474 lines
72 KiB
XML
474 lines
72 KiB
XML
<document id="936F50FC916993B660F99603A308452D" ID-CLB-Dataset="22129" ID-DOI="10.5852/ejt.2013.68" ID-GBIF-Dataset="7f2671b2-d26a-4f72-ad1e-5dd3fd67ffc8" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827674" ID-ZooBank="AD45942C-4285-4BAD-9728-EB41EE22F226" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589478025346" checkinUser="carolina" docAuthor="Stöhr, Sabine, Boissin, Emilie & Hoareau, Thierry B." docDate="2013" docId="03D287F7452E842E5C06FCC2FE90F9FA" docLanguage="en" docName="ejt-2013-68.pdf.imf" docOrigin="European Journal of Taxonomy 68" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiocoma (Breviturma) krohi Stöhr & Boissin & Hoareau & Ecomar & Usr 2013, sp. nov." docType="treatment" docUuid="D662D2A3-330D-4693-8757-E6112C4BABC1" docUuidSource="ZooBank" docVersion="8" lastPageNumber="20" masterDocId="FFEBFF8F453E843D5E2AFFE7FF99FFD9" masterDocTitle="Taxonomic revision and phylogeny of the Ophiocoma brevipes group (Echinodermata, Ophiuroidea), with description of a new subgenus (Breviturma) and a new species" masterLastPageNumber="26" masterPageNumber="1" pageNumber="17" updateTime="1698839756557" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="99788766AD84AC749B09573691481674" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="7549634DCB3C273B7AFD2112A4FCE271">
|
||
<mods:title id="BEC12A1AE6BF4AED269DD10581789F24">Taxonomic revision and phylogeny of the Ophiocoma brevipes group (Echinodermata, Ophiuroidea), with description of a new subgenus (Breviturma) and a new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6EF1704D5D23F18CAD23249970948E77" type="personal">
|
||
<mods:role id="AC55F88A79C9A30CC684501BAAE6A616">
|
||
<mods:roleTerm id="D1D03C6B49B983253FFF90144B2527FE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7C6436692A6802D4F5B6BF5FBB25327A">Stöhr, Sabine</mods:namePart>
|
||
<mods:nameIdentifier id="9522F47C2D989824DD88F3BD73ADE13C" type="LSID">412800EB-AACE-4313-9810-61F89B740405</mods:nameIdentifier>
|
||
<mods:affiliation id="B41EF1814522BEF64288BD0215975D8C">Swedish Museum of Natural History, Department of Zoology, Box 50007, 104 05 Stockholm, Sweden Email: sabine. stohr @ nrm. se (corresponding author) & urn: lsid: zoobank. org: author: 412800 EB-AACE- 4313 - 9810 - 61 F 89 B 740405</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="487731B4EABF4E0013C5454E780347BC" type="personal">
|
||
<mods:role id="A8BDA4A5A13BBB31144C3AE14487384B">
|
||
<mods:roleTerm id="8E335D8B3D994327CD861E771FA0B37A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="84CAC8D3600C077660694B5228DBCE32">Boissin, Emilie</mods:namePart>
|
||
<mods:nameIdentifier id="88CE244FE5E01FE1B519D041D373EF7F" type="LSID">B35FF544-AE44-43B2-B1CA-3769A281F386</mods:nameIdentifier>
|
||
<mods:affiliation id="6296256F66A54ACB13E9FB5BB5568DFC">Laboratoire ECOMAR, Université de La Réunion, BP 7151, 97715 Saint-Denis, La Réunion, France & USR 3278 - CRIOBE-CNRS-EPHE, Laboratoire d’Excellence « CORAIL », Université de Perpignan-CBETM, 58 rue Paul Alduy, 66860 Perpignan Cedex & Email: eboissin @ gmail. com & urn: lsid: zoobank. org: author: B 35 FF 544 - AE 44 - 43 B 2 - B 1 CA- 3769 A 281 F 386</mods:affiliation>
|
||
<mods:nameIdentifier id="4FC5E7530045617BE5ED4C4CEFD48502" type="email">eboissin@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="30DFF0ADCECAF4FC36755B424270A164" type="personal">
|
||
<mods:role id="C13C5FC66DFE80B4922F24FDAD855211">
|
||
<mods:roleTerm id="56B01113CC4070020BC91686C7C5FCEE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4558E32D6718CC6B815025838EC554B5">Hoareau, Thierry B.</mods:namePart>
|
||
<mods:nameIdentifier id="4B771128C375CA3AB89D3C3F880F30B2" type="LSID">44578720-231E-4D99-8F38-2A94B7C4E4F2</mods:nameIdentifier>
|
||
<mods:affiliation id="584BFA8494A691948C9142CB7D7BD3A4">Laboratoire ECOMAR, Université de La Réunion, BP 7151, 97715 Saint-Denis, La Réunion, France & Molecular Ecology and Evolution Programme, Department of Genetics, University of Pretoria, Private bag X 20, Hatfield, Pretoria 0028, South Africa & Email: thoareau @ gmail. com & urn: lsid: zoobank. org: author: 44578720 - 231 E- 4 D 99 - 8 F 38 - 2 A 94 B 7 C 4 E 4 F 2</mods:affiliation>
|
||
<mods:nameIdentifier id="984696478E6F925C2FFE18DDFCBE432C" type="email">thoareau@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F0143CFA4FC469747E6CE64D07764D73">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F7526B8C2D1EDE336E1F424D0E201F11" type="host">
|
||
<mods:titleInfo id="12F29F5CE268C719FA09D3CA65BDD747">
|
||
<mods:title id="869C71F25DF6B51C459B3785E5195D22">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="3390A91EE70D1D1C4F922AE13BF4D413">
|
||
<mods:date id="42D9068CDBE4574C442F346BE379730E">2013</mods:date>
|
||
<mods:detail id="27DD4007243034A9628B45923CD72D52" type="pubDate">
|
||
<mods:number id="3F85AFE83E961AD173E21B2C8FD9904C">2013-12-09</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CFA0FB8680A08EC2071374A120C9764D" type="issue">
|
||
<mods:number id="512E238FBA10D761FDA693EF8216B202">68</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BFBA30A6EDA3E8CF7B1EDAD99B72A2DD" unit="page">
|
||
<mods:start id="1EF2F4B13AA519E7F26AC3ED9D60B6E9">1</mods:start>
|
||
<mods:end id="871E27AB63F03D6FE0A7E83D3D7A3AB4">26</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D0317CEF9A2E98D5BFA43CF024D47C3F">journal article</mods:classification>
|
||
<mods:identifier id="62802C495078DA89DDC836116F4E322F" type="CLB-Dataset">22129</mods:identifier>
|
||
<mods:identifier id="CB75D049EBAAA23C1449063277B44CAD" type="DOI">10.5852/ejt.2013.68</mods:identifier>
|
||
<mods:identifier id="73B269900E0D65A9C7DAC4D2F200D1A2" type="GBIF-Dataset">7f2671b2-d26a-4f72-ad1e-5dd3fd67ffc8</mods:identifier>
|
||
<mods:identifier id="3B8BCD371A257CA53B40DFA3822A2A45" type="ISSN">2118-9773</mods:identifier>
|
||
<mods:identifier id="D1BCC26B48D0E79AFE75DD0B933B5432" type="Zenodo-Dep">3827674</mods:identifier>
|
||
<mods:identifier id="4FD199DF57F7F3B1304120932FACBC72" type="ZooBank">AD45942C-4285-4BAD-9728-EB41EE22F226</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D287F7452E842E5C06FCC2FE90F9FA" ID-DOI="http://doi.org/10.5281/zenodo.3846763" ID-GBIF-Taxon="164152681" ID-Zenodo-Dep="3846763" LSID="urn:lsid:zoobank.org:act:D662D2A3-330D-4693-8757-E6112C4BABC1" httpUri="http://treatment.plazi.org/id/03D287F7452E842E5C06FCC2FE90F9FA" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
|
||
<subSubSection id="C361656A452E842D5C06FCC2FB91FC99" box="[556,1032,805,832]" pageId="16" pageNumber="17" type="nomenclature">
|
||
<paragraph id="8BC436E1452E842D5C06FCC2FB91FC99" blockId="16.[366,1222,805,904]" box="[556,1032,805,832]" pageId="16" pageNumber="17">
|
||
<heading id="D08C818D452E842D5C06FCC2FB91FC99" box="[556,1032,805,832]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
|
||
<taxonomicName id="4C7B4D62452E842D5C06FCC2FC3FFC99" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[556,934,805,832]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov." subGenus="Breviturma">
|
||
<emphasis id="B90FEAF3452E842D5C06FCC2FC3FFC99" bold="true" box="[556,934,805,832]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90FEAF3452E842D5C06FCC2FD21FC99" bold="true" box="[556,696,805,832]" italics="true" pageId="16" pageNumber="17">Ophiocoma</emphasis>
|
||
(
|
||
<emphasis id="B90FEAF3452E842D5CE3FCC2FCCDFCE6" bold="true" box="[713,852,805,831]" italics="true" pageId="16" pageNumber="17">Breviturma</emphasis>
|
||
)
|
||
<emphasis id="B90FEAF3452E842D5D4FFCC2FC3FFC99" bold="true" box="[869,934,805,832]" italics="true" pageId="16" pageNumber="17">krohi</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788452E842D5D87FCC1FB91FC99" box="[941,1032,806,832]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842D5F44FCAEFC6DFC5E" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BC436E1452E842D5F44FCAEFB5FFCBD" blockId="16.[366,1222,805,904]" box="[366,1222,841,868]" pageId="16" pageNumber="17">
|
||
<uri id="FFEA3AE3452E842D5F44FCAEFB5FFCBD" box="[366,1222,841,868]" pageId="16" pageNumber="17">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFDD0C34452E842D5CA7FCAEFB5FFCBD" box="[653,1222,841,868]" pageId="16" pageNumber="17">D662D2A3-330D-4693-8757-E6112C4BABC1</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5C15FC8AFC6DFC5E" blockId="16.[366,1222,805,904]" box="[575,1012,877,904]" pageId="16" pageNumber="17">
|
||
<figureCitation id="13402A64452E842D5C15FC8AFD1EFC5E" box="[575,647,877,903]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="16" pageNumber="17">Figs 2</figureCitation>
|
||
G-L, 4K-O, 5C, F, I, L, O, R, 6
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842D5E97FC48FD48FC34" pageId="16" pageNumber="17" type="reference_group">
|
||
<paragraph id="8BC436E1452E842D5E97FC48FD4AFC13" blockId="16.[189,723,943,1005]" box="[189,723,943,970]" pageId="16" pageNumber="17">
|
||
<treatmentCitationGroup id="AB6B11CF452E842D5E97FC48FD4AFC13" box="[189,723,943,970]" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C7B4D62452E842D5E97FC48FE36FC10" authorityName="Peters" authorityYear="1851" box="[189,431,943,969]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3452E842D5E97FC48FE36FC10" box="[189,431,943,969]" italics="true" pageId="16" pageNumber="17">Ophiocoma brevipes</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADA10F0452E842D5FE7FC48FD49FC13" author="Stohr S. & Conand C. & Boissin E." box="[461,720,943,970]" page="553" pageId="16" pageNumber="17" year="2008">
|
||
<bibRefCitation id="EFEA4B10452E842D5FE7FC48FD49FC13" author="Stohr S. & Conand C. & Boissin E." box="[461,720,943,970]" pageId="16" pageNumber="17" pagination="545 - 560" refId="ref15768" refString="Stohr S., Conand C. & Boissin E. 2008. Brittle stars (Echinodermata: Ophiuroidea) from La Reunion and the systematic position of Ophiocanops Koehler, 1922. Zoological Journal of the Linnean Society 153 (3): 545 - 560. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2008.00401. x" type="journal article" year="2008">
|
||
Stöhr
|
||
<emphasis id="B90FEAF3452E842D5C3EFC57FDD7FC10" box="[532,590,943,969]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2008: 553
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97FC34FD48FC34" blockId="16.[189,723,943,1005]" box="[189,721,979,1005]" pageId="16" pageNumber="17">
|
||
<treatmentCitationGroup id="AB6B11CF452E842D5E97FC34FD48FC34" box="[189,721,979,1005]" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C7B4D62452E842D5E97FC34FEDFFC34" authorityName="L. Agassiz" authorityYear="1835" box="[189,326,979,1005]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B90FEAF3452E842D5E97FC34FEDFFC34" box="[189,326,979,1005]" italics="true" pageId="16" pageNumber="17">Ophiocoma</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. –
|
||
<treatmentCitation id="0ADA10F0452E842D5FEFFC34FD57FC34" author="Hoareau T. B. & Boissin E. & Paulay G. & Bruggemann J. H." box="[453,718,979,1005]" page="2" pageId="16" pageNumber="17" year="2013">
|
||
<bibRefCitation id="EFEA4B10452E842D5FEFFC34FD57FC34" author="Hoareau T. B. & Boissin E. & Paulay G. & Bruggemann J. H." box="[453,718,979,1005]" pageId="16" pageNumber="17" pagination="2167 - 2197" refId="ref15158" refString="Hoareau T. B., Boissin E., Paulay G. & Bruggemann J. H. 2013. The Southwestern Indian Ocean as a potential marine evolutionary hotspot: perspectives from comparative phylogeography of reef brittlestars. Journal of Biogeography 40 (11): 2167 - 2197. http: // dx. doi. org / 10.1111 / jbi. 12155" type="journal article" year="2013">
|
||
Hoareau
|
||
<emphasis id="B90FEAF3452E842D5C05FC33FDF3FC34" box="[559,618,979,1005]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2013: 2
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842D5E97FBF2FC57FB13" pageId="16" pageNumber="17" type="diagnosis">
|
||
<paragraph id="8BC436E1452E842D5E97FBF2FEAFFBF6" blockId="16.[189,310,1045,1071]" box="[189,310,1045,1071]" pageId="16" pageNumber="17">
|
||
<heading id="D08C818D452E842D5E97FBF2FEAFFBF6" bold="true" box="[189,310,1045,1071]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B90FEAF3452E842D5E97FBF2FEAFFBF6" bold="true" box="[189,310,1045,1071]" pageId="16" pageNumber="17">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97FBA3FC57FB13" blockId="16.[189,1399,1092,1226]" pageId="16" pageNumber="17">
|
||
Species of
|
||
<taxonomicName id="4C7B4D62452E842D5F16FBA3FDFAFB86" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[316,611,1092,1119]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="subGenus" status="subgen. nov." subGenus="Breviturma">
|
||
<emphasis id="B90FEAF3452E842D5F16FBA3FE5CFB87" box="[316,453,1092,1118]" italics="true" pageId="16" pageNumber="17">Ophiocoma</emphasis>
|
||
(
|
||
<emphasis id="B90FEAF3452E842D5FFFFBA2FDC3FB87" box="[469,602,1093,1118]" italics="true" pageId="16" pageNumber="17">Breviturma</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788452E842D5C40FBA3FD65FB86" box="[618,764,1092,1119]" pageId="16" pageNumber="17" rank="subGenus">subgen. nov.</taxonomicNameLabel>
|
||
with variegated to uniform dark brown colour pattern; arm spines may be weakly annulated, but not as clearly light and dark banded as in
|
||
<taxonomicName id="4C7B4D62452E842D5AAEFB8EFAB4FB5B" authorityName="de Loriol" authorityYear="1899" box="[1156,1325,1128,1155]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="doederleini">
|
||
<emphasis id="B90FEAF3452E842D5AAEFB8EFAB4FB5B" box="[1156,1325,1128,1155]" italics="true" pageId="16" pageNumber="17">O. doederleini</emphasis>
|
||
</taxonomicName>
|
||
; up to four arm spines (in large specimens beyond segment 15), sometimes five at segments 6-9 (rarely 5-8); relatively coarse disc granulation. Maximum size about
|
||
<quantity id="4C839B04452E842D5D65FB57FC3BFB13" box="[847,930,1200,1226]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="17" unit="mm" value="16.0">16 mm</quantity>
|
||
dd.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842D5E97FB15FE47FA86" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="8BC436E1452E842D5E97FB15FEDDFAD5" blockId="16.[189,324,1266,1292]" box="[189,324,1266,1292]" pageId="16" pageNumber="17">
|
||
<heading id="D08C818D452E842D5E97FB15FEDDFAD5" bold="true" box="[189,324,1266,1292]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B90FEAF3452E842D5E97FB15FEDDFAD5" bold="true" box="[189,324,1266,1292]" pageId="16" pageNumber="17">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97FAC6FE47FA86" blockId="16.[189,1399,1313,1375]" pageId="16" pageNumber="17">
|
||
The species is named in honour of Dr. Andreas Kroh,
|
||
<collectingRegion id="49BFF803452E842D5D5BFAC6FC5EFAE2" box="[881,967,1313,1339]" country="Austria" name="Wien" pageId="16" pageNumber="17">Vienna</collectingRegion>
|
||
, for his important contributions to echinoderm systematics.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842D5E97FA60FE05F8EB" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="8BC436E1452E842D5E97FA60FEF5FA78" blockId="16.[189,364,1415,1441]" box="[189,364,1415,1441]" pageId="16" pageNumber="17">
|
||
<heading id="D08C818D452E842D5E97FA60FEF5FA78" bold="true" box="[189,364,1415,1441]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B90FEAF3452E842D5E97FA60FEF5FA78" bold="true" box="[189,364,1415,1441]" pageId="16" pageNumber="17">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B133CBC452E842D5EC0FA50FD81F9C1" ID-GBIF-Occurrence="2619830304" collectingDate="2009-03-25" collectorName="E. Boissin" country="La Reunion Island" latitude="-21.340834" location="La Reunion Island" longLatPrecision="21" longitude="55.45861" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BC436E1452E842D5EC0FA50FEC5FA08" blockId="16.[189,1398,1463,1560]" box="[234,348,1463,1489]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90FEAF3452E842D5EC0FA50FEC5FA08" bold="true" box="[234,348,1463,1489]" pageId="16" pageNumber="17">
|
||
<typeStatus id="54C08843452E842D5EC0FA50FEC5FA08" box="[234,348,1463,1489]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97FA3CFD81F9C1" blockId="16.[189,1398,1463,1560]" pageId="16" pageNumber="17">
|
||
<collectingCountry id="F36C7671452E842D5E97FA3CFE00FA2D" box="[189,409,1498,1525]" name="La Reunion Island" pageId="16" pageNumber="17">La Réunion Island</collectingCountry>
|
||
, St. Pierre,
|
||
<geoCoordinate id="EE4F5026452E842D5C0AFA3DFD3CFA2D" box="[544,677,1498,1524]" degrees="21" direction="south" minutes="20" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="27" value="-21.340834">21°20’27 S</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026452E842D5C86FA3DFCACFA2C" box="[684,821,1498,1525]" degrees="55" direction="east" minutes="27" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="31" value="55.45861">55°27’31 E</geoCoordinate>
|
||
, collectors
|
||
<collectorName id="268E5337452E842D5D91FA3CFBAFFA2D" box="[955,1078,1498,1525]" pageId="16" pageNumber="17">E. Boissin</collectorName>
|
||
, T. Hoareau,
|
||
<date id="FFC51021452E842D5AF8FA3DFAEBFA2D" box="[1234,1394,1498,1525]" pageId="16" pageNumber="17" value="2009-03-25">
|
||
<collectingDate id="EF81E9C9452E842D5AF8FA3DFAEBFA2D" box="[1234,1394,1498,1525]" pageId="16" pageNumber="17" value="2009-03-25">25 Mar. 2009</collectingDate>
|
||
</date>
|
||
, in ethanol (MNHN-IE-4300).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BC436E1452E842D5EC0F9A6FEFEF982" blockId="16.[189,1398,1601,1842]" box="[234,359,1601,1627]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90FEAF3452E842D5EC0F9A6FEFEF982" bold="true" box="[234,359,1601,1627]" pageId="16" pageNumber="17">
|
||
<typeStatus id="54C08843452E842D5EC0F9A6FEFEF982" box="[234,359,1601,1627]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5E97F982FE05F8EB" blockId="16.[189,1398,1601,1842]" pageId="16" pageNumber="17">
|
||
From same locality as
|
||
<typeStatus id="54C08843452E842D5FEDF983FDB7F9A7" box="[455,558,1636,1662]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
8 spms: 1 spm, dissected, fragments on SEM stubs and arms in ethanol (SMNH-Type-8535); 1 spm gold coated and uncoated arms dry (SMNH-Type-8534); 2 spms in ethanol (SMNH-Type-8531, 8536); 1 spm in ethanol (
|
||
<collectionCode id="ED6AAE24452E842D5CFBF94BFD63F91F" box="[721,762,1708,1734]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="17" type="Museum">UF</collectionCode>
|
||
13938); 1 spm in ethanol (
|
||
<collectionCode id="ED6AAE24452E842D5A0DF94BFBE5F91F" box="[1063,1148,1708,1734]" pageId="16" pageNumber="17">URUN</collectionCode>
|
||
2009-11108); 2 spms in ethanol (MNHN-IE-4301, 4302).
|
||
<collectingCountry id="F36C7671452E842D5C59F937FC9AF933" box="[627,771,1744,1770]" name="Réunion" pageId="16" pageNumber="17">La Réunion</collectingCountry>
|
||
, Cap Homard,
|
||
<geoCoordinate id="EE4F5026452E842D5D97F937FBCBF933" box="[957,1106,1744,1770]" degrees="21" direction="south" minutes="02" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="07" value="-21.035276">21°02’07” S</geoCoordinate>
|
||
<geoCoordinate id="EE4F5026452E842D5A76F937FB6CF933" box="[1116,1269,1744,1771]" degrees="55" direction="east" minutes="13" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="06" value="55.218334">55°13’06” E</geoCoordinate>
|
||
, collectors E. Boissin, T. Hoareau,
|
||
<date id="FFC51021452E842D5FF0F913FDE2F8D7" box="[474,635,1780,1806]" pageId="16" pageNumber="17" value="2009-02-23">23 Feb. 2009</date>
|
||
: 2 spms in ethanol (SMNH-Type-8532, 8533); 1 spm in ethanol (MNHN-IE-4303).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452E842E5E97F8BEFAAFFB96" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BC436E1452E842D5E97F8BEFED7F8AA" blockId="16.[189,334,1881,1907]" box="[189,334,1881,1907]" pageId="16" pageNumber="17">
|
||
<heading id="D08C818D452E842D5E97F8BEFED7F8AA" bold="true" box="[189,334,1881,1907]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B90FEAF3452E842D5E97F8BEFED7F8AA" bold="true" box="[189,334,1881,1907]" pageId="16" pageNumber="17">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842D5EC0F86EFEC5F87A" blockId="16.[189,1399,1929,2027]" box="[234,348,1929,1955]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90FEAF3452E842D5EC0F86EFEC5F87A" bold="true" box="[234,348,1929,1955]" pageId="16" pageNumber="17">
|
||
<typeStatus id="54C08843452E842D5EC0F86EFEC5F87A" box="[234,348,1929,1955]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452E842C5E97F84AFAC1FC81" blockId="16.[189,1399,1929,2027]" lastBlockId="17.[189,1399,267,857]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
12.7 mm dd, radial shields completely obscured by granules, granule density 104
|
||
<superScript id="7C0E9BA9452E842D5A81F84AFB7FF860" attach="left" box="[1195,1254,1963,1991]" fontSize="6" pageId="16" pageNumber="17">mm-2</superScript>
|
||
. Arm spine sequence 3(2), 3(2), 3, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3. Dorsal disc with dark and lighter brown lines radiating from a dark central patch, some white spots at disc edge (
|
||
<figureCitation id="13402A64452F842C5DD4FEECFBF8FEFF" box="[1022,1121,267,294]" captionStart="Fig" captionStartId="17.[189,232,1965,1991]" captionTargetBox="[189,1399,1317,1924]" captionTargetId="figure@17.[189,1399,1317,1924]" captionTargetPageId="17" captionText="Fig. 6. Holotype of Ophiocoma (Breviturma subgen. nov.) krohi sp. nov. (MNHN-IE-4300). A. Dorsal disc. B. Dorsal arm. C. Ventral arm, note the pattern on the arm spines. D. Ventral disc. E. Jaws." figureDoi="http://doi.org/10.5281/zenodo.3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18">Fig. 6A</figureCitation>
|
||
). Dorsal arms medium brown with dark and white transverse bands (
|
||
<figureCitation id="13402A64452F842C5CF4FEC9FCA5FE90" box="[734,828,302,329]" captionStart="Fig" captionStartId="17.[189,232,1965,1991]" captionTargetBox="[189,1399,1317,1924]" captionTargetId="figure@17.[189,1399,1317,1924]" captionTargetPageId="17" captionText="Fig. 6. Holotype of Ophiocoma (Breviturma subgen. nov.) krohi sp. nov. (MNHN-IE-4300). A. Dorsal disc. B. Dorsal arm. C. Ventral arm, note the pattern on the arm spines. D. Ventral disc. E. Jaws." figureDoi="http://doi.org/10.5281/zenodo.3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18">Fig. 6B</figureCitation>
|
||
). Ventral disc dark brown mottled, ventral arms and oral frame lighter brown, oral shields dark. Ventral interradii completely, but sparsely granulated (
|
||
<figureCitation id="13402A64452F842C5EECFE92FED1FE56" box="[198,328,373,399]" captionStart="Fig" captionStartId="17.[189,232,1965,1991]" captionTargetBox="[189,1399,1317,1924]" captionTargetId="figure@17.[189,1399,1317,1924]" captionTargetPageId="17" captionText="Fig. 6. Holotype of Ophiocoma (Breviturma subgen. nov.) krohi sp. nov. (MNHN-IE-4300). A. Dorsal disc. B. Dorsal arm. C. Ventral arm, note the pattern on the arm spines. D. Ventral disc. E. Jaws." figureDoi="http://doi.org/10.5281/zenodo.3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18">Fig. 6D, E</figureCitation>
|
||
). Dorsal spines brown, ventral spines weakly annulated brown on white. DAPs oval, W:L 1.8:1, contiguous. Dorsalmost spine a little longer than a segment, middle spines about 2 segments long, ventralmost spine slightly shorter. Arm spines flattened, slightly tapered with truncated tips, dorsalmost spine wider than other spines. Oral shields trapezoid, distally almost as wide as long, proximally just over half as wide as long, lateral edges straight (except madreporite), proximal and distal edges convex (
|
||
<figureCitation id="13402A64452F842C5EECFDC2FEBFFD99" box="[198,294,549,576]" captionStart="Fig" captionStartId="17.[189,232,1965,1991]" captionTargetBox="[189,1399,1317,1924]" captionTargetId="figure@17.[189,1399,1317,1924]" captionTargetPageId="17" captionText="Fig. 6. Holotype of Ophiocoma (Breviturma subgen. nov.) krohi sp. nov. (MNHN-IE-4300). A. Dorsal disc. B. Dorsal arm. C. Ventral arm, note the pattern on the arm spines. D. Ventral disc. E. Jaws." figureDoi="http://doi.org/10.5281/zenodo.3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18">Fig. 6D</figureCitation>
|
||
). Madreporite larger than other oral shields, more rounded, all edges convex. At tip of jaw a cluster of 8-10 small, round tooth papillae. Along each jaw edge 4 rectangular, wider than high, oral papillae, and distally a smaller papilla at an angle to the others (
|
||
<figureCitation id="13402A64452F842C5DA4FD8CFC7EFD5F" box="[910,999,619,646]" captionStart="Fig" captionStartId="17.[189,232,1965,1991]" captionTargetBox="[189,1399,1317,1924]" captionTargetId="figure@17.[189,1399,1317,1924]" captionTargetPageId="17" captionText="Fig. 6. Holotype of Ophiocoma (Breviturma subgen. nov.) krohi sp. nov. (MNHN-IE-4300). A. Dorsal disc. B. Dorsal arm. C. Ventral arm, note the pattern on the arm spines. D. Ventral disc. E. Jaws." figureDoi="http://doi.org/10.5281/zenodo.3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18">Fig. 6E</figureCitation>
|
||
).Adoral shields uneven triangular, framing the lateral edges of the oral shield, not meeting proximally. Bursal slits extend from oral shield to disc edge. First ventral arm plate outside mouth-slit, small, winglike pentagonal, middle of distal edge notched. Following plates pentagonal, about as long as greatest width, with obtuse proximal angle, lateral edges notched, distal edge convex, contiguous (
|
||
<figureCitation id="13402A64452F842C5D16FD1FFC00FCCA" box="[828,921,760,787]" captionStart="Fig" captionStartId="17.[189,232,1965,1991]" captionTargetBox="[189,1399,1317,1924]" captionTargetId="figure@17.[189,1399,1317,1924]" captionTargetPageId="17" captionText="Fig. 6. Holotype of Ophiocoma (Breviturma subgen. nov.) krohi sp. nov. (MNHN-IE-4300). A. Dorsal disc. B. Dorsal arm. C. Ventral arm, note the pattern on the arm spines. D. Ventral disc. E. Jaws." figureDoi="http://doi.org/10.5281/zenodo.3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18">Fig. 6C</figureCitation>
|
||
). 2 oval scales at a slightly open angle at the lateral arm plate of each tentacle pore proximally; the inner scale decreases gradually and is absent on the distalmost segments. Tube feet expanded, smooth, with distal constrictions and terminal bulb.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452F842C5E97FC67FBC4FC43" blockId="17.[189,1117,896,922]" box="[189,1117,896,922]" pageId="17" pageNumber="18">
|
||
The
|
||
<typeStatus id="54C08843452F842C5ED8FC67FEFBFC43" box="[242,354,896,922]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="54C08843452F842C5C1DFC67FD07FC43" box="[567,670,896,922]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
in most characters, except as follows:
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452F842C5EC0FC26FE57FC02" blockId="17.[189,1399,961,1199]" box="[234,462,961,987]" pageId="17" pageNumber="18">
|
||
<emphasis id="B90FEAF3452F842C5EC0FC26FE57FC02" bold="true" box="[234,462,961,987]" pageId="17" pageNumber="18">SMNH-Type-8534</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452F842C5E97FC02FABCFB76" blockId="17.[189,1399,961,1199]" pageId="17" pageNumber="18">
|
||
1 spm,
|
||
<quantity id="4C839B04452F842C5F3EFC02FEFFFC26" box="[276,358,997,1023]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="mm" value="11.0">11 mm</quantity>
|
||
dd, dark brown radiating stripes on dorsal disc, interradii mottled light and dark brown, overall small white dots; dorsal arms dark brown with some lighter markings, spines white with diffuse brown pattern, some with annulations (not figured). Ventral interradii dark brown mottled with white, fully granulated; adoral shields and oral plates unevenly brown, oral shields dark brown with white edges, oral papillae white. Ventral arm plates and tentacle scales brown with white edges. Arm spine sequence 3, 3, 3, 4, 4, 5, 5, 4, 4, 4, 4, 4, 4, 4, 4, 4. Granule density 132-136
|
||
<superScript id="7C0E9BA9452F842C5A1AFB72FBF5FB78" attach="left" box="[1072,1132,1171,1199]" fontSize="6" pageId="17" pageNumber="18">mm-2</superScript>
|
||
on light image.
|
||
</paragraph>
|
||
<caption id="DF046669452F842C5E97F84AFABCF832" ID-DOI="http://doi.org/10.5281/zenodo.3827686" ID-Zenodo-Dep="3827686" httpUri="https://zenodo.org/record/3827686/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1965,1991]" targetBox="[189,1399,1317,1924]" targetPageId="17">
|
||
<paragraph id="8BC436E1452F842C5E97F84AFABCF832" blockId="17.[189,1398,1965,2027]" pageId="17" pageNumber="18">
|
||
<emphasis id="B90FEAF3452F842C5E97F84AFE92F81E" bold="true" box="[189,267,1965,1991]" pageId="17" pageNumber="18">Fig. 6.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C7B4D62452F842C5F83F84AFDABF81E" authorityName="L. Agassiz" authorityYear="1835" box="[425,562,1965,1991]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B90FEAF3452F842C5F83F84AFDABF81E" box="[425,562,1965,1991]" italics="true" pageId="17" pageNumber="18">Ophiocoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C7B4D62452F842C5C10F849FC2AF81E" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[570,947,1965,1992]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov." subGenus="Breviturma">
|
||
(
|
||
<emphasis id="B90FEAF3452F842C5C69F849FD51F81E" box="[579,712,1966,1991]" italics="true" pageId="17" pageNumber="18">Breviturma</emphasis>
|
||
subgen. nov.)
|
||
<emphasis id="B90FEAF3452F842C5D5CF84AFC2AF81E" box="[886,947,1965,1991]" italics="true" pageId="17" pageNumber="18">krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788452F842C5D90F849FB8FF811" box="[954,1046,1966,1992]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(MNHN-IE-4300).
|
||
<emphasis id="B90FEAF3452F842C5B2EF84AFA80F81E" bold="true" box="[1284,1305,1965,1991]" pageId="17" pageNumber="18">A</emphasis>
|
||
. Dorsal disc.
|
||
<emphasis id="B90FEAF3452F842C5ED1F836FE96F832" bold="true" box="[251,271,2001,2027]" pageId="17" pageNumber="18">B</emphasis>
|
||
. Dorsal arm.
|
||
<emphasis id="B90FEAF3452F842C5F85F837FE5DF833" bold="true" box="[431,452,2000,2026]" pageId="17" pageNumber="18">C</emphasis>
|
||
. Ventral arm, note the pattern on the arm spines.
|
||
<emphasis id="B90FEAF3452F842C5A2FF836FB83F832" bold="true" box="[1029,1050,2001,2027]" pageId="17" pageNumber="18">D</emphasis>
|
||
. Ventral disc.
|
||
<emphasis id="B90FEAF3452F842C5AE8F836FB4FF832" bold="true" box="[1218,1238,2001,2027]" pageId="17" pageNumber="18">E</emphasis>
|
||
. Jaws.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC436E1452C842F5E97FEECFD96FE4C" blockId="18.[189,1399,266,405]" pageId="18" pageNumber="19">
|
||
Dorsal disc granule diameter
|
||
<quantity id="4C839B04452C842F5C37FEECFD0CFEFF" box="[541,661,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.3" metricValueMin="5.7" pageId="18" pageNumber="19" unit="mm" value="65.0" valueMax="73.0" valueMin="57.0">57-73 µm</quantity>
|
||
, granule density 132-136
|
||
<superScript id="7C0E9BA9452C842F5DE6FEEBFB91FEC1" attach="left" box="[972,1032,266,294]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
, on SEM image (
|
||
<figureCitation id="13402A64452C842F5AF4FEECFAF1FEFF" box="[1246,1384,267,294]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="18" pageNumber="19">Fig. 4K, M</figureCitation>
|
||
); third arm spine 1.3 mm long, DAP 1.2 mm wide (
|
||
<figureCitation id="13402A64452C842F5D33FED7FCEEFE92" box="[793,887,304,331]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="18" pageNumber="19">Fig. 4L</figureCitation>
|
||
). Oral shields as wide as long (
|
||
<figureCitation id="13402A64452C842F5ADBFED7FACBFE92" box="[1265,1362,304,331]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="18" pageNumber="19">Fig. 4N</figureCitation>
|
||
). 5 tooth papillae, 5 lateral oral papillae at each jaw edge, outer papilla widest. Few granules on proximal ends of oral plates (
|
||
<figureCitation id="13402A64452C842F5F89FE9CFD9BFE4C" box="[419,514,379,405]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[189,1399,380,1783]" captionTargetId="figure@11.[189,1399,380,1783]" captionTargetPageId="11" captionText="Fig. 4. SEM images of three species of Ophiocoma (Breviturma) subgen. nov. A-E. Ophiocoma brevipes Peters, 1851 (SMNH-133230). A. Dorsal disc. B. Dorsal arms. C. Dorsal disc granules. D. Jaws. E. Ventral arm. F-I. Ophiocoma dentata Müller & Troschel, 1842 (SMNH-133232). F. Dorsal disc. G. Dorsal arm. H. Dorsal disc granules. I. Jaws. J. Ventral arm. K-N. Ophiocoma krohi sp. nov. paratype (SMNH-Type-8534). K. Dorsal disc. L. Dorsal arms. M. Dorsal disc granules. N. Jaws. O. Ventral arm. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827682" httpUri="https://zenodo.org/record/3827682/files/figure.png" pageId="18" pageNumber="19">Fig. 4N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5EC0FE59FE57FE01" blockId="18.[189,1399,446,695]" box="[234,462,446,472]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEAF3452C842F5EC0FE59FE57FE01" bold="true" box="[234,462,446,472]" pageId="18" pageNumber="19">SMNH-Type-8535</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97FE04FB4EFD6E" blockId="18.[189,1399,446,695]" pageId="18" pageNumber="19">
|
||
1 spm,
|
||
<quantity id="4C839B04452C842F5F3FFE04FEF0FE27" box="[277,361,483,510]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="mm" value="13.0">13 mm</quantity>
|
||
dd, dorsal disc dark brown, white spots on radial shields; dorsal arms banded dark and light brown; spines dark brown, on distal arm annulated (not figured). Granule diameter
|
||
<quantity id="4C839B04452C842F5AD2FDEFFAE8FDFA" box="[1272,1393,520,547]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" metricValueMax="7.4" metricValueMin="5.7" pageId="18" pageNumber="19" unit="mm" value="65.5" valueMax="74.0" valueMin="57.0">57-74 µm</quantity>
|
||
, density 104
|
||
<superScript id="7C0E9BA9452C842F5F61FDC9FE1FFDE3" attach="left" box="[331,390,556,584]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
. Ventral interradii dark brown; adoral shields light brown with dark spots; oral shields dark brown with white edges; oral papillae white. Ventral arm plates brown with white edges and lighter areas. Arm spine sequence 3, 3, 3, 4, 4, 5, 5, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4. Up to 5 tooth papillae, 5 lateral oral papillae at each jaw edge, outer papilla widest. Few granules on distal ends of oral plates.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97FD07FEA8FC26" blockId="18.[189,1399,736,1023]" pageId="18" pageNumber="19">
|
||
Oral plates, adradial distal part with strong folds, abradial face with large muscle flange with horizontal striations that end in deep incisions in the distal edge; 1.9 mm wide, 2.3 mm high (
|
||
<figureCitation id="13402A64452C842F5A9BFCE2FA88FCF9" box="[1201,1297,773,800]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="18" pageNumber="19">Fig. 5C</figureCitation>
|
||
). Dental plate short, with two tooth foramina on external dorsal half, dorsalmost one oval, wider than long, other small, round; internally a wide septum divides each foramen into two pairs of vertical oval holes, upper ones larger than lower ones (
|
||
<figureCitation id="13402A64452C842F5C71FC93FD2CFC56" box="[603,693,884,911]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="18" pageNumber="19">Fig. 5F</figureCitation>
|
||
). Radial shield angular, with large round condyle at internal distal end (
|
||
<figureCitation id="13402A64452C842F5F14FC7DFE09FC6D" box="[318,400,921,948]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="18" pageNumber="19">Fig. 5I</figureCitation>
|
||
). Adradial genital plate long, flat, thin, thickened at distal end, with deep pit; abradial genital plate thin, blade-like, curved distally (
|
||
<figureCitation id="13402A64452C842F5CE0FC58FC84FC03" box="[714,797,959,986]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="18" pageNumber="19">Fig 5L</figureCitation>
|
||
). Proximal vertebrae 2.0 mm wide and 1.5 mm high (
|
||
<figureCitation id="13402A64452C842F5EECFC03FEBDFC26" box="[198,292,996,1023]" captionStart="Fig" captionStartId="14.[189,232,1648,1674]" captionTargetBox="[257,1330,266,1607]" captionTargetId="figure@14.[257,1330,266,1607]" captionTargetPageId="14" captionText="Fig. 5. SEM images of skeletal elements of Ophiocoma (Breviturma subgen. nov.) brevipes Peters, 1851 (SMNH-133230), O. (B.) dentata Müller & Troschel, 1842 (SMNH-133232) and O. krohi sp. nov. paratype (SMNH-Type-8535). A-C. Oral plates, abradial aspect at left, orientated with dorsal side up. A. O. brevipes. B. O. dentata. C. O. krohi sp. nov. D-F. Dental plates, external aspect left, internal aspect right, dorsal edge upwards. D. O. brevipes. E. O. dentata. F. O. krohi sp. nov. G-I. Radial shields, internal aspect, proximal edge upwards. G. O. brevipes. H. O. dentata. I. O. krohi sp. nov. J-L.Vertebrae from proximal part of arm, proximal aspect left, dorsal upwards. J. O. brevipes. K. O. dentata. L. O. krohi sp. nov. M-O. Genital plates, abradial plate left, adradial plate right, distal ends up (towards radial shields). M. O. brevipes. N. O. dentata. O. O. krohi sp. nov. P-R. Lateral arm plates from proximal arm, external aspect left, internal aspect right, distal end right. P. O. brevipes. Q. O. dentata. R. O. krohi sp. nov. For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827684" httpUri="https://zenodo.org/record/3827684/files/figure.png" pageId="18" pageNumber="19">Fig. 5Q</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5EC0FBC0FE57FB98" blockId="18.[189,1399,1063,1275]" box="[234,462,1063,1089]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEAF3452C842F5EC0FBC0FE57FB98" bold="true" box="[234,462,1063,1089]" pageId="18" pageNumber="19">SMNH-Type-8531</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97FBABFC7FFB22" blockId="18.[189,1399,1063,1275]" pageId="18" pageNumber="19">
|
||
1 spm, 6.8 mm dd, granule density 152
|
||
<superScript id="7C0E9BA9452C842F5C8AFBAAFD42FB80" attach="left" box="[672,731,1099,1127]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 5, 5, 4, 4, 4, 4, 3, 3, 3, 3, 3, 3. Dorsal disc with central dark brown patch, with white central spot, dark brown radiating lines, medium brown interradial lines on light brown background, white spots at disc edge (
|
||
<figureCitation id="13402A64452C842F5B39FB71FAF0FB68" box="[1299,1385,1174,1201]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="18" pageNumber="19">Fig. 2I</figureCitation>
|
||
). Dorsal arms with transverse dark and lighter brown bands. Ventral disc light brown with irregular cream patches, oral frame and ventral arms lighter brown, oral shields dark.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5EC0FAC3FE57FAE7" blockId="18.[189,1399,1316,1454]" box="[234,462,1316,1342]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEAF3452C842F5EC0FAC3FE57FAE7" bold="true" box="[234,462,1316,1342]" pageId="18" pageNumber="19">SMNH-Type-8532</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97FAAEFAEEFA74" blockId="18.[189,1399,1316,1454]" pageId="18" pageNumber="19">
|
||
1 spm, 9.6 mm dd, granule density 116-120
|
||
<superScript id="7C0E9BA9452C842F5CE9FAADFD67FA8F" attach="left" box="[707,766,1352,1380]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3. Dorsal disc dark brown mottled, dorsal arms lighter brown with dark bands. Ventral disc light and dark brown mottled (not figured). Ventral arms and oral frame light brown, oral shields with dark patch.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5EC0FA30FE2DFA28" blockId="18.[189,1399,1495,1633]" box="[234,436,1495,1521]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEAF3452C842F5EC0FA30FE2DFA28" bold="true" box="[234,436,1495,1521]" pageId="18" pageNumber="19">MNHN-IE-4303</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97FA1BFBF8F9B9" blockId="18.[189,1399,1495,1633]" pageId="18" pageNumber="19">
|
||
1 spm, 8.1 mm dd, granule density 124
|
||
<superScript id="7C0E9BA9452C842F5CBCFA1BFD48F9D1" attach="left" box="[662,721,1530,1558]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3. Dorsal disc light brown mottled, dorsal arms light brown with dark bands. Ventral disc light and dark brown mottled, arms and oral frame crème, oral shields with dark patches.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5EC0F96EFE57F97A" blockId="18.[189,1398,1673,1811]" box="[234,462,1673,1699]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEAF3452C842F5EC0F96EFE57F97A" bold="true" box="[234,462,1673,1699]" pageId="18" pageNumber="19">SMNH-Type-8533</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97F948FE65F8CA" blockId="18.[189,1398,1673,1811]" pageId="18" pageNumber="19">
|
||
1 spm,
|
||
<quantity id="4C839B04452C842F5F38F948FEFCF910" box="[274,357,1711,1737]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="mm" value="10.0">10 mm</quantity>
|
||
dd, granule density 140-164
|
||
<superScript id="7C0E9BA9452C842F5C96F948FD61F962" attach="left" box="[700,760,1709,1737]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 5, 5, 5, 5, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3. Dorsal disc variegated white and brown, arms dark and lighter brown banded, spines annulated (
|
||
<figureCitation id="13402A64452C842F5FBAF91EFE77F8CA" box="[400,494,1785,1811]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="18" pageNumber="19">Fig. 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5EC0F8DBFE57F88F" blockId="18.[189,1398,1852,2027]" box="[234,462,1852,1878]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEAF3452C842F5EC0F8DBFE57F88F" bold="true" box="[234,462,1852,1878]" pageId="18" pageNumber="19">SMNH-Type-8536</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452C842F5E97F886FA86F832" blockId="18.[189,1398,1852,2027]" pageId="18" pageNumber="19">
|
||
1 spm, 15.5 mm dd, granule density 64-96
|
||
<superScript id="7C0E9BA9452C842F5CEEF885FC99F8B7" attach="left" box="[708,768,1888,1916]" fontSize="6" pageId="18" pageNumber="19">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3. Dorsal disc uniformly dark brown, except on radial shields white granules (
|
||
<figureCitation id="13402A64452C842F5EECF84CFEBEF81F" box="[198,295,1963,1990]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="18" pageNumber="19">Fig. 2K</figureCitation>
|
||
). DAPs dark brown, all spines weakly annulated with irregular dark stripes on white. Oral frame white with dark marks on oral shields. Ventral arm plates dark brown with white borders.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5EC0FEECFE2DFEFC" blockId="19.[189,1399,267,437]" box="[234,436,267,293]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90FEAF3452D842E5EC0FEECFE2DFEFC" bold="true" box="[234,436,267,293]" pageId="19" pageNumber="20">MNHN-IE-4302</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5E97FEC8FB8BFE6C" blockId="19.[189,1399,267,437]" pageId="19" pageNumber="20">
|
||
1 spm,
|
||
<quantity id="4C839B04452D842E5F3EFEC8FEF1FE90" box="[276,360,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="19" pageNumber="20" unit="mm" value="13.0">13 mm</quantity>
|
||
dd, granule density 96-104
|
||
<superScript id="7C0E9BA9452D842E5C9FFEC8FD69FEE2" attach="left" box="[693,752,301,329]" fontSize="6" pageId="19" pageNumber="20">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3, 3, 3. Light brown disc with irregular dark spots or blotches. Dorsal arms medium brown with widely spaced dark brown transverse bands (
|
||
<figureCitation id="13402A64452D842E5C55FE90FD46FE48" box="[639,735,375,401]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="19" pageNumber="20">Fig. 2H</figureCitation>
|
||
). Ventral disc medium and dark brown mottled, ventral arms and oral frame light brown to cream, oral shields with dark pattern.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5EC0FE3AFE2DFE2E" blockId="19.[189,1399,477,647]" box="[234,436,477,503]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90FEAF3452D842E5EC0FE3AFE2DFE2E" bold="true" box="[234,436,477,503]" pageId="19" pageNumber="20">MNHN-IE-4301</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5E97FDE6FEC5FD5E" blockId="19.[189,1399,477,647]" pageId="19" pageNumber="20">
|
||
1 spm, 11.8 mm dd, granule density 72-96
|
||
<superScript id="7C0E9BA9452D842E5C86FDE6FD7EFDD4" attach="left" box="[684,743,511,539]" fontSize="6" pageId="19" pageNumber="20">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 5, 5, 5, 5(4), 4, 4(5), 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3. Dorsal disc dark brown mottled, dorsal arms dark brown with few white marks, spines dark brown (
|
||
<figureCitation id="13402A64452D842E5F8BFDAEFE6CFDBA" box="[417,501,585,611]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="19" pageNumber="20">Fig. 2J</figureCitation>
|
||
). Oral frame cream white with light brown marks on oral shields, ventral arm plates brown.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5EC0FD48FE48FD10" blockId="19.[189,1398,687,893]" box="[234,465,687,713]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90FEAF3452D842E5EC0FD48FE48FD10" bold="true" box="[234,465,687,713]" pageId="19" pageNumber="20">URUN 2009-11108</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5E97FD34FCF3FCA4" blockId="19.[189,1398,687,893]" pageId="19" pageNumber="20">
|
||
1 spm, 11.3 mm dd, granule density 112-120
|
||
<superScript id="7C0E9BA9452D842E5CFCFD33FC88FD06" attach="left" box="[726,785,721,750]" fontSize="6" pageId="19" pageNumber="20">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 4, 4, 4, 4, 4, 4, 4, 4, 3, 3, 3, 3, 3, 3, 3, 3. Dorsal disc with dark brown star-shaped pattern, interradii brown with irregular white spots, irregular white spots in centre of disc, white granules on radial shields (not figured). Dorsal arms mostly dark brown, some lighter patches, spines dark brown. Oral frame white, brown patches on oral shields. Ventral arm plates brown with white margins.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5EC0FC42FEC2FC66" blockId="19.[189,1398,933,1104]" box="[234,347,933,959]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90FEAF3452D842E5EC0FC42FEC2FC66" bold="true" box="[234,347,933,959]" pageId="19" pageNumber="20">UF13938</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5E97FC2EFAAFFB96" blockId="19.[189,1398,933,1104]" pageId="19" pageNumber="20">
|
||
1 spm, 10.3 mm dd, granule density 112-116
|
||
<superScript id="7C0E9BA9452D842E5CE0FC2DFC9FFC0F" attach="left" box="[714,774,968,996]" fontSize="6" pageId="19" pageNumber="20">mm-2</superScript>
|
||
. Arm spine sequence 3, 3, 3, 4, 4, 5, 5, 5, 5(4), 4, 4, 4, 4, 4, 3, 3, 3. Dorsal disc light brown with irregular dark patches and spots, dorsal arm light brown with widely spaced narrow dark brown bands (
|
||
<figureCitation id="13402A64452D842E5C9BFBF6FC97FBF2" box="[689,782,1041,1067]" captionStart="Fig" captionStartId="7.[189,232,1754,1780]" captionTargetBox="[189,1398,427,1713]" captionTargetId="figure@7.[189,1398,427,1713]" captionTargetPageId="7" captionText="Fig. 2. Colour variations in species of Ophiocoma (Breviturma) subgen. nov. A-C. Ophiocoma doederleini Rowe & Pawson, 1977. A. Reticulated specimen (UF5318). B. Close-up of annulated arm spines. C. Densely reticulated specimen (UF6518). D. Ophiocoma brevipes Peters, 1851 (SMNH- 133230). E-F. Ophiocoma dentata Müller & Troschel, 1842. E. Variegated specimen (SMNH-133232). F. Uniformly reddish brown specimen (URUN2009-11047). G-L. Ophiocoma krohi sp. nov. paratypes. G. With large patches (SMNH-Type-8533). H. Small irregular spots (MNHN-IE-4302). I. Star-shaped radiating pattern (SMNH-Type-8531). J. Densely mottled (MNHN-IE-4301). K. Uniformly dark brown (SMNH-Type-8536). L. Light brown, sparsely mottled (UF13938). For institution codes see main text." figureDoi="http://doi.org/10.5281/zenodo.3827678" httpUri="https://zenodo.org/record/3827678/files/figure.png" pageId="19" pageNumber="20">Fig. 2L</figureCitation>
|
||
). Spines light brown, weakly annulated. Oral frame white with light brown pattern on oral shields, ventral arm plates light brown with white margins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361656A452D842E5E97FB90FE90F9FA" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="8BC436E1452D842E5E97FB90FEB6FB48" blockId="19.[189,303,1143,1169]" box="[189,303,1143,1169]" pageId="19" pageNumber="20">
|
||
<heading id="D08C818D452D842E5E97FB90FEB6FB48" bold="true" box="[189,303,1143,1169]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B90FEAF3452D842E5E97FB90FEB6FB48" bold="true" box="[189,303,1143,1169]" pageId="19" pageNumber="20">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5E97FB40FC1BFAAC" blockId="19.[189,1399,1191,1397]" pageId="19" pageNumber="20">
|
||
<taxonomicName id="4C7B4D62452D842E5E97FB40FE16FB18" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[189,399,1191,1217]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3452D842E5E97FB40FE16FB18" box="[189,399,1191,1217]" italics="true" pageId="19" pageNumber="20">Ophiocoma krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788452D842E5FB1FB40FE63FB18" box="[411,506,1191,1217]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C7B4D62452D842E5C89FB40FCB7FB18" authorityName="Peters" authorityYear="1851" box="[675,814,1191,1217]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3452D842E5C89FB40FCB7FB18" box="[675,814,1191,1217]" italics="true" pageId="19" pageNumber="20">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D62452D842E5D5BFB40FC6BFB18" authorityName="Muller & Troschel" authorityYear="1842" box="[881,1010,1191,1217]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452D842E5D5BFB40FC6BFB18" box="[881,1010,1191,1217]" italics="true" pageId="19" pageNumber="20">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
in maximum size, granule size and density, and in colour pattern. Although quite variable in colour, it has never been observed with reticulated or spotted pattern as
|
||
<taxonomicName id="4C7B4D62452D842E5C1DFB08FD2DFAD0" authorityName="Muller & Troschel" authorityYear="1842" box="[567,692,1263,1289]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452D842E5C1DFB08FD2DFAD0" box="[567,692,1263,1289]" italics="true" pageId="19" pageNumber="20">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D62452D842E5CC4FB08FC01FAD0" authorityName="de Loriol" authorityYear="1899" box="[750,920,1263,1289]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="doederleini">
|
||
<emphasis id="B90FEAF3452D842E5CC4FB08FC01FAD0" box="[750,920,1263,1289]" italics="true" pageId="19" pageNumber="20">O. doederleini</emphasis>
|
||
</taxonomicName>
|
||
. We examined
|
||
<specimenCount id="9D7DFD68452D842E5A65FB08FB77FAD1" box="[1103,1262,1262,1289]" pageId="19" pageNumber="20" type="generic">12 specimens</specimenCount>
|
||
and half of them showed up to five arm spines on 2-4 segments, the other half had only four. Thus they all differ in spine number from
|
||
<taxonomicName id="4C7B4D62452D842E5F8BFAD0FDBFFA89" authorityName="Peters" authorityYear="1851" box="[417,550,1334,1361]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="brevipes">
|
||
<emphasis id="B90FEAF3452D842E5F8BFAD0FDBFFA89" box="[417,550,1334,1361]" italics="true" pageId="19" pageNumber="20">O. brevipes</emphasis>
|
||
</taxonomicName>
|
||
, but some are similar to
|
||
<taxonomicName id="4C7B4D62452D842E5D6EFAD0FC26FA89" authorityName="Muller & Troschel" authorityYear="1842" box="[836,959,1334,1361]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452D842E5D6EFAD0FC26FA89" box="[836,959,1334,1361]" italics="true" pageId="19" pageNumber="20">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
in lacking a fifth spine, although they may have four spines farther out on the arm than
|
||
<taxonomicName id="4C7B4D62452D842E5CD5FABCFCE2FAAD" authorityName="Muller & Troschel" authorityYear="1842" box="[767,891,1370,1397]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="dentata">
|
||
<emphasis id="B90FEAF3452D842E5CD5FABCFCE2FAAD" box="[767,891,1370,1397]" italics="true" pageId="19" pageNumber="20">O. dentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC436E1452D842E5E97FA7AFE90F9FA" blockId="19.[189,1399,1437,1571]" pageId="19" pageNumber="20">
|
||
The specimens reported by
|
||
<bibRefCitation id="EFEA4B10452D842E5C2EFA7AFD47FA6E" author="Stohr S. & Conand C. & Boissin E." box="[516,734,1437,1463]" pageId="19" pageNumber="20" pagination="545 - 560" refId="ref15768" refString="Stohr S., Conand C. & Boissin E. 2008. Brittle stars (Echinodermata: Ophiuroidea) from La Reunion and the systematic position of Ophiocanops Koehler, 1922. Zoological Journal of the Linnean Society 153 (3): 545 - 560. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2008.00401. x" type="journal article" year="2008">
|
||
Stöhr
|
||
<emphasis id="B90FEAF3452D842E5C66FA79FD11FA6E" box="[588,648,1437,1463]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
are most likely conspecific with
|
||
<taxonomicName id="4C7B4D62452D842E5A4DFA7AFB51FA6E" authority="Stöhr & Boissin & Hoareau & Ecomar & Usr, 2013" authorityName="Stöhr & Boissin & Hoareau & Ecomar & Usr" authorityYear="2013" box="[1127,1224,1437,1463]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="krohi" status="sp. nov.">
|
||
<emphasis id="B90FEAF3452D842E5A4DFA7AFB51FA6E" box="[1127,1224,1437,1463]" italics="true" pageId="19" pageNumber="20">O. krohi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C5788452D842E5AFAFA7AFAB5FA6E" box="[1232,1324,1437,1463]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
based on colour pattern, the presence of five spines and their small size. The species has a wide geographic distribution on coral reefs across the Indian and Pacific Oceans at depths of
|
||
<quantity id="4C839B04452D842E5A4AFA02FB2EFA26" box="[1120,1207,1509,1535]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="19" pageNumber="20" unit="m" value="15.0" valueMax="30.0" valueMin="0.0">0-30 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFEA4B10452D842E5AE3FA02FF64F9FA" author="Hoareau T. B. & Boissin E. & Paulay G. & Bruggemann J. H." pageId="19" pageNumber="20" pagination="2167 - 2197" refId="ref15158" refString="Hoareau T. B., Boissin E., Paulay G. & Bruggemann J. H. 2013. The Southwestern Indian Ocean as a potential marine evolutionary hotspot: perspectives from comparative phylogeography of reef brittlestars. Journal of Biogeography 40 (11): 2167 - 2197. http: // dx. doi. org / 10.1111 / jbi. 12155" type="journal article" year="2013">
|
||
Hoareau
|
||
<emphasis id="B90FEAF3452D842E5B13FA01FAEEFA26" box="[1337,1399,1509,1535]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |