Ophiocoma</emphasis>
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(
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<emphasis id="B90FEAF3453284315C72F959FD7AF901" bold="true" box="[600,739,1726,1752]" italics="true" pageId="12" pageNumber="13">Breviturma</emphasis>
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)
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<emphasis id="B90FEAF3453284315CDEF959FCD7F900" bold="true" box="[756,846,1726,1753]" italics="true" pageId="12" pageNumber="13">dentata</emphasis>
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</emphasis>
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<bibRefCitation id="EFEA4B10453284315D7FF959FBE0F900" author="Muller J. H. & Troschel F. H." box="[853,1145,1726,1753]" pageId="12" pageNumber="13" refId="ref15348" refString="Muller J. H. & Troschel F. H. 1842. System der Asteriden. F. Vieweg & Sohn, Braunschweig." type="book" year="1842">Müller & Troschel, 1842</bibRefCitation>
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</taxonomicName>
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</heading>
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<subSubSection id="C361656A453284315C66F905FC7EF925" box="[588,999,1762,1789]" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BC436E1453284315C66F905FC7EF925" blockId="12.[443,1145,1726,1789]" box="[588,999,1762,1789]" pageId="12" pageNumber="13">
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<figureCitation id="13402A64453284315C66F905FD5FF924" box="[588,710,1762,1789]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="12" pageNumber="13">Figs 2E, F</figureCitation>
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,
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<figureCitation id="13402A64453284315CF8F905FD78F925" box="[722,737,1762,1788]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="12" pageNumber="13">4</figureCitation>
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F-J, 5B, E, H, K, N, Q
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</paragraph>
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<paragraph id="8BC436E1453284315E97F8C3FC22F8E7" blockId="12.[189,955,1828,1961]" box="[189,955,1828,1854]" pageId="12" pageNumber="13">
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<treatmentCitation id="0ADA10F0453284315E97F8C3FC99F8E7" author="Muller J. H. & Troschel F. H." box="[189,768,1828,1854]" page="99" pageId="12" pageNumber="13" year="1842">
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<taxonomicName id="4C7B4D62453284315E97F8C3FC99F8E7" authority="Muller & Troschel, 1842: 99" authorityName="Muller & Troschel" authorityPageNumber="99" authorityYear="1842" box="[189,768,1828,1854]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="dentata">
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<emphasis id="B90FEAF3453284315E97F8C3FE3CF8E7" box="[189,421,1828,1854]" italics="true" pageId="12" pageNumber="13">Ophiocoma dentata</emphasis>
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<bibRefCitation id="EFEA4B10453284315F86F8C3FC99F8E7" author="Muller J. H. & Troschel F. H." box="[428,768,1828,1854]" pageId="12" pageNumber="13" refId="ref15348" refString="Muller J. H. & Troschel F. H. 1842. System der Asteriden. F. Vieweg & Sohn, Braunschweig." type="book" year="1842">Müller & Troschel, 1842: 99</bibRefCitation>
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</taxonomicName>
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, pl. 7, figs 3-3a.
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</paragraph>
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<paragraph id="8BC436E1453284315E97F8AFFD4DF8BB" blockId="12.[189,955,1828,1961]" box="[189,724,1863,1890]" pageId="12" pageNumber="13">
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<taxonomicName id="4C7B4D62453284315E97F8AFFE5DF8B8" authorityName="von Martens" authorityYear="1870" box="[189,452,1864,1890]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="ternispina">
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<emphasis id="B90FEAF3453284315E97F8AFFE5DF8B8" box="[189,452,1864,1890]" italics="true" pageId="12" pageNumber="13">Ophiocoma ternispina</emphasis>
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</taxonomicName>
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v.
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<treatmentCitation id="0ADA10F0453284315FCDF8AFFD49F8BB" author="Martens E. von" box="[487,720,1863,1890]" page="252" pageId="12" pageNumber="13" year="1870">
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<bibRefCitation id="EFEA4B10453284315FCDF8AFFD49F8BB" author="Martens E. von" box="[487,720,1863,1890]" pageId="12" pageNumber="13" pagination="244 - 262" refId="ref15327" refString="Martens E. von 1870. Die Ophiuriden des indischen Oceans. Archiv fur Naturgeschichte 36: 244 - 262." type="journal article" year="1870">Martens, 1870: 252</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC436E1453284315E97F88CFD14F85C" blockId="12.[189,955,1828,1961]" box="[189,653,1899,1926]" pageId="12" pageNumber="13">
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<treatmentCitationGroup id="AB6B11CF453284315E97F88CFD14F85C" box="[189,653,1899,1926]" pageId="12" pageNumber="13">
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<treatmentCitation id="0ADA10F0453284315E97F88CFD13F85C" author="Lyman T." box="[189,650,1899,1926]" page="80" pageId="12" pageNumber="13" year="1862">
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<taxonomicName id="4C7B4D62453284315E97F88CFD13F85C" authority="Lyman, 1862: 80" authorityName="Lyman" authorityPageNumber="80" authorityYear="1862" box="[189,650,1899,1926]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="insularia">
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<emphasis id="B90FEAF3453284315E97F88CFE2EF85C" box="[189,439,1899,1925]" italics="true" pageId="12" pageNumber="13">Ophiocoma insularia</emphasis>
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<bibRefCitation id="EFEA4B10453284315F94F88BFD13F85C" author="Lyman T." box="[446,650,1899,1926]" pageId="12" pageNumber="13" pagination="75 - 86" refId="ref15258" refString="Lyman T. 1862. Descriptions of new Ophiuridae. Proceedings of the Boston Society of Natural History 8: 75 - 86." type="journal article" year="1862">Lyman, 1862: 80</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC436E1453284315E97F868FD10F870" blockId="12.[189,955,1828,1961]" box="[189,649,1935,1961]" pageId="12" pageNumber="13">
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<treatmentCitationGroup id="AB6B11CF453284315E97F868FD10F870" box="[189,649,1935,1961]" pageId="12" pageNumber="13">
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<treatmentCitation id="0ADA10F0453284315E97F868FD1CF870" author="Smith E. A." box="[189,645,1935,1961]" page="39" pageId="12" pageNumber="13" year="1876">
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<taxonomicName id="4C7B4D62453284315E97F868FD1CF870" authority="Smith, 1876: 39" authorityName="Smith" authorityPageNumber="39" authorityYear="1876" box="[189,645,1935,1961]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="variegata">
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<emphasis id="B90FEAF3453284315E97F868FE24F870" box="[189,445,1935,1961]" italics="true" pageId="12" pageNumber="13">Ophiocoma variegata</emphasis>
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<bibRefCitation id="EFEA4B10453284315FEFF868FD1CF870" author="Smith E. A." box="[453,645,1935,1961]" pageId="12" pageNumber="13" pagination="39 - 40" refId="ref15680" refString="Smith E. A. 1876. Descriptions of two new species of Ophiocoma. Annals and Magazine of Natural History, series 4 18: 39 - 40." type="journal article" year="1876">Smith, 1876: 39</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC436E1453284315E97F836FC99F832" blockId="12.[189,768,2001,2027]" box="[189,768,2001,2027]" pageId="12" pageNumber="13">
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<treatmentCitationGroup id="AB6B11CF453284315E97F836FC99F832" box="[189,768,2001,2027]" pageId="12" pageNumber="13">
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non
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<taxonomicName id="4C7B4D62453284315EDAF836FD9CF832" authorityName="Marktanner-Turneretscher" authorityYear="1887" box="[240,517,2001,2027]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="marmorata">
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<emphasis id="B90FEAF3453284315EDAF836FD9CF832" box="[240,517,2001,2027]" italics="true" pageId="12" pageNumber="13">Ophiocoma marmorata</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0ADA10F0453284315C08F836FD65F832" author="Devaney D. M." box="[546,764,2001,2027]" page="17" pageId="12" pageNumber="13" year="1970">
|
||
<bibRefCitation id="EFEA4B10453284315C08F836FD65F832" author="Devaney D. M." box="[546,764,2001,2027]" pageId="12" pageNumber="13" refId="ref14989" refString="Devaney D. M. 1970. Studies on ophiocomid brittlestars. I. A new genus (Clarkcoma) of Ophiocominae with a reevaluation of the genus Ophiocoma. Smithsonian Contributions to Zoology 51, Smithsonian Institution Press, Washington. http: // dx. doi. org / 10.5479 / si. 00810282.51" type="book" year="1970">Devaney 1970: 17</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97FEECFC88FE66" pageId="13" pageNumber="14" type="diagnosis">
|
||
<paragraph id="8BC436E1453384305E97FEECFEAFFEFC" blockId="13.[189,310,267,293]" box="[189,310,267,293]" pageId="13" pageNumber="14">
|
||
<heading id="D08C818D453384305E97FEECFEAFFEFC" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B90FEAF3453384305E97FEECFEAFFEFC" bold="true" box="[189,310,267,293]" pageId="13" pageNumber="14">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97FEDDFC88FE66" blockId="13.[189,1399,314,447]" pageId="13" pageNumber="14">
|
||
Species of
|
||
<taxonomicName id="4C7B4D62453384305F6EFEDDFDE9FE8D" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[324,624,314,340]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="subGenus" status="subgen. nov." subGenus="Breviturma">
|
||
<emphasis id="B90FEAF3453384305F6EFEDDFE54FE8D" box="[324,461,314,340]" italics="true" pageId="13" pageNumber="14">Ophiocoma</emphasis>
|
||
(
|
||
<emphasis id="B90FEAF3453384305FC8FEDCFDFEFE8D" box="[482,615,315,340]" italics="true" pageId="13" pageNumber="14">Breviturma</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453384305C50FEDDFC88FE8D" box="[634,785,314,340]" pageId="13" pageNumber="14" rank="subGenus">subgen. nov.</taxonomicNameLabel>
|
||
characterized by rarely possessing more than four arm spines and if present only at few segments; arm spines never annulated; variable in colour pattern (uniformly or marbled brown, reticulated of spotted); dorsal disc granule density less than 150
|
||
<superScript id="7C0E9BA9453384305B19FE65FAF6FE54" attach="left" box="[1331,1391,383,412]" fontSize="6" pageId="13" pageNumber="14">mm-2</superScript>
|
||
. Maximum size at least
|
||
<quantity id="4C839B04453384305FE7FE42FDB9FE66" box="[461,544,421,447]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="14" unit="mm" value="28.0">28 mm</quantity>
|
||
dd (
|
||
<bibRefCitation id="EFEA4B10453384305C79FE42FC9DFE66" author="Devaney D. M." box="[595,772,421,447]" pageId="13" pageNumber="14" refId="ref14989" refString="Devaney D. M. 1970. Studies on ophiocomid brittlestars. I. A new genus (Clarkcoma) of Ophiocominae with a reevaluation of the genus Ophiocoma. Smithsonian Contributions to Zoology 51, Smithsonian Institution Press, Washington. http: // dx. doi. org / 10.5479 / si. 00810282.51" type="book" year="1970">Devaney 1970</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97FE00FAB2FD4F" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BC436E1453384305E97FE00FE75FDD9" blockId="13.[189,492,486,513]" box="[189,492,486,513]" pageId="13" pageNumber="14">
|
||
<heading id="D08C818D453384305E97FE00FE75FDD9" bold="true" box="[189,492,486,513]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B90FEAF3453384305E97FE00FE75FDD9" bold="true" box="[189,492,486,513]" pageId="13" pageNumber="14">Type material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305EC0FDF1FDD8FD8A" blockId="13.[189,577,533,595]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="3B133CBC453384305EC0FDF1FDD8FD8A" ID-GBIF-Occurrence="2619830301" collectionCode="ZMB" pageId="13" pageNumber="14" specimenCode="ZMB Ech 931" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="B90FEAF3453384305EC0FDF1FD8BFDF6" bold="true" box="[234,530,533,560]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305EC0FDF1FEC5FDE9" box="[234,348,534,560]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="DBDD9E9A453384305F4EFDF1FD8BFDF6" box="[356,530,533,560]" pageId="13" pageNumber="14">ZMB Ech 931</specimenCode>
|
||
</emphasis>
|
||
<typeStatus id="54C08843453384305E97FDDEFEB3FD8A" box="[189,298,569,595]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<emphasis id="B90FEAF3453384305F7BFDDEFDD8FD8A" box="[337,577,569,595]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7B4D62453384305F7BFDDEFDA4FD8A" authorityName="Muller & Troschel" authorityYear="1842" box="[337,573,569,595]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="dentata">Ophiocoma dentata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97FD9CFAB2FD4F" blockId="13.[189,1323,635,662]" box="[189,1323,635,662]" pageId="13" pageNumber="14">
|
||
COLLECTING DATA.
|
||
<collectingRegion id="49BFF803453384305FA5FD9CFE75FD4C" box="[399,492,635,661]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="14">Celebes</collectingRegion>
|
||
(
|
||
<collectingRegion id="49BFF803453384305FD1FD9CFDF2FD4F" box="[507,619,635,662]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="14">Sulawesi</collectingRegion>
|
||
),
|
||
<geoCoordinate id="EE4F5026453384305C55FD9CFC89FD4C" box="[639,784,635,662]" degrees="1" direction="south" minutes="50.875" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-1.8479166">1° 50.875’ S</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026453384305D32FD9CFC50FD4F" box="[792,969,635,662]" degrees="120" direction="east" minutes="31.675" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="120.527916">120° 31.675’ E</geoCoordinate>
|
||
(from atlas), collector Deppe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97FD5AFDC5FCBC" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BC436E1453384305E97FD5AFDC5FCBC" blockId="13.[189,1399,701,870]" pageId="13" pageNumber="14">
|
||
MORPHOLOGICAL DATA.
|
||
<quantity id="4C839B04453384305FEAFD5AFDA0FD0E" box="[448,569,701,727]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="13" pageNumber="14" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18-19 mm</quantity>
|
||
dd (irregular disc), granules on dorsal disc mostly rubbed off, light and dark brown pattern on dorsal disc, dorsal arms medium brown, ventral interradii fully granulated, white, DAP W:L 2:1, arm spine sequence 3, 3, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3, 3, oral shield longer than wide, ventralmost and 2
|
||
<superScript id="7C0E9BA9453384305FF6FCC1FE74FCED" attach="left" box="[476,493,806,820]" fontSize="6" pageId="13" pageNumber="14">nd</superScript>
|
||
ventral spine longest, longer than an arm segment, dorsal spines thickened, ventral ones slender, not annulated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305EC0FC6AFE95FBE8" pageId="13" pageNumber="14" type="materials_examined">
|
||
<materialsCitation id="3B133CBC453384305EC0FC6AFE95FBE8" ID-GBIF-Occurrence="2619830302" collectionCode="ZMB" country="Indonesia" latitude="8.356667" location="Flores-Sea" longLatPrecision="92" longitude="122.963615" municipality="Larantuka" pageId="13" pageNumber="14" specimenCode="ZMB Ech 1815" specimenCount="2" typeStatus="holotype">
|
||
<paragraph id="8BC436E1453384305EC0FC6AFDB8FC7E" blockId="13.[189,608,909,971]" box="[234,545,909,935]" pageId="13" pageNumber="14">
|
||
<emphasis id="B90FEAF3453384305EC0FC6AFDB8FC7E" bold="true" box="[234,545,909,935]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305EC0FC6AFEC5FC7E" box="[234,348,909,935]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="DBDD9E9A453384305F4EFC6AFDB8FC7E" box="[356,545,909,935]" pageId="13" pageNumber="14">ZMB Ech 1815</specimenCode>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97FC56FDF9FC12" blockId="13.[189,608,909,971]" box="[189,608,945,971]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305E97FC56FEB3FC12" box="[189,298,945,971]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<emphasis id="B90FEAF3453384305F7BFC56FDF9FC12" box="[337,608,945,971]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7B4D62453384305F7BFC56FDC5FC12" authorityName="von Martens" authorityYear="1870" box="[337,604,945,971]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="ternispina">Ophiocoma ternispina</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97FC14FE95FBE8" blockId="13.[189,1399,1010,1073]" pageId="13" pageNumber="14">
|
||
COLLECTING DATA.
|
||
<collectingCountry id="F36C7671453384305FBDFC14FD95FBD4" box="[407,524,1011,1037]" name="Indonesia" pageId="13" pageNumber="14">Indonesia</collectingCountry>
|
||
,
|
||
<location id="8EA4603A453384305C36FC14FD04FBD4" LSID="urn:lsid:plazi:treatment:03D287F74532842D5F91F959FC11FD27:8EA4603A453384305C36FC14FD04FBD4" box="[540,669,1011,1037]" country="Indonesia" latitude="8.356667" longLatPrecision="92" longitude="122.963615" municipality="Larantuka" name="Flores-Sea" pageId="13" pageNumber="14">Flores-Sea</location>
|
||
,
|
||
<collectingMunicipality id="6BA0AC9B453384305C86FC14FCBFFBD4" box="[684,806,1011,1037]" pageId="13" pageNumber="14">Larantuka</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F36C7671453384305D1FFC14FC18FBD4" box="[821,897,1011,1037]" name="Indonesia" pageId="13" pageNumber="14">Flores</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE4F5026453384305DBBFC14FB8BFBD4" box="[913,1042,1011,1037]" degrees="8" direction="north" minutes="21.4" orientation="latitude" pageId="13" pageNumber="14" precision="92" value="8.356667">8° 21.4’ N</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026453384305A34FC14FB4FFBD4" box="[1054,1238,1010,1037]" degrees="122" direction="east" minutes="57.817" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="122.963615">122° 57.817’ E</geoCoordinate>
|
||
(from atlas), 1 spm.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97FBBEFDA3FAC6" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BC436E1453384305E97FBBEFC61FB04" blockId="13.[189,1398,1112,1246]" pageId="13" pageNumber="14">
|
||
MORPHOLOGICAL DATA.
|
||
<quantity id="4C839B04453384305FEFFBBFFD80FBAA" box="[453,537,1112,1139]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="mm" value="16.0">16 mm</quantity>
|
||
dd, disc light brown, triangular brown spots and curved lines on DAPs, spines weakly annulated, DAP W:L 2:1, disc irregular round, ventral interradii completely granulated, arm spine sequence 3, 3, 3, (3)4, 4, 4, (4)3, 3, 3, 3, 3, 3, 3, 3, 3,3, 3 (numbers vary between arms), 2
|
||
<superScript id="7C0E9BA9453384305B4FFB79FAEFFB75" attach="none" box="[1381,1398,1182,1196]" fontSize="6" pageId="13" pageNumber="14">nd</superScript>
|
||
ventral spine longest, slender spines, oral shield twice as long as wide.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305EC0FAE2FDA3FAC6" blockId="13.[189,582,1285,1347]" box="[234,570,1285,1311]" pageId="13" pageNumber="14">
|
||
<emphasis id="B90FEAF3453384305EC0FAE2FDA3FAC6" bold="true" box="[234,570,1285,1311]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305EC0FAE2FED6FAC6" box="[234,335,1285,1311]" pageId="13" pageNumber="14" type="syntype">Syntype</typeStatus>
|
||
SMNH-Type-5244
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97FACEFE2FFA70" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BC436E1453384305E97FACEFDDFFA9A" blockId="13.[189,582,1285,1347]" box="[189,582,1320,1347]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305E97FACEFE84FA9A" box="[189,285,1321,1347]" pageId="13" pageNumber="14" type="syntype">Syntype</typeStatus>
|
||
of
|
||
<emphasis id="B90FEAF3453384305F6EFACEFDDFFA9A" box="[324,582,1321,1347]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7B4D62453384305F6EFACEFDDBFA9A" authorityName="Lyman" authorityYear="1862" box="[324,578,1321,1347]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="insularia">Ophiocoma insularia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97FA8CFE2FFA70" blockId="13.[189,1399,1386,1449]" pageId="13" pageNumber="14">
|
||
COLLECTING DATA.
|
||
<collectingRegion id="49BFF803453384305FB8FA8DFE73FA5D" box="[402,490,1386,1412]" country="United States of America" name="Hawaii" pageId="13" pageNumber="14">Hawaii</collectingRegion>
|
||
, 1860, 1 spm (originally donated to
|
||
<collectingRegion id="49BFF803453384305DB5FA8DFB87FA5D" box="[927,1054,1386,1412]" country="Sweden" name="Stockholm" pageId="13" pageNumber="14">Stockholm</collectingRegion>
|
||
by Museum of Comparative Zoology at Harvard).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97FA37FD7FF98C" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BC436E1453384305E97FA37FD7FF98C" blockId="13.[189,1399,1488,1621]" pageId="13" pageNumber="14">
|
||
MORPHOLOGICAL DATA. 17.6 mm dd, disc light brown marbled, arms light brown, dorsal and ventral side similar in colour, disc fully granulated dorsally and ventrally. No annulations on arm spines, spine sequence 3, 3, 3, 4, 4, 4, 5(4), 5(4), 5(4), 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3 (two arms with only 4 spines). Oral shield oval. Granule density 96-120
|
||
<superScript id="7C0E9BA9453384305C89F9DCFD46F99E" attach="left" box="[675,735,1593,1621]" fontSize="6" pageId="13" pageNumber="14">mm-2</superScript>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305EC0F99BFD7BF924" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BC436E1453384305EC0F99BFE67F94F" blockId="13.[189,594,1660,1722]" box="[234,510,1660,1686]" pageId="13" pageNumber="14">
|
||
<emphasis id="B90FEAF3453384305EC0F99BFE67F94F" bold="true" box="[234,510,1660,1686]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305EC0F99BFEC3F94F" box="[234,346,1660,1686]" pageId="13" pageNumber="14" type="syntype">Syntypes</typeStatus>
|
||
MCZN 1668
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97F947FDCBF963" blockId="13.[189,594,1660,1722]" box="[189,594,1696,1722]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305E97F947FEB0F963" box="[189,297,1696,1722]" pageId="13" pageNumber="14" type="syntype">Syntypes</typeStatus>
|
||
of
|
||
<emphasis id="B90FEAF3453384305F7AF947FDCBF963" box="[336,594,1696,1722]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7B4D62453384305F7AF947FDD7F963" authorityName="Lyman" authorityYear="1862" box="[336,590,1696,1722]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="insularia">Ophiocoma insularia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453384305E97F905FD7BF924" blockId="13.[189,738,1762,1789]" box="[189,738,1762,1789]" pageId="13" pageNumber="14">
|
||
COLLECTING DATA.
|
||
<collectingRegion id="49BFF803453384305FA5F905FE7EF925" box="[399,487,1762,1788]" country="United States of America" name="Hawaii" pageId="13" pageNumber="14">Hawaii</collectingRegion>
|
||
, Maui, 1860, 7 spms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384305E97F8C3FE67F81E" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BC436E1453384305E97F8C3FC01F85C" blockId="13.[189,1399,1828,1925]" pageId="13" pageNumber="14">MORPHOLOGICAL DATA. 13.3-16.6 mm dd, all dark brown to black on dorsal disc and arms, including spines, ventral side slightly lighter brown. Both middle spines about equal, longer than dorsal and ventral spines. (Photos provided by Adam Baldinger, MCZN).</paragraph>
|
||
<paragraph id="8BC436E1453384305EC0F84AFE67F81E" blockId="13.[189,594,1965,2027]" box="[234,510,1965,1991]" pageId="13" pageNumber="14">
|
||
<emphasis id="B90FEAF3453384305EC0F84AFE67F81E" bold="true" box="[234,510,1965,1991]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305EC0F84AFEC3F81E" box="[234,346,1965,1991]" pageId="13" pageNumber="14" type="syntype">Syntypes</typeStatus>
|
||
MCZN 1666
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453384325E97F836FD0FFEFF" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BC436E1453384305E97F836FDCBF832" blockId="13.[189,594,1965,2027]" box="[189,594,2001,2027]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54C08843453384305E97F836FEB0F832" box="[189,297,2001,2027]" pageId="13" pageNumber="14" type="syntype">Syntypes</typeStatus>
|
||
of
|
||
<emphasis id="B90FEAF3453384305F7AF836FDCBF832" box="[336,594,2001,2027]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7B4D62453384305F7AF836FDD7F832" authorityName="Lyman" authorityYear="1862" box="[336,590,2001,2027]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="insularia">Ophiocoma insularia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="DF046669453084335E97F997FD23F832" ID-DOI="http://doi.org/10.5281/zenodo.3827684" ID-Zenodo-Dep="3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1648,1674]" targetBox="[257,1330,266,1607]" targetPageId="14">
|
||
<paragraph id="8BC436E1453084335E97F997FD23F832" blockId="14.[189,1399,1648,2027]" pageId="14" pageNumber="15">
|
||
<emphasis id="B90FEAF3453084335E97F997FE97F952" bold="true" box="[189,270,1648,1675]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
|
||
SEM images of skeletal elements of
|
||
<taxonomicName id="4C7B4D62453084335CCAF996FCF0F952" authorityName="L. Agassiz" authorityYear="1835" box="[736,873,1649,1675]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B90FEAF3453084335CCAF996FCF0F952" box="[736,873,1649,1675]" italics="true" pageId="14" pageNumber="15">Ophiocoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C7B4D62453084335D5EF996FF61F976" authority="Peters, 1851" authorityName="Peters" authorityYear="1851" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes" subGenus="Breviturma">
|
||
(
|
||
<emphasis id="B90FEAF3453084335D54F995FB9AF952" box="[894,1027,1650,1675]" italics="true" pageId="14" pageNumber="15">Breviturma</emphasis>
|
||
subgen. nov.)
|
||
<emphasis id="B90FEAF3453084335A90F996FA85F952" box="[1210,1308,1649,1675]" italics="true" pageId="14" pageNumber="15">brevipes</emphasis>
|
||
<bibRefCitation id="EFEA4B10453084335B0DF996FF61F976" author="Peters W." pageId="14" pageNumber="15" pagination="463 - 466" refId="ref15488" refString="Peters W. 1851. Ubersicht der an der Kuste von Mossambique eingesammelten Ophiuren, unter denen sich zwei neue Gattungen befinden. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen der Koniglich Preussischen Akademie der Wissenschaften zu Berlin 1851: 463 - 466." type="journal article" year="1851">Peters, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
(SMNH-133230),
|
||
<taxonomicName id="4C7B4D62453084335FFCF973FC37F977" authority="Muller & Troschel, 1842" authorityName="Muller & Troschel" authorityYear="1842" box="[470,942,1684,1710]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata" subGenus="Breviturma">
|
||
<emphasis id="B90FEAF3453084335FFCF973FE6AF977" box="[470,499,1684,1710]" italics="true" pageId="14" pageNumber="15">O.</emphasis>
|
||
(
|
||
<emphasis id="B90FEAF3453084335C29F972FD84F977" box="[515,541,1685,1710]" italics="true" pageId="14" pageNumber="15">B.</emphasis>
|
||
)
|
||
<emphasis id="B90FEAF3453084335C07F973FD1CF977" box="[557,645,1684,1710]" italics="true" pageId="14" pageNumber="15">dentata</emphasis>
|
||
<bibRefCitation id="EFEA4B10453084335CA6F973FC37F977" author="Muller J. H. & Troschel F. H." box="[652,942,1684,1710]" pageId="14" pageNumber="15" refId="ref15348" refString="Muller J. H. & Troschel F. H. 1842. System der Asteriden. F. Vieweg & Sohn, Braunschweig." type="book" year="1842">Müller & Troschel, 1842</bibRefCitation>
|
||
</taxonomicName>
|
||
(SMNH-133232) and
|
||
<taxonomicName id="4C7B4D62453084335A9CF973FA8FF977" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[1206,1302,1684,1710]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335A9CF973FA8FF977" box="[1206,1302,1684,1710]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335B37F973FAEEF977" box="[1309,1399,1684,1710]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype (SMNH-Type-8535).
|
||
<emphasis id="B90FEAF3453084335C0CF950FDC3F908" bold="true" box="[550,602,1719,1745]" pageId="14" pageNumber="15">A-C</emphasis>
|
||
. Oral plates, abradial aspect at left, orientated with dorsal side up.
|
||
<emphasis id="B90FEAF3453084335B70F950FAF6F908" bold="true" box="[1370,1391,1719,1745]" pageId="14" pageNumber="15">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335E97F93CFEDDF92D" authorityName="Peters" authorityYear="1851" box="[189,324,1754,1781]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453084335E97F93CFEDDF92D" box="[189,324,1754,1781]" italics="true" pageId="14" pageNumber="15">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335F79F93DFEFEF92D" bold="true" box="[339,359,1754,1780]" pageId="14" pageNumber="15">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335F5CF93CFE6AF92D" authorityName="Muller & Troschel" authorityYear="1842" box="[374,499,1754,1781]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3453084335F5CF93CFE6AF92D" box="[374,499,1754,1781]" italics="true" pageId="14" pageNumber="15">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335C28F93DFD8EF92D" bold="true" box="[514,535,1754,1780]" pageId="14" pageNumber="15">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335C0DF93CFD11F92D" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[551,648,1754,1781]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335C0DF93CFD11F92D" box="[551,648,1754,1781]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335CBAF93CFD72F92C" box="[656,747,1755,1781]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B90FEAF3453084335CD9F93DFCBDF92D" bold="true" box="[755,804,1754,1780]" pageId="14" pageNumber="15">D-F</emphasis>
|
||
. Dental plates, external aspect left, internal aspect right, dorsal edge upwards.
|
||
<emphasis id="B90FEAF3453084335FD1F919FD89F8C1" bold="true" box="[507,528,1790,1816]" pageId="14" pageNumber="15">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335C37F919FD39F8CE" authorityName="Peters" authorityYear="1851" box="[541,672,1789,1816]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453084335C37F919FD39F8CE" box="[541,672,1789,1816]" italics="true" pageId="14" pageNumber="15">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335C87F919FD58F8C1" bold="true" box="[685,705,1790,1816]" pageId="14" pageNumber="15">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335CE4F919FCDEF8CE" authorityName="Muller & Troschel" authorityYear="1842" box="[718,839,1789,1816]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3453084335CE4F919FCDEF8CE" box="[718,839,1789,1816]" italics="true" pageId="14" pageNumber="15">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335D7EF919FCFFF8C1" bold="true" box="[852,870,1790,1816]" pageId="14" pageNumber="15">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335D59F919FC4BF8CE" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[883,978,1789,1816]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335D59F919FC4BF8CE" box="[883,978,1789,1816]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335DFDF919FBB6F8C1" box="[983,1071,1790,1816]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B90FEAF3453084335A1EF91AFBFEF8CE" bold="true" box="[1076,1127,1789,1815]" pageId="14" pageNumber="15">G-I.</emphasis>
|
||
Radial shields, internal aspect, proximal edge upwards.
|
||
<emphasis id="B90FEAF3453084335C10F8C7FDC8F8E3" bold="true" box="[570,593,1824,1850]" pageId="14" pageNumber="15">G</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335C4BF8C6FD7EF8E2" authorityName="Peters" authorityYear="1851" box="[609,743,1825,1851]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453084335C4BF8C6FD7EF8E2" box="[609,743,1825,1851]" italics="true" pageId="14" pageNumber="15">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335CDDF8C6FC97F8E2" bold="true" box="[759,782,1825,1851]" pageId="14" pageNumber="15">H</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335D37F8C6FC03F8E2" authorityName="Muller & Troschel" authorityYear="1842" box="[797,922,1825,1851]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3453084335D37F8C6FC03F8E2" box="[797,922,1825,1851]" italics="true" pageId="14" pageNumber="15">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335D80F8C6FC2CF8E2" bold="true" box="[938,949,1825,1851]" pageId="14" pageNumber="15">I</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335DEFF8C6FBBFF8E2" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[965,1062,1825,1851]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335DEFF8C6FBBFF8E2" box="[965,1062,1825,1851]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335A04F8C6FB13F8E2" box="[1070,1162,1825,1851]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B90FEAF3453084335AB8F8C6FB27F8E2" bold="true" box="[1170,1214,1825,1851]" pageId="14" pageNumber="15">J-L</emphasis>
|
||
.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards.
|
||
<emphasis id="B90FEAF3453084335DAFF8A3FC0DF887" bold="true" box="[901,916,1860,1886]" pageId="14" pageNumber="15">J</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335D8EF8A3FBB5F887" authorityName="Peters" authorityYear="1851" box="[932,1068,1860,1886]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453084335D8EF8A3FBB5F887" box="[932,1068,1860,1886]" italics="true" pageId="14" pageNumber="15">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335A16F8A3FBCAF887" bold="true" box="[1084,1107,1860,1886]" pageId="14" pageNumber="15">K</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335A49F8A3FB79F887" authorityName="Muller & Troschel" authorityYear="1842" box="[1123,1248,1860,1886]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3453084335A49F8A3FB79F887" box="[1123,1248,1860,1886]" italics="true" pageId="14" pageNumber="15">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335ADBF8A3FA9CF887" bold="true" box="[1265,1285,1860,1886]" pageId="14" pageNumber="15">L</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335B3EF8A3FAEEF887" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[1300,1399,1860,1886]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335B3EF8A3FAEEF887" box="[1300,1399,1860,1886]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335E97F88FFE85F85B" box="[189,284,1896,1922]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B90FEAF3453084335F0CF880FEFBF858" bold="true" box="[294,354,1895,1921]" pageId="14" pageNumber="15">M-O</emphasis>
|
||
. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields).
|
||
<emphasis id="B90FEAF3453084335F00F86DFEDFF87D" bold="true" box="[298,326,1930,1956]" pageId="14" pageNumber="15">M</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335F7DF86CFE46F87D" authorityName="Peters" authorityYear="1851" box="[343,479,1930,1957]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453084335F7DF86CFE46F87D" box="[343,479,1930,1957]" italics="true" pageId="14" pageNumber="15">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335FDBF86DFD9FF87D" bold="true" box="[497,518,1930,1956]" pageId="14" pageNumber="15">N</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335C3DF86CFD0FF87D" authorityName="Muller & Troschel" authorityYear="1842" box="[535,662,1930,1957]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3453084335C3DF86CFD0FF87D" box="[535,662,1930,1957]" italics="true" pageId="14" pageNumber="15">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335C8DF86DFD27F87D" bold="true" box="[679,702,1930,1956]" pageId="14" pageNumber="15">O</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335CFAF86CFCAAF87D" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[720,819,1930,1957]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335CFAF86CFCAAF87D" box="[720,819,1930,1957]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335D17F86CFC03F87C" box="[829,922,1931,1957]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B90FEAF3453084335D8EF86DFC4CF87D" bold="true" box="[932,981,1930,1956]" pageId="14" pageNumber="15">P-R</emphasis>
|
||
. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right.
|
||
<emphasis id="B90FEAF3453084335DB9F849FC3CF811" bold="true" box="[915,933,1966,1992]" pageId="14" pageNumber="15">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335D99F849FBA1F81E" authorityName="Peters" authorityYear="1851" box="[947,1080,1965,1992]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453084335D99F849FBA1F81E" box="[947,1080,1965,1992]" italics="true" pageId="14" pageNumber="15">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335A6CF84AFBC4F81E" bold="true" box="[1094,1117,1965,1991]" pageId="14" pageNumber="15">Q</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335A41F849FB7FF81E" authorityName="Muller & Troschel" authorityYear="1842" box="[1131,1254,1965,1992]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3453084335A41F849FB7FF81E" box="[1131,1254,1965,1992]" italics="true" pageId="14" pageNumber="15">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90FEAF3453084335ADEF849FA90F811" bold="true" box="[1268,1289,1966,1992]" pageId="14" pageNumber="15">R</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D62453084335B3DF849FAEEF81E" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[1303,1399,1965,1992]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453084335B3DF849FAEEF81E" box="[1303,1399,1965,1992]" italics="true" pageId="14" pageNumber="15">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453084335E97F836FE81F832" box="[189,280,2001,2027]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
For institution codes see main text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC436E1453184325E97FEECFD0FFEFF" blockId="15.[189,662,267,294]" box="[189,662,267,294]" pageId="15" pageNumber="16">
|
||
COLLECTING DATA.
|
||
<collectingRegion id="49BFF803453184325FA5FEECFE7EFEFC" box="[399,487,267,293]" country="United States of America" name="Hawaii" pageId="15" pageNumber="16">Hawaii</collectingRegion>
|
||
, 1860, 2 spms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325E97FEA8FC12FE6A" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="8BC436E1453184325E97FEA8FC12FE6A" blockId="15.[189,1399,334,435]" pageId="15" pageNumber="16">MORPHOLOGICAL DATA. 19.5 and 31.6 mm dd, dark brown disc and arms, with lighter abraded patches on dorsal disc, dark brown on ventral disc and arms; a note among the labels reads “4-5 spines, 3 upper about equal”. (Photos provided by Adam Baldinger, MCZN).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325E97FE3BFE2DFDA8" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="8BC436E1453184325E97FE3BFE60FE2F" blockId="15.[189,505,476,502]" box="[189,505,476,502]" pageId="15" pageNumber="16">
|
||
<heading id="D08C818D453184325E97FE3BFE60FE2F" bold="true" box="[189,505,476,502]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B90FEAF3453184325E97FE3BFE60FE2F" bold="true" box="[189,505,476,502]" pageId="15" pageNumber="16">Other material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453184325EC0FDEBFE3DFDFF" blockId="15.[189,1398,524,625]" box="[234,420,524,550]" pageId="15" pageNumber="16">
|
||
<emphasis id="B90FEAF3453184325EC0FDEBFE3DFDFF" bold="true" box="[234,420,524,550]" pageId="15" pageNumber="16">
|
||
<collectionCode id="ED6AAE24453184325EC0FDEBFED5FDFF" box="[234,332,524,550]" pageId="15" pageNumber="16">SMNH-</collectionCode>
|
||
133232
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453184325E97FDD6FE2DFDA8" blockId="15.[189,1398,524,625]" pageId="15" pageNumber="16">
|
||
COLLECTING DATA.
|
||
<collectingCountry id="F36C7671453184325FA4FDD5FD82FD92" box="[398,539,561,588]" name="Réunion" pageId="15" pageNumber="16">La Réunion</collectingCountry>
|
||
, Etang Salé,
|
||
<geoCoordinate id="EE4F5026453184325C99FDD6FCDCFD92" box="[691,837,561,587]" degrees="21" direction="south" minutes="16" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="17" value="-21.271389">21°16’17” S</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026453184325D66FDD6FC7BFD95" box="[844,994,561,588]" degrees="55" direction="east" minutes="19" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="56" value="55.332222">55°19’56” E</geoCoordinate>
|
||
, collectors E. Boissin, T. Hoareau,
|
||
<date id="FFC51021453184325E97FDB1FEC3FDA9" box="[189,346,598,624]" pageId="15" pageNumber="16" value="2009-02-24">24 Feb. 2009</date>
|
||
, 1 spm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325E97FD7DFC72FA48" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="8BC436E1453184325E97FD7DFC12FCB4" blockId="15.[189,1399,665,877]" pageId="15" pageNumber="16">
|
||
MORPHOLOGICAL DATA.
|
||
<quantity id="4C839B04453184325F94FD7DFD94FD6D" box="[446,525,666,692]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="11.0">11 mm</quantity>
|
||
dd, dorsal disc with a brown multipointed star-shape, white centre spot with brown dots, interradii white with brown marbled pattern (
|
||
<figureCitation id="13402A64453184325D65FD59FC31FD00" box="[847,936,702,729]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="15" pageNumber="16">Fig. 2E</figureCitation>
|
||
); dorsal arm white with rectangular dark brown outlines, spines brown at base, fading into white towards their tip. Ventral interradii brown and white marbled, fully granulated to oral shields, oral frame white, oral shields with 2 parallel longitudinal brown lines. Ventral arm white with median brown dots and short lines. Arm spine sequence 3, 3, 3, 3, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3, 3, 3. Granule density 216-224
|
||
<superScript id="7C0E9BA9453184325D62FCB4FC1DFC86" attach="left" box="[840,900,849,877]" fontSize="6" pageId="15" pageNumber="16">mm-2</superScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453184325E97FC71FC72FA48" blockId="15.[189,1399,916,1425]" pageId="15" pageNumber="16">
|
||
Dorsal disc granule diameter
|
||
<quantity id="4C839B04453184325C0AFC72FD00FC69" box="[544,665,917,944]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" metricValueMax="5.4" metricValueMin="4.9" pageId="15" pageNumber="16" unit="mm" value="51.5" valueMax="54.0" valueMin="49.0">49-54 µm</quantity>
|
||
, granule density 120-144
|
||
<superScript id="7C0E9BA9453184325DFEFC71FB96FC7B" attach="left" box="[980,1039,916,944]" fontSize="6" pageId="15" pageNumber="16">mm-2</superScript>
|
||
, on SEM image (
|
||
<figureCitation id="13402A64453184325AC2FC71FAF1FC69" box="[1256,1384,918,944]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="15" pageNumber="16">Fig. 4F, H</figureCitation>
|
||
); third spine on proximal segments
|
||
<quantity id="4C839B04453184325C7FFC5CFD02FC0C" box="[597,667,955,981]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
|
||
long, DAPs 1.4-1.6 mm wide
|
||
<figureCitation id="13402A64453184325A24FC5CFBF7FC0C" box="[1038,1134,955,981]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="15" pageNumber="16">Fig. 4G</figureCitation>
|
||
). Oral shields as wide as long; 5-6 tooth papillae, 5 lateral oral papillae at each jaw edge, outer papilla widest (
|
||
<figureCitation id="13402A64453184325AF0FC07FAB7FC23" box="[1242,1326,992,1018]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="15" pageNumber="16">Fig. 4I</figureCitation>
|
||
). Few granules on proximal ends of oral plates. Oral plates, adradial distal part with strong folds, abradial face with large muscle flange with horizontal striations (
|
||
<figureCitation id="13402A64453184325D74FBCDFC23FB9D" box="[862,954,1065,1092]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5B</figureCitation>
|
||
), 1.6 mm wide, 2.0 mm high. Dental plate short, externally with two large tooth foramina in dorsal half, dorsalmost foramen oval, wider than long, other foramen square; internally each foramen divided by a wide septum, dorsalmost one into two wide vertical oval slits (septum broken), other foramen into two short parallel holes (
|
||
<figureCitation id="13402A64453184325A89FB7EFB67FB6A" box="[1187,1278,1176,1203]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5E</figureCitation>
|
||
). External ventral half of dental plate with smooth horizontal ridges, forming two wide ovals, the ventralmost one half the size of the other; a cluster of tiny depressions at ventral end of plate (
|
||
<figureCitation id="13402A64453184325A70FB04FB20FB24" box="[1114,1209,1250,1277]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5E</figureCitation>
|
||
<figureCitation id="13402A64453184325A93FB05FB5DFB24" box="[1209,1220,1250,1277]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[195,1380,1071,1817]" captionTargetId="figure@3.[189,1398,1071,1820]" captionTargetPageId="3" captionText="Fig. 1. Phylogenetic reconstruction of Ophiocoma (Breviturma) subgen. nov. (two species of the scolopendrina group as outgroup), estimated by maximum likelihood and neighbour-joining analysis of concatenated sequences of the mitochondrial cytochrome c oxidase subunit I and rDNA 16S (in total 1125 bp). Values at branches indicate bootstrap values for maximum likelihood (above branches) and neighbour-joining (below branches). Specimen codes are resolved in the Supplementary file." figureDoi="http://doi.org/10.5281/zenodo.3827676" httpUri="https://zenodo.org/record/3827676/files/figure.png" pageId="15" pageNumber="16">1</figureCitation>
|
||
). Radial shield triangular with concave proximal edge, internally with distal round condyle (
|
||
<figureCitation id="13402A64453184325A7AFAEFFB36FAFB" box="[1104,1199,1287,1314]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5H</figureCitation>
|
||
), not as ball-like pronounced as in
|
||
<taxonomicName id="4C7B4D62453184325FBFFACAFD84FA9E" authorityName="Peters" authorityYear="1851" box="[405,541,1325,1351]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3453184325FBFFACAFD84FA9E" box="[405,541,1325,1351]" italics="true" pageId="15" pageNumber="16">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D62453184325C76FACAFD59FA9E" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[604,704,1325,1351]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3453184325C76FACAFD59FA9E" box="[604,704,1325,1351]" italics="true" pageId="15" pageNumber="16">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788453184325CE0FACAFCBEFA9E" box="[714,807,1325,1351]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Adradial genital plate (
|
||
<figureCitation id="13402A64453184325A6FFACAFB3EFA9E" box="[1093,1191,1324,1351]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5K</figureCitation>
|
||
<figureCitation id="13402A64453184325A8DFACBFB28FA9E" box="[1191,1201,1324,1351]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[195,1380,1071,1817]" captionTargetId="figure@3.[189,1398,1071,1820]" captionTargetPageId="3" captionText="Fig. 1. Phylogenetic reconstruction of Ophiocoma (Breviturma) subgen. nov. (two species of the scolopendrina group as outgroup), estimated by maximum likelihood and neighbour-joining analysis of concatenated sequences of the mitochondrial cytochrome c oxidase subunit I and rDNA 16S (in total 1125 bp). Values at branches indicate bootstrap values for maximum likelihood (above branches) and neighbour-joining (below branches). Specimen codes are resolved in the Supplementary file." figureDoi="http://doi.org/10.5281/zenodo.3827676" httpUri="https://zenodo.org/record/3827676/files/figure.png" pageId="15" pageNumber="16">1</figureCitation>
|
||
) long, flat, thin, thickened at distal end, with condyle and pit; abradial genital plate (
|
||
<figureCitation id="13402A64453184325DCDFAB5FBDAFAB5" box="[999,1091,1361,1388]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5K</figureCitation>
|
||
<figureCitation id="13402A64453184325A6EFAB6FBCAFAB5" box="[1092,1107,1361,1388]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="15" pageNumber="16">2</figureCitation>
|
||
) thin, blade-like, curved distally. Proximal vertebrae 1.6 mm wide and 1.2 mm high (
|
||
<figureCitation id="13402A64453184325DAAFA90FC47FA48" box="[896,990,1398,1425]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="15" pageNumber="16">Fig. 5N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325EC0FA5DFE2DF9C7" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="8BC436E1453184325EC0FA5DFE4BFA0D" blockId="15.[189,1398,1466,1566]" box="[234,466,1466,1492]" pageId="15" pageNumber="16">
|
||
<emphasis id="B90FEAF3453184325EC0FA5DFE4BFA0D" bold="true" box="[234,466,1466,1492]" pageId="15" pageNumber="16">
|
||
<collectionCode id="ED6AAE24453184325EC0FA5DFEA6FA0D" box="[234,319,1466,1492]" pageId="15" pageNumber="16">URUN</collectionCode>
|
||
2009-11047
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453184325E97FA38FE2DF9C7" blockId="15.[189,1398,1466,1566]" pageId="15" pageNumber="16">
|
||
COLLECTING DATA.
|
||
<collectingCountry id="F36C7671453184325FA4FA38FD82FA20" box="[398,539,1503,1529]" name="Réunion" pageId="15" pageNumber="16">La Réunion</collectingCountry>
|
||
, Etang Salé,
|
||
<geoCoordinate id="EE4F5026453184325C99FA38FCDCFA20" box="[691,837,1503,1529]" degrees="21" direction="south" minutes="16" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="17" value="-21.271389">21°16’17” S</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026453184325D66FA38FC7BFA20" box="[844,994,1503,1530]" degrees="55" direction="east" minutes="19" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="56" value="55.332222">55°19’56” E</geoCoordinate>
|
||
, collectors E. Boissin, T. Hoareau,
|
||
<date id="FFC51021453184325E97F9E3FEC3F9C7" box="[189,346,1540,1566]" pageId="15" pageNumber="16" value="2009-02-24">24 Feb. 2009</date>
|
||
, 1 spm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325E97F9AFFDF6F909" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="8BC436E1453184325E97F9AFFDF6F909" blockId="15.[189,1399,1607,1745]" pageId="15" pageNumber="16">
|
||
MORPHOLOGICAL DATA. 23.7 mm dd, dorsal disc uniformly reddish brown (
|
||
<figureCitation id="13402A64453184325A6AF9A0FB07F9B8" box="[1088,1182,1607,1633]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="15" pageNumber="16">Fig. 2F</figureCitation>
|
||
), dorsal arm dark brown, ventral disc and arms dark brown. Granule density 44-60
|
||
<superScript id="7C0E9BA9453184325D84F98AFC73F9A0" attach="left" box="[942,1002,1643,1671]" fontSize="6" pageId="15" pageNumber="16">mm-2</superScript>
|
||
, granule diameter
|
||
<quantity id="4C839B04453184325AE9F98BFAA1F95E" box="[1219,1336,1644,1671]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="15" pageNumber="16" unit="mm" value="80.0" valueMax="90.0" valueMin="70.0">70-90 µm</quantity>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4 (5), 4 (5), 4, 4, 4, 4, 4, 3, 3, 3, 3 (5 spines on one side of one arm, once at segment 7 and once at segment 8).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325EC0F91DFECCF887" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="8BC436E1453184325EC0F91DFE4DF8CA" blockId="15.[189,1398,1785,1886]" box="[234,468,1785,1812]" pageId="15" pageNumber="16">
|
||
<emphasis id="B90FEAF3453184325EC0F91DFE4DF8CA" bold="true" box="[234,468,1785,1812]" pageId="15" pageNumber="16">
|
||
<collectionCode id="ED6AAE24453184325EC0F91DFEA6F8CD" box="[234,319,1786,1812]" pageId="15" pageNumber="16">URUN</collectionCode>
|
||
2007-00380
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1453184325E97F8F8FECCF887" blockId="15.[189,1398,1785,1886]" pageId="15" pageNumber="16">
|
||
COLLECTING DATA.
|
||
<collectingCountry id="F36C7671453184325FBEF8F8FDBAF8E1" box="[404,547,1822,1849]" name="Réunion" pageId="15" pageNumber="16">La Réunion</collectingCountry>
|
||
, La Saline Trou d’eau,
|
||
<geoCoordinate id="EE4F5026453184325D69F8F9FC4EF8E0" box="[835,983,1822,1849]" degrees="21" direction="south" minutes="06" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="09" value="-21.1025">21°06’09” S</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026453184325DC8F8F9FBE3F8E0" box="[994,1146,1822,1849]" degrees="55" direction="east" minutes="14" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="24" value="55.24">55°14’24” E</geoCoordinate>
|
||
, collector G. Paulay, 2007, 1 spm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A453184325E97F860FB17F832" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="8BC436E1453184325E97F860FB17F832" blockId="15.[189,1399,1927,2027]" pageId="15" pageNumber="16">
|
||
MORPHOLOGICAL DATA. 24.2 mm dd, dorsal disc brown, dorsal arms dark brown with widely spaced light markings, spine slighter brown, ventral disc brown, oral frame lighter, ventral arms brown. Granule density 52-68
|
||
<superScript id="7C0E9BA9453184325F4FF836FE38F804" attach="left" box="[357,417,1999,2027]" fontSize="6" pageId="15" pageNumber="16">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3, 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842D5E97FEECFC11FD27" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8BC436E1452E842D5E97FEECFEB6FEFC" blockId="16.[189,303,267,293]" box="[189,303,267,293]" pageId="16" pageNumber="17">
|
||
<heading id="D08C818D452E842D5E97FEECFEB6FEFC" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B90FEAF3452E842D5E97FEECFEB6FEFC" bold="true" box="[189,303,267,293]" pageId="16" pageNumber="17">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97FEDDFE57FD41" blockId="16.[189,1399,314,664]" pageId="16" pageNumber="17">
|
||
The types of
|
||
<taxonomicName id="4C7B4D62452E842D5F78FEDCFE47FE8D" authorityName="Lyman" authorityYear="1862" box="[338,478,314,341]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="insularia">
|
||
<emphasis id="B90FEAF3452E842D5F78FEDCFE47FE8D" box="[338,478,314,341]" italics="true" pageId="16" pageNumber="17">O. insularia</emphasis>
|
||
</taxonomicName>
|
||
are clearly all conspecific. Their colouration and large size suggest that they are also conspecific with
|
||
<taxonomicName id="4C7B4D62452E842D5F96FEB9FDA1FEA1" authorityName="Muller & Troschel" authorityYear="1842" box="[444,568,350,376]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452E842D5F96FEB9FDA1FEA1" box="[444,568,350,376]" italics="true" pageId="16" pageNumber="17">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C7B4D62452E842D5C62FEB9FCA9FEA1" authorityName="Muller & Troschel" authorityYear="1842" box="[584,816,350,376]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452E842D5C62FEB9FCA9FEA1" box="[584,816,350,376]" italics="true" pageId="16" pageNumber="17">Ophiocoma dentata</emphasis>
|
||
</taxonomicName>
|
||
occurs in several colour morphs, from uniformly dark brown to brown marbled discs with banded arms, and reticulated or spotted forms that resemble
|
||
<taxonomicName id="4C7B4D62452E842D5E97FE41FEF3FE19" authorityName="de Loriol" authorityYear="1899" box="[189,362,422,448]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="doederleini">
|
||
<emphasis id="B90FEAF3452E842D5E97FE41FEF3FE19" box="[189,362,422,448]" italics="true" pageId="16" pageNumber="17">O. doederleini</emphasis>
|
||
</taxonomicName>
|
||
. It can always be distinguished from the latter species by the absence of annulations on the arm spines, and shorter arm spines. In addition, only the largest individuals sometimes have more than four spines on few arm segments. The conspecificity of the various colour morphs is also confirmed by molecular data (Boissin & Hoareau, unpublished). Another species currently believed to be conspecific with
|
||
<taxonomicName id="4C7B4D62452E842D5F84FDD1FDB4FD96" authorityName="Muller & Troschel" authorityYear="1842" box="[430,557,565,592]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452E842D5F84FDD1FDB4FD96" box="[430,557,565,592]" italics="true" pageId="16" pageNumber="17">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="4C7B4D62452E842D5C7EFDD1FD73FD96" authorityName="Smith" authorityYear="1876" box="[596,746,566,592]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="variegata">
|
||
<emphasis id="B90FEAF3452E842D5C7EFDD1FD73FD96" box="[596,746,566,592]" italics="true" pageId="16" pageNumber="17">O. variegata</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<typeStatus id="54C08843452E842D5D19FDD1FCFFFD89" box="[819,870,566,592]" pageId="16" pageNumber="17">type</typeStatus>
|
||
locality Rodriguez Island. According to the original description (
|
||
<bibRefCitation id="EFEA4B10452E842D5F9FFDBEFDDDFDAD" author="Smith E. A." box="[437,580,601,628]" pageId="16" pageNumber="17" pagination="39 - 40" refId="ref15680" refString="Smith E. A. 1876. Descriptions of two new species of Ophiocoma. Annals and Magazine of Natural History, series 4 18: 39 - 40." type="journal article" year="1876">Smith 1876</bibRefCitation>
|
||
), the
|
||
<typeStatus id="54C08843452E842D5CA2FDBDFD22FDAD" box="[648,699,602,628]" pageId="16" pageNumber="17">type</typeStatus>
|
||
measured
|
||
<quantity id="4C839B04452E842D5D17FDBEFC08FDAD" box="[829,913,601,628]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="28.0">28 mm</quantity>
|
||
dd and had only four arm spines, which agrees with
|
||
<taxonomicName id="4C7B4D62452E842D5F61FD9AFE5EFD4E" authorityName="Muller & Troschel" authorityYear="1842" box="[331,455,637,663]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452E842D5F61FD9AFE5EFD4E" box="[331,455,637,663]" italics="true" pageId="16" pageNumber="17">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97FD27FC11FD27" blockId="16.[189,1398,703,766]" pageId="16" pageNumber="17">
|
||
<bibRefCitation id="EFEA4B10452E842D5E97FD27FEE2FD00" author="Devaney D. M." box="[189,379,703,730]" pageId="16" pageNumber="17" refId="ref14989" refString="Devaney D. M. 1970. Studies on ophiocomid brittlestars. I. A new genus (Clarkcoma) of Ophiocominae with a reevaluation of the genus Ophiocoma. Smithsonian Contributions to Zoology 51, Smithsonian Institution Press, Washington. http: // dx. doi. org / 10.5479 / si. 00810282.51" type="book" year="1970">Devaney (1970)</bibRefCitation>
|
||
synonymized
|
||
<taxonomicName id="4C7B4D62452E842D5C0CFD27FD56FD00" authorityName="Marktanner-Turneretscher" authorityYear="1887" box="[550,719,704,730]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="marmorata">
|
||
<emphasis id="B90FEAF3452E842D5C0CFD27FD56FD00" box="[550,719,704,730]" italics="true" pageId="16" pageNumber="17">O. marmorata</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C7B4D62452E842D5D38FD27FC17FD00" authorityName="Muller & Troschel" authorityYear="1842" box="[786,910,703,730]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452E842D5D38FD27FC17FD00" box="[786,910,703,730]" italics="true" pageId="16" pageNumber="17">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
, but as we discuss below, the
|
||
<typeStatus id="54C08843452E842D5ADAFD58FACEFD00" box="[1264,1367,703,729]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7B4D62452E842D5E97FD04FEFCFD24" authorityName="Marktanner-Turneretscher" authorityYear="1887" box="[189,357,739,765]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="marmorata">
|
||
<emphasis id="B90FEAF3452E842D5E97FD04FEFCFD24" box="[189,357,739,765]" italics="true" pageId="16" pageNumber="17">O. marmorata</emphasis>
|
||
</taxonomicName>
|
||
is most likely not conspecific with
|
||
<taxonomicName id="4C7B4D62452E842D5D2FFD04FC18FD24" authorityName="Muller & Troschel" authorityYear="1842" box="[773,897,739,765]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452E842D5D2FFD04FC18FD24" box="[773,897,739,765]" italics="true" pageId="16" pageNumber="17">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |