249 lines
31 KiB
XML
249 lines
31 KiB
XML
<document id="7681073E547A31AF1AB444309E139AA0" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFFADAF00D1DBDC13FE3116D2" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Genieridium Vaz-De-Mello 2008, new genus" docType="treatment" docVersion="4" lastPageNumber="27" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="26" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8E0923A9CACECD4C5C7BB69A3FB9953F">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
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<mods:name id="B6366BB575EFDD9E8A910B01D3945A61" type="personal">
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<mods:roleTerm id="2CF34B59F80910DEF0DB92A92FC19232">Author</mods:roleTerm>
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<mods:namePart id="2569512FC19A3BF3C62CDE91FFC49581">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="CC713E0ABDC4871A31D976DD08CB28CA">text</mods:typeOfResource>
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<mods:titleInfo id="733D5E391FC9889131472327226AAE4A">
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<mods:title id="65361BD0170489D587DB4A106FCAF1BE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="07194AB14CE27C7D6D05AFEB1896F4FF">
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<mods:date id="348A61CB28CBB6C4C9091F9E5C197AB8">2008</mods:date>
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<mods:detail id="5A4BE4CE66371EB406FB12114EE13850" type="pubDate">
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<mods:number id="855F9FB7D985E854B723B9644807C2DF">2008-12-05</mods:number>
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<mods:detail id="41244CDA68E69A0A4E6EC710AE6730C5" type="volume">
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<mods:number id="856752C94A77FB100A91EC4CEE8F030A">1955</mods:number>
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<mods:end id="DFABA7A9A93FDA0421F58157EAFE0FB4">75</mods:end>
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<mods:classification id="3C51629C8913425894DA3C5FA326E20F">journal article</mods:classification>
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<mods:identifier id="928B6ED821A4C3741FD85A4CF385F6D0" type="ISSN">11755334</mods:identifier>
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</mods:mods>
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<treatment id="03D287CEFFADAF00D1DBDC13FE3116D2" ID-DOI="http://doi.org/10.5281/zenodo.5241995" ID-Zenodo-Dep="5241995" LSID="urn:lsid:plazi:treatment:03D287CEFFADAF00D1DBDC13FE3116D2" httpUri="http://treatment.plazi.org/id/03D287CEFFADAF00D1DBDC13FE3116D2" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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<subSubSection id="C3616553FFADAF03D1DBDC13FE7010D9" box="[151,470,871,897]" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8BC436D8FFADAF03D1DBDC13FE7010D9" blockId="25.[151,470,871,897]" box="[151,470,871,897]" pageId="25" pageNumber="26">
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<heading id="D08C81B4FFADAF03D1DBDC13FE7010D9" bold="true" box="[151,470,871,897]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="B90FEACAFFADAF03D1DBDC13FE7010D9" bold="true" box="[151,470,871,897]" pageId="25" pageNumber="26">
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6.
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<taxonomicName id="4C7B4D5BFFADAF03D1F8DC13FEEB10D9" ID-CoL="4MST" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[180,333,871,897]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90FEACAFFADAF03D1F8DC13FEEB10D9" bold="true" box="[180,333,871,897]" italics="true" pageId="25" pageNumber="26">Genieridium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23C57B1FFADAF03D018DC13FE7010D9" box="[340,470,871,897]" pageId="25" pageNumber="26" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFADAF03D1DBDCC6FEAB1654" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph id="8BC436D8FFADAF03D1DBDCC6FEAB1654" blockId="25.[151,1436,946,2012]" pageId="25" pageNumber="26">
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<emphasis id="B90FEACAFFADAF03D1DBDCC6FEBF1094" bold="true" box="[151,281,946,972]" pageId="25" pageNumber="26">Diagnosis:</emphasis>
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Moderate to large-sized species (only rare specimens of
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<taxonomicName id="4C7B4D5BFFADAF03D288DCC6FBE91093" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Vaz-De-Mello" baseAuthorityYear="2008" box="[964,1103,946,971]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cryptops">
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<emphasis id="B90FEACAFFADAF03D288DCC6FBE91093" box="[964,1103,946,971]" italics="true" pageId="25" pageNumber="26">G. cryptops</emphasis>
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</taxonomicName>
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measure less than
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<quantity id="4C839B3DFFADAF03D463DCC6FA291094" box="[1327,1423,946,972]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="25" pageNumber="26" unit="mm" value="3.7">3.7 mm</quantity>
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), oval-elongate shape (
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<figureCitation id="13402A5DFFADAF03D0D4DCAEFD8A10AC" box="[408,556,986,1012]" captionStart="FIGURES 15–22" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 15–22. Dorsal views. 15–16: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 21–22: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="25" pageNumber="26">Figs. 21–22</figureCitation>
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); clypeal teeth two (Figs. 55, 60, 62, 64, 70) or none, in the latter case clypeus widely emarginated (Figs. 57, 68). Eyes, in dorsal view, absent or very small, distanced by more than 12 times eye width. Pronotum separated from hypomeron by weak carina (Fig. 59). Elytra convex at least in the anterior half. Length of mesotibiae and metatibiae at least three times apical width. Males with mesal apical protibial teeth; all claws bent (mesoclaws and metaclaws less modified than proclaws); and last tarsomere of prolegs modified to receive claws. Length of last abdominal sternite along midline equal to or less than medial width of metafemur; pygidium, in ventral view, occupying more than two-fifths of abdominal horizontal length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFADAF00D18ADA6EFDDC1272" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
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<paragraph id="8BC436D8FFADAF03D18ADA6EFC0514EC" blockId="25.[151,1436,946,2012]" pageId="25" pageNumber="26">
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<emphasis id="B90FEACAFFADAF03D18ADA6EFEC6166C" bold="true" box="[198,352,1306,1332]" pageId="25" pageNumber="26">Description:</emphasis>
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Length
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<quantity id="4C839B3DFFADAF03D08FDA6EFDF5166C" box="[451,595,1306,1332]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.5" metricValueMin="3.5" pageId="25" pageNumber="26" unit="mm" value="5.0" valueMax="6.5" valueMin="3.5">3.5–6.5 mm</quantity>
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, body oval elongated, convex (
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<figureCitation id="13402A5DFFADAF03D28DDA6EFBE9166C" box="[961,1103,1306,1332]" captionStart="FIGURES 15–22" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 15–22. Dorsal views. 15–16: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 21–22: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="25" pageNumber="26">Figs. 21–22</figureCitation>
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), color grey to black, brown in teneral specimens, shiny or opaque, lacking any traces of metallic sheen. Clypeus with two teeth or none, always with central anterior concavity coincident with emargination, which is also present in toothless species (Figs. 57, 68). Clypeus laterally regularly rounded, continuous with genal border. Clypeofrontal and clypeogenal sutures indistinct, head lacking any traces of carinae or tubercles. Eyes, in dorsal view, absent or very small, when present elongated, more than three times their width; interocular width at least 12 times eye width (Figs. 55, 57, 60, 62, 64, 68, 70). Pronotum lacking anterior or posterior beads, simply convex, with lateral callosities distinguished only by lack of punctation; disc covered by simple setose punctures and separated of hypomeron by weak but distinct carina interrupted by regular setose punctures. Hypomeron ventrally with strong lateral longitudinal carina; mesoepimera with transverse anterior carina stronger mesally. Metasternum with simple setose punctures on disc. Elytra with one or two rows of setose punctures, discal interstriae flat all along; striae apically deeper and slightly wider than on disc. Pseudoepipleura gradually narrowed posteriorly, superiorly with complete row of setose punctures. Protibiae with three strong teeth distributed along at least apical three-fifths, basally denticulate, with ventral scale-like setae. Mesotibiae and metatibiae strongly widened apically, apical width more than third of tibial length. Parameres flattened, right-angled in relation to phallobase, shorter than half of phallobase length. Spermatheca simply C-shaped, in one case (
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<taxonomicName id="4C7B4D5BFFADAF03D447D807FA3214D4" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Vaz-De-Mello" baseAuthorityYear="2008" box="[1291,1428,1906,1932]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="cryptops">
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<emphasis id="B90FEACAFFADAF03D447D807FA8114D4" box="[1291,1319,1907,1932]" italics="true" pageId="25" pageNumber="26">G.</emphasis>
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cryptops
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</taxonomicName>
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) with elongated base and apex. Coxites triangular and symmetrical.
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</paragraph>
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<paragraph id="8BC436D8FFADAF00D18AD8B6FDDC1272" blockId="25.[151,1436,946,2012]" lastBlockId="26.[151,1437,152,1418]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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<emphasis id="B90FEACAFFADAF03D18AD8B6FE621484" bold="true" box="[198,452,1986,2012]" pageId="25" pageNumber="26">Sexual dimorphism:</emphasis>
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Males have proclaws bent at midlength, and mesoclaws and metaclaws also modified, either bent at midlength or larger than in females and with stronger basal angle; protarsus with last tarsomere modified to receive claws; metasternal disc strongly concave (less concave or flat in females), pygidium narrower than in females, and each tibia with apicomesal tooth, sometimes modified in laminate anteriorly directed apodeme in metatibia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFAEAF00D18ADE43FE611222" pageId="26" pageNumber="27" type="type_taxon">
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<paragraph id="8BC436D8FFAEAF00D18ADE43FE611222" blockId="26.[151,1437,152,1418]" pageId="26" pageNumber="27">
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<emphasis id="B90FEACAFFAEAF00D18ADE43FECD1209" bold="true" box="[198,363,311,337]" pageId="26" pageNumber="27">
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<typeStatus id="54C0887AFFAEAF00D18ADE43FEA51209" box="[198,259,311,337]" pageId="26" pageNumber="27">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C7B4D5BFFAEAF00D03FDE4CFCA9120A" ID-CoL="7693N" authority="Balthasar, 1942" authorityName=": Balthasar" authorityYear="1938" box="[371,783,312,338]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bidens">
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<emphasis id="B90FEACAFFAEAF00D03FDE4CFDF61209" box="[371,592,312,337]" italics="true" pageId="26" pageNumber="27">Pedaridium bidens</emphasis>
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<bibRefCitation id="EFEA4B29FFAEAF00D315DE4CFCA9120A" author="Balthasar, V." box="[601,783,312,338]" pageId="26" pageNumber="27" pagination="36 - 44" refId="ref47394" refString="Balthasar, V. (1942) Neue Scarabeiden aus Sud-Amerika. 72. Beitrag zur Kenntnis der Scarabeiden (Col.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 39, 36 - 44." type="journal article" year="1942">Balthasar, 1942</bibRefCitation>
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</taxonomicName>
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(present designation) =
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<taxonomicName id="4C7B4D5BFFAEAF00D561DE4CFF781222" ID-CoL="3FKVH" authority="(Balthasar, 1942)" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Vaz-De-Mello" baseAuthorityYear="1938" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bidens" status="comb. nov.">
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<emphasis id="B90FEACAFFAEAF00D561DE4CFAB31209" box="[1069,1301,312,337]" italics="true" pageId="26" pageNumber="27">Genieridium bidens</emphasis>
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(
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<bibRefCitation id="EFEA4B29FFAEAF00D469DE4CFF731222" author="Balthasar, V." pageId="26" pageNumber="27" pagination="36 - 44" refId="ref47394" refString="Balthasar, V. (1942) Neue Scarabeiden aus Sud-Amerika. 72. Beitrag zur Kenntnis der Scarabeiden (Col.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 39, 36 - 44." type="journal article" year="1942">Balthasar, 1942</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="B90FEACAFFAEAF00D1A6DE2BFE191221" bold="true" box="[234,447,351,377]" pageId="26" pageNumber="27">
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<taxonomicNameLabel id="A23C57B1FFAEAF00D1A6DE2BFE191221" box="[234,447,351,377]" pageId="26" pageNumber="27" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFAEAF00D18ADEF3FE3E1292" pageId="26" pageNumber="27" type="etymology">
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<paragraph id="8BC436D8FFAEAF00D18ADEF3FE3E1292" blockId="26.[151,1437,152,1418]" pageId="26" pageNumber="27">
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<emphasis id="B90FEACAFFAEAF00D18ADEF3FEF112F9" bold="true" box="[198,343,391,417]" pageId="26" pageNumber="27">Etymology:</emphasis>
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After François Génier, good friend and scarabeidologist, companion in so many works and ideas. Gender neutral.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFAEAF00D18ADEA3FEFF11E2" pageId="26" pageNumber="27" type="distribution">
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<paragraph id="8BC436D8FFAEAF00D18ADEA3FEFF11E2" blockId="26.[151,1437,152,1418]" pageId="26" pageNumber="27">
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<emphasis id="B90FEACAFFAEAF00D18ADEA3FECF12A9" bold="true" box="[198,361,471,497]" pageId="26" pageNumber="27">Distribution:</emphasis>
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<collectingCountry id="F36C7648FFAEAF00D03EDEACFE4E12AA" box="[370,488,472,498]" name="Colombia" pageId="26" pageNumber="27">Colombia</collectingCountry>
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,
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<collectingCountry id="F36C7648FFAEAF00D0B9DEACFDD612AA" box="[501,624,472,498]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
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, non-Amazonian
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<collectingCountry id="F36C7648FFAEAF00D20FDEACFC2812AA" box="[835,910,472,498]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
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, eastern
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<collectingCountry id="F36C7648FFAEAF00D2BBDEACFBF612AA" box="[1015,1104,472,498]" name="Bolivia" pageId="26" pageNumber="27">Bolivia</collectingCountry>
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,
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<collectingCountry id="F36C7648FFAEAF00D511DEACFB6D12AA" box="[1117,1227,472,498]" name="Paraguay" pageId="26" pageNumber="27">Paraguay</collectingCountry>
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, and northeastern
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<collectingCountry id="F36C7648FFAEAF00D1DBDD74FEB61142" box="[151,272,512,538]" name="Argentina" pageId="26" pageNumber="27">Argentina</collectingCountry>
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. The genus shows a disjunct distribution, with one group in the Caribbean subregion (Northwestern South American dominion), occupying the Cauca, Maracaibo, Venezuelan Coast and Venezuelan Llanos provinces, and the other in the Chacoan, Parana and Amazonian subregions, occupying the Pantanal, Caatinga, Cerrado,
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<collectingRegion id="49BFF83AFFAEAF00D005DD0CFE3111CA" box="[329,407,632,658]" country="Argentina" name="Chaco" pageId="26" pageNumber="27">Chaco</collectingRegion>
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, Brazilian Atlantic Forest, Parana Forest, and
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<taxonomicName id="4C7B4D5BFFAEAF00D28DDD0CFA8611CA" authority="Forest" authorityName="Forest" box="[961,1312,632,658]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="angustifolia">
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<emphasis id="B90FEACAFFAEAF00D28DDD0CFB6811C9" box="[961,1230,632,657]" italics="true" pageId="26" pageNumber="27">Araucaria angustifolia</emphasis>
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Forest
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</taxonomicName>
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provinces (
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<figureCitation id="13402A5DFFAEAF00D1D3DDD4FEEB11E2" box="[159,333,672,698]" captionStart-0="FIGURES 103–104" captionStart-1="FIGURES 105–106" captionStart-2="FIGURES 107–108" captionStartId-0="67.[151,269,1939,1963]" captionStartId-1="68.[151,269,1958,1982]" captionStartId-2="69.[151,269,1969,1993]" captionTargetBox-0="[181,1405,194,1916]" captionTargetBox-1="[177,1411,194,1930]" captionTargetBox-2="[166,1417,182,1945]" captionTargetId-0="figure-34@67.[181,1405,194,1916]" captionTargetId-1="figure-35@68.[177,1411,194,1930]" captionTargetId-2="figure-37@69.[166,1420,182,1945]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionTargetPageId-2="69" captionText-0="FIGURES 103–104. Distribution maps. 103. Degallieridium lilliputanum (circles) and Feeridium woodruffi (triangles); 104. Genieridium bidens (circles) and G. paranense (triangles)." captionText-1="FIGURES 105–106. Distribution maps. 105. Genieridium cryptops; 106. G. margareteae (circles) and G. zanunciorum (triangles)." captionText-2="FIGURES 107–108. Distribution maps. 107. Genieridium medinae (circles) and G. bordoni (triangles); 108. Nunoidium argentinum (circles), Pereiraidium almeidai (squares) and Silvinha unica (triangles)." pageId="26" pageNumber="27">Figs. 104–107</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFAEAF00D18ADDB3FE3116D2" pageId="26" pageNumber="27" type="discussion">
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<paragraph id="8BC436D8FFAEAF00D18ADDB3FDE317C2" blockId="26.[151,1437,152,1418]" pageId="26" pageNumber="27">
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<emphasis id="B90FEACAFFAEAF00D18ADDB3FEE711B9" bold="true" box="[198,321,711,737]" pageId="26" pageNumber="27">Remarks:</emphasis>
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The position of
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<taxonomicName id="4C7B4D5BFFAEAF00D359DDBCFD0F11B9" authorityName="Vaz-De-Mello" authorityYear="2008" box="[533,681,712,737]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFFAEAF00D359DDBCFD0F11B9" box="[533,681,712,737]" italics="true" pageId="26" pageNumber="27">Genieridium</emphasis>
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</taxonomicName>
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was not resolved, and it was recovered in a trichotomy with
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D1DBDD84FE871051" authorityName="Harold" authorityYear="1868" box="[151,289,752,777]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFAEAF00D1DBDD84FE871051" box="[151,289,752,777]" italics="true" pageId="26" pageNumber="27">Pedaridium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B90FEACAFFAEAF00D013DD84FE4F1051" box="[351,489,752,777]" italics="true" pageId="26" pageNumber="27">
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D013DD84FE431051" authorityName="Vaz-De-Mello" authorityYear="2008" box="[351,485,752,777]" class="Insecta" family="Scarabaeidae" genus="Nunoidium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Nunoidium</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The new genus main external synapomorphies are the elongation and inclined orientation of pygidium, and the punctation interrupting the pronotal-hypomeral carina. Apart from characters mentioned as synapomorphies of
|
||
<emphasis id="B90FEACAFFAEAF00D369DC34FD661001" box="[549,704,832,857]" italics="true" pageId="26" pageNumber="27">
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D369DC34FD1A1001" authorityName="Vaz-De-Mello" authorityYear="2008" box="[549,700,832,857]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Genieridium</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
it can be distinguished from
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D550DC34FB001001" authorityName="Harold" authorityYear="1868" box="[1052,1190,832,857]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFAEAF00D550DC34FB001001" box="[1052,1190,832,857]" italics="true" pageId="26" pageNumber="27">Pedaridium</emphasis>
|
||
</taxonomicName>
|
||
by the much shorter phallobase, differently shaped internal sac sclerites (
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D240DC1CFC3010D9" authorityName="Harold" authorityYear="1868" box="[780,918,872,897]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFAEAF00D240DC1CFC3010D9" box="[780,918,872,897]" italics="true" pageId="26" pageNumber="27">Pedaridium</emphasis>
|
||
</taxonomicName>
|
||
lacks all but the pseudoflagellum), lack of basal “tooth” in the spermatheca, elytra conjointly convex (
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D200DCE4FC7010F1" authorityName="Harold" authorityYear="1868" box="[844,982,912,937]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFAEAF00D200DCE4FC7010F1" box="[844,982,912,937]" italics="true" pageId="26" pageNumber="27">Pedaridium</emphasis>
|
||
</taxonomicName>
|
||
has clearly tectiform elytra, separately flattened), lack of longer setae on the anterior part of pronotum and lack of a pronotal lateral expansion. From
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D1DBDC94FEBF10A1" authorityName="Vaz-De-Mello" authorityYear="2008" box="[151,281,992,1017]" class="Insecta" family="Scarabaeidae" genus="Nunoidium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFAEAF00D1DBDC94FEBF10A1" box="[151,281,992,1017]" italics="true" pageId="26" pageNumber="27">Nunoidium</emphasis>
|
||
</taxonomicName>
|
||
it is readily distinguished by the lack of a posterior pronotal bead, reduction of lateral bead, reduced eyes, and presence of sexually dimorphic claws. This genus includes two clearly delimited monophyletic, biogeographically-defined groups: species from
|
||
<collectingCountry id="F36C7648FFAEAF00D22CDB44FC7E1712" box="[864,984,1072,1098]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
|
||
and
|
||
<collectingCountry id="F36C7648FFAEAF00D56CDB44FB321712" box="[1056,1172,1072,1098]" name="Colombia" pageId="26" pageNumber="27">Colombia</collectingCountry>
|
||
are characterized by strong modification of male metalegs (with flattened, blade-like apicomesal tooth) and southern species by the reduction of the seventh elytral stria.
|
||
</paragraph>
|
||
<paragraph id="8BC436D8FFAEAF00D18ADBD3FE3116D2" blockId="26.[151,1437,152,1418]" pageId="26" pageNumber="27">
|
||
<emphasis id="B90FEACAFFAEAF00D18ADBD3FEC91799" bold="true" box="[198,367,1191,1217]" pageId="26" pageNumber="27">Composition:</emphasis>
|
||
Besides the
|
||
<typeStatus id="54C0887AFFAEAF00D343DBDCFDE7179A" box="[527,577,1192,1218]" pageId="26" pageNumber="27">type</typeStatus>
|
||
species, the genus is presently defined to include
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D5E0DBDCFF7817B2" authority="(Arrow, 1913)" baseAuthorityName="Arrow" baseAuthorityYear="1913" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cryptops" status="comb. nov.">
|
||
<emphasis id="B90FEACAFFAEAF00D5E0DBDCFA931799" box="[1196,1333,1192,1217]" italics="true" pageId="26" pageNumber="27">G. cryptops</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFAEAF00D404DBDCFF7317B2" author="Arrow, G. J." pageId="26" pageNumber="27" pagination="456 - 466" refId="ref47114" refString="Arrow, G. J. (1913) Some new species of Lamellicorn beetles from Brazil. The Annals and Magazine of Natural History, Series 8, 11, 456 - 466." type="journal article" year="1913">Arrow, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90FEACAFFAEAF00D1A6DBBBFE6717B1" bold="true" box="[234,449,1231,1257]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A23C57B1FFAEAF00D1A6DBBBFE6717B1" box="[234,449,1231,1257]" pageId="26" pageNumber="27" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D09CDBA4FCBB17B2" authority="(Arrow, 1932)" baseAuthorityName="Arrow" baseAuthorityYear="1932" box="[464,797,1232,1258]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="paranense" status="comb. nov.">
|
||
<emphasis id="B90FEACAFFAEAF00D09CDBA4FDCD17B1" box="[464,619,1232,1257]" italics="true" pageId="26" pageNumber="27">G. paranense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFAEAF00D336DBA4FCB217B2" author="Arrow, G. J." box="[634,788,1232,1258]" pageId="26" pageNumber="27" pagination="223 - 226" refId="ref47190" refString="Arrow, G. J. (1932) New species of Lamellicorn beetles (subfamily Coprinae) from South America. Stylops, 1 (10), 223 - 226." type="journal article" year="1932">Arrow, 1932</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90FEACAFFAEAF00D265DBBBFBA617B1" bold="true" box="[809,1024,1231,1257]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A23C57B1FFAEAF00D265DBBBFBA617B1" box="[809,1024,1231,1257]" pageId="26" pageNumber="27" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D55CDBA4FAFB17B2" authority="(Martinez, 1992)" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Martinez" baseAuthorityYear="1992" box="[1040,1373,1232,1258]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bordoni" status="comb. nov.">
|
||
<emphasis id="B90FEACAFFAEAF00D55CDBA4FB2A17B1" box="[1040,1164,1232,1257]" italics="true" pageId="26" pageNumber="27">G. bordoni</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFAEAF00D5D7DBA4FAF217B2" author="Martinez, A." box="[1179,1364,1232,1258]" pageId="26" pageNumber="27" pagination="21 - 25" refId="ref49829" refString="Martinez, A. (1992) Una nueva especie de Pedaridium (Col. Scarabaeidae, Coprini). Gayana Zoologica, 56 (1 - 2), 21 - 25." type="journal article" year="1992">Martínez, 1992</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90FEACAFFAEAF00D425DBBBFE951649" bold="true" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A23C57B1FFAEAF00D425DBBBFE951649" pageId="26" pageNumber="27" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D00FDB8CFE5B1649" baseAuthorityName="Vaz-de-Mello & Canhedo" baseAuthorityYear="1998" box="[323,509,1272,1297]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zanunciorum">
|
||
<emphasis id="B90FEACAFFAEAF00D00FDB8CFE5B1649" box="[323,509,1272,1297]" italics="true" pageId="26" pageNumber="27">G. zanunciorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFAEAF00D340DB8CFC2D164A" author="Vaz-de-Mello, F. Z. & Canhedo, V. L." box="[524,907,1272,1298]" pageId="26" pageNumber="27" pagination="95 - 100" refId="ref51217" refString="Vaz-de-Mello, F. Z. & Canhedo, V. L. (1998) Duas novas especies de Pedaridium (Coleoptera, Scarabaeidae, Coprinae, Ateuchini). Iheringia, Serie Zoologia, 84, 95 - 100." type="journal article" year="1998">Vaz-de-Mello & Canhedo, 1998</bibRefCitation>
|
||
),
|
||
<emphasis id="B90FEACAFFAEAF00D2ECDB83FBD01649" bold="true" box="[928,1142,1271,1297]" pageId="26" pageNumber="27">new combination</emphasis>
|
||
;
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D5CADB8CFA9C1649" baseAuthorityName="Genier & Vaz-de-Mello" baseAuthorityYear="2002" box="[1158,1338,1272,1297]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="margareteae">
|
||
<emphasis id="B90FEACAFFAEAF00D5CADB8CFA9C1649" box="[1158,1338,1272,1297]" italics="true" pageId="26" pageNumber="27">G. margareteae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFAEAF00D406DB8CFE001662" author="Genier, F. & Vaz-de-Mello, F. Z." pageId="26" pageNumber="27" pagination="185 - 196" refId="ref48361" refString="Genier, F. & Vaz-de-Mello, F. Z. (2002) A review of Arrow's types of Trichillum and Pedaridium (Coleoptera: Scarabaeinae, Ateuchini) with description of two new species of Pedaridium. Acta Zoologica Cracoviensia, 45 (3), 185 - 196." type="journal article" year="2002">Génier & Vaz-de-Mello, 2002</bibRefCitation>
|
||
),
|
||
<emphasis id="B90FEACAFFAEAF00D0F7DA6BFD371661" bold="true" box="[443,657,1311,1337]" pageId="26" pageNumber="27">new combination</emphasis>
|
||
; and
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D399DA54FCFF1661" baseAuthorityName="Gill & Vaz-de-Mello" baseAuthorityYear="2002" box="[725,857,1312,1337]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="medinae">
|
||
<emphasis id="B90FEACAFFAEAF00D399DA54FCFF1661" box="[725,857,1312,1337]" italics="true" pageId="26" pageNumber="27">G. medinae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFAEAF00D225DA54FB091662" author="Gill, B. D. & Vaz-de-Mello, F. Z." box="[873,1199,1312,1338]" pageId="26" pageNumber="27" pagination="47 - 51" refId="ref48413" refString="Gill, B. D. & Vaz-de-Mello, F. Z. (2003) An unusual new species of Pedaridium Harold (Coleoptera: Scarabaeidae: Ateuchini) from Colombia. Journal of the Entomological Society of Ontario, 133, 47 - 51." type="journal article" year="2003">Gill & Vaz-de-Mello, 2003</bibRefCitation>
|
||
),
|
||
<emphasis id="B90FEACAFFAEAF00D589DA6BFA3A1661" bold="true" box="[1221,1436,1311,1337]" pageId="26" pageNumber="27">new combination</emphasis>
|
||
(all described as
|
||
<taxonomicName id="4C7B4D5BFFAEAF00D02ADA3CFE561639" authorityName="Harold" authorityYear="1868" box="[358,496,1352,1377]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFAEAF00D02ADA3CFE561639" box="[358,496,1352,1377]" italics="true" pageId="26" pageNumber="27">Pedaridium</emphasis>
|
||
</taxonomicName>
|
||
). Because no undescribed species of this genus are known to me, I present a small synopsis below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |