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<document id="226578388CF7CA2CA40F98CA74F73619" ID-DOI="10.5281/zenodo.4665549" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665549" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712630742" checkinUser="felipe" docAuthor="Peigné, Stéphane, Morlo, Michael, Chaimanee, Yaowalak, Ducrocq, Stéphane, Tun, Soe Thura &amp; Jaeger, Jean-Jacques" docDate="2007" docId="03D287BCA70CFFEE186EFB18FDE7F952" docLanguage="en" docName="Geodiversitas.29.3.441-458.pdf" docOrigin="Geodiversitas 29 (3)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Hyaenodontidae Leidy 1869" docType="treatment" docVersion="3" lastPageNumber="454" masterDocId="FFEBFFC4A707FFE31B28FFEBFFBAFF98" masterDocTitle="New discoveries of hyaenodontids (Creodonta, Mammalia) from the Pondaung Formation, middle Eocene, Myanmar - paleobiogeographic implications" masterLastPageNumber="458" masterPageNumber="441" pageNumber="452" updateTime="1698900954563" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="00891AE29C8E4AADD0172FD97B32196C">New discoveries of hyaenodontids (Creodonta, Mammalia) from the Pondaung Formation, middle Eocene, Myanmar - paleobiogeographic implications</mods:title>
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<mods:namePart id="43FD469F2FF5D6F12F5737E589464847">Peigné, Stéphane</mods:namePart>
<mods:affiliation id="465BC04734A527FC1B3A762FA563F76A">Muséum national dHistoire naturelle, Département Histoire de la Terre, USM 0203 / UMR 5143 Paléobiodiversité et Paléoenvironnements, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) and Université de Poitiers, UFR SFA, UMR 6046 CNRS, Géobiologie, Biochronologie et Paléontologie humaine, 40 avenue du Recteur Pineau, F- 86022 Poitiers cedex (France) peigne @ mnhn. fr</mods:affiliation>
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<mods:namePart id="3264D176260D15B06E84A199AEAA6D65">Morlo, Michael</mods:namePart>
<mods:affiliation id="4E75E7D299241927E61D5637EF076DA3">Forschungsinstitut Senckenberg, Abteilung Messelforschung, Senckenberganlage 25, D- 60325 Frankfurt (Germany) michael. morlo @ senckenberg. de</mods:affiliation>
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<mods:namePart id="223BD98930A2568EE5B892581C7DD2B4">Chaimanee, Yaowalak</mods:namePart>
<mods:affiliation id="81EAC6018AFF4D618AF9740122C62438">Department of Mineral Resources, Geological Survey Division, Rama IV Road, Bangkok 10400 (Thailand) yaowalak @ dmr. go. th</mods:affiliation>
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<mods:namePart id="3FE076E760CBCA0F30A7C2FCF5D47E1F">Ducrocq, Stéphane</mods:namePart>
<mods:affiliation id="3C519932CF1FD0AF22CF668EC36CEABE">Université de Poitiers, UFR SFA, UMR 6046 CNRS, Géobiologie, Biochronologie et Paléontologie humaine, 40 avenue du Recteur Pineau, F- 86022 Poitiers cedex (France) stephane. ducrocq @ univ-poitiers. fr</mods:affiliation>
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<mods:namePart id="FF55BD60679E4B090A49B3D213BA40B1">Tun, Soe Thura</mods:namePart>
<mods:affiliation id="0EDE53DC54B921D38B8D3A9DBFBD3A1C">Department of Geology, University of Yangon, Yangon 11041 (Myanmar) htar @ myanmar. com. mm</mods:affiliation>
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<mods:namePart id="10FE2688A0BE813C456B29FC06F34FFD">Jaeger, Jean-Jacques</mods:namePart>
<mods:affiliation id="2BDF160A85C96B354CC3ABA304CC41F7">Université de Poitiers, UFR SFA, UMR 6046 CNRS, Géobiologie, Biochronologie et Paléontologie humaine, 40 avenue du Recteur Pineau, F- 86022 Poitiers cedex (France) jean-jacques. jaeger @ univ-poitiers. fr</mods:affiliation>
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<mods:date id="9C714F7F1D4AFB952DD5F407333BE97D">2007</mods:date>
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<paragraph id="8BC436AAA70CFFE8186EFB18FBDFFA96" blockId="11.[838,1125,1267,1294]" box="[838,1125,1267,1294]" pageId="11" pageNumber="452">
<heading id="D08C81C6A70CFFE8186EFB18FBDFFA96" box="[838,1125,1267,1294]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="452" reason="2">
<taxonomicName id="4C7B4D29A70CFFE8186EFB18FC40FA96" authorityName="Leidy" authorityYear="1869" box="[838,1018,1267,1294]" class="Mammalia" family="Hyaenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="11" pageNumber="452" phylum="Chordata" rank="family">Hyaenodontidae</taxonomicName>
sp. indet.
</heading>
</paragraph>
<paragraph id="8BC436AAA70CFFE819E2FAC6FB15FAC7" blockId="11.[714,1247,1324,1375]" pageId="11" pageNumber="452">
REFERRED MATERIAL. — From Thamingyauk:NMM Tmk- 500 (isolated subcomplete right M3,
<figureCitation id="13402A2FA70CFFE81F1FFAACFB19FAC7" box="[1079,1187,1351,1375]" captionStart="FIG" captionStartId="5.[151,162,1627,1644]" captionTargetBox="[269,1132,258,1568]" captionTargetId="graphics-153@5.[255,1147,249,1598]" captionTargetPageId="5" captionText="FIG. 2. — A-K, Kyawdawia lupina Egi et al., 2005 from Pondaung Formation,Myanmar.From Bahin:NMM Bhn-1040,right M1 in occlusal (A) and labial (B) views; from Thandaung: NMM Tdg-801, fragment of left M3 in distal (C) and occlusal (D) views; left M2 in occlusal (E), lingual (F) and labial (G) views; NMM Tdg-802, fragment of right P4 in labial (H) and occlusal (I) views; from Kyaukmagyi: NMM Kmi-500, fragment of right M2 in occlusal (J) and lingual (K) views; L, M, Hyaenodontidae indet. from Pondaung Formation, Myanmar." figureDoi="http://doi.org/10.5281/zenodo.4665553" httpUri="https://zenodo.org/record/4665553/files/figure.png" pageId="11" pageNumber="452">Fig. 2L, M</figureCitation>
).
</paragraph>
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<subSubSection id="C3616521A70CFFE819E2FA9CFBF2FA31" pageId="11" pageNumber="452" type="distribution">
<paragraph id="8BC436AAA70CFFE819E2FA9CFBF2FA31" blockId="11.[714,1248,1399,1449]" pageId="11" pageNumber="452">
DISTRIBUTION AND AGE.  Pondaung Formation,
<collectingCountry id="F36C763AA70CFFE81F9EFA9CFCB2FA31" name="Myanmar" pageId="11" pageNumber="452">Myanmar</collectingCountry>
(see
<figureCitation id="13402A2FA70CFFE81811FA7AFCCAFA31" box="[825,880,1425,1449]" captionStart="FIG" captionStartId="3.[151,162,1353,1370]" captionTargetBox="[209,1184,252,1307]" captionTargetId="figure-13@3.[208,1192,252,1310]" captionTargetPageId="3" captionText="FIG. 1. — Geographical map of the Pondaung area (Myanmar) showing the vertebrate fossil localities in the Pondaung Formation that have yielded creodont material; in these names, “Kyitchaung” means “locality” in Myanmar language: 1, Kyawdaw Kyitchaung (Kdw); 2, Thandaung Kyitchaung (Tdg); 3, Kyaukmagyi Kyitchaung (Kmi); 4, Lema Kyitchaung (Lma); 5, UCMP V83106; 6, fossiliferous site 2 km N of Thadut (Orienspterodon); 7, Paukkaung Kyitchaung (Pk 2); 8, Yarshe Kyitchaung (Yse); 9, Thamingyauk Kyitchaung (Tmk); 10, Pangan Kyitchaung (Pgn); 11, Taungnigyin Kyitchaung (= Pangan 2; Pgn 2); 12, Myinthagya Kyitchaung (Mta). Modified after Aung Naing Soe (2004)." figureDoi="http://doi.org/10.5281/zenodo.4665551" httpUri="https://zenodo.org/record/4665551/files/figure.png" pageId="11" pageNumber="452">Fig. 1</figureCitation>
); late middle Eocene.
</paragraph>
</subSubSection>
<subSubSection id="C3616521A70CFFEE19E2FA39FDE7F952" lastPageId="13" lastPageNumber="454" pageId="11" pageNumber="452" type="description">
<paragraph id="8BC436AAA70CFFE819E2FA39FCE1FA72" blockId="11.[713,1247,1490,1706]" box="[714,859,1490,1516]" pageId="11" pageNumber="452">DESCRIPTION</paragraph>
<paragraph id="8BC436AAA70CFFE819E2FA1AFB65F932" blockId="11.[713,1247,1490,1706]" pageId="11" pageNumber="452">The tooth is complete except for the mesiolabial corner. It is large and transversely elongated. The parastylar wing, which develops transversely, is broken, but it is likely to be short. The paracone is slightly larger than the metacone, which is located roughly distally to it. Both cusps are separated by an</paragraph>
<paragraph id="8BC436AAA70CFFE819E0F944FB67F952" blockId="11.[712,1245,1711,1738]" box="[712,1245,1711,1738]" pageId="11" pageNumber="452">open V-shaped valley continuing by a small notch,</paragraph>
<paragraph id="8BC436AAA70BFFEF1864FF98FB72FF1C" blockId="12.[844,1224,115,133]" box="[844,1224,115,133]" pageId="12" pageNumber="453">
Creodonts from the Pondaung Formation,
<collectingCountry id="F36C763AA70BFFEF1FAAFF9FFB72FF1C" box="[1154,1224,116,132]" name="Myanmar" pageId="12" pageNumber="453">Myanmar</collectingCountry>
</paragraph>
<caption id="DF046622A70BFFEF1BBFF999FB65F928" ID-DOI="http://doi.org/10.5281/zenodo.4665557" ID-Zenodo-Dep="4665557" httpUri="https://zenodo.org/record/4665557/files/figure.png" pageId="12" pageNumber="453" startId="12.[151,162,1650,1667]" targetBox="[150,1242,258,1609]" targetPageId="12">
<paragraph id="8BC436AAA70BFFEF1BBFF999FB65F928" blockId="12.[151,1247,1650,1713]" pageId="12" pageNumber="453">
FIG. 4. — Comparison between the cladogram obtained by
<bibRefCitation id="EFEA4B5BA70BFFEF19BBF999FCADF91B" author="EGI N. &amp; HOLROYD P. A. &amp; AUNG NAING SOE &amp; TAKAI M. &amp; CIOCHON R. L." box="[659,791,1650,1667]" pageId="12" pageNumber="453" pagination="337 - 358" refId="ref9902" refString="EGI N., HOLROYD P. A., AUNG NAING SOE, TAKAI M. &amp; CIOCHON R. L. 2005. - Proviverrine hyaenodontids (Creodonta: Mammalia) from the Eocene of Myanmar and a phylogenetic analysis of the Proviverrines from the Para-Tethys area. Journal of Systematic Palaeontology 3: 337 - 358." type="journal article" year="2005">
Egi
<emphasis id="B90FEAB8A70BFFEF199CF999FD62F91B" box="[692,728,1650,1667]" italics="true" pageId="12" pageNumber="453">et al</emphasis>
. (2005)
</bibRefCitation>
(
<emphasis id="B90FEAB8A70BFFEF180BF999FC8AF91B" bold="true" box="[803,816,1650,1667]" pageId="12" pageNumber="453">A</emphasis>
) and that proposed here (
<emphasis id="B90FEAB8A70BFFEF1F38F999FBA7F91B" bold="true" box="[1040,1053,1650,1667]" pageId="12" pageNumber="453">B</emphasis>
). The position of
<taxonomicName id="4C7B4D29A70BFFEF1F9CF999FB60F91B" authorityName="Van Valen" authorityYear="1965" box="[1204,1242,1650,1667]" class="Mammalia" family="Hyaenodontidae" genus="Arfia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="12" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="B90FEAB8A70BFFEF1F9CF999FB60F91B" box="[1204,1242,1650,1667]" italics="true" pageId="12" pageNumber="453">Arfia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B4D29A70BFFEF1BBFF963FF42F901" authorityName="Cope" authorityYear="1874" box="[151,248,1672,1689]" class="Mammalia" family="Hyaenodontidae" genus="Prototomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="12" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="B90FEAB8A70BFFEF1BBFF963FF42F901" box="[151,248,1672,1689]" italics="true" pageId="12" pageNumber="453">Prototomus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7B4D29A70BFFEF1A0FF963FE23F901" authorityName="Matthew &amp; Granger" authorityYear="1915" box="[295,409,1672,1689]" class="Mammalia" family="Hyaenodontidae" genus="Prolimnocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="12" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="B90FEAB8A70BFFEF1A0FF963FE23F901" box="[295,409,1672,1689]" italics="true" pageId="12" pageNumber="453">Prolimnocyon</emphasis>
</taxonomicName>
as ancestor of
<taxonomicName id="4C7B4D29A70BFFEF1930F963FD84F901" authorityName="Van Valen" authorityYear="1965" box="[536,574,1672,1689]" class="Mammalia" family="Hyaenodontidae" genus="Arfia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="12" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="B90FEAB8A70BFFEF1930F963FD84F901" box="[536,574,1672,1689]" italics="true" pageId="12" pageNumber="453">Arfia</emphasis>
</taxonomicName>
-like South Asian proviverrine-Hyainailourinae,
<taxonomicName id="4C7B4D29A70BFFEF18E8F963FB9BF901" authorityName="Cope" authorityYear="1874" box="[960,1057,1672,1689]" class="Mammalia" family="Hyaenodontidae" genus="Prototomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="12" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="B90FEAB8A70BFFEF18E8F963FB9BF901" box="[960,1057,1672,1689]" italics="true" pageId="12" pageNumber="453">Prototomus</emphasis>
</taxonomicName>
-like
<taxonomicName id="4C7B4D29A70BFFEF1F61F962FB03F902" authorityName="Schlosser" authorityYear="1886" box="[1097,1209,1673,1690]" class="Mammalia" family="Hyaenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="12" pageNumber="453" phylum="Chordata" rank="subFamily" subFamily="Proviverrinae">Proviverrinae</taxonomicName>
, and Limnocyoninae,respectively, illustrate the current status of these genera.Dotted line represents imprecise relationships of Masrasec-
</paragraph>
</caption>
<paragraph id="8BC436AAA70BFFEF1BBFF95DFE3CF95F" blockId="12.[151,390,1718,1735]" box="[151,390,1718,1735]" pageId="12" pageNumber="453">
<emphasis id="B90FEAB8A70BFFEF1BBFF95DFF14F95F" box="[151,174,1718,1735]" italics="true" pageId="12" pageNumber="453">tor</emphasis>
. See text for explanation.
</paragraph>
<paragraph id="8BC436AAA70AFFEE1B86FF98FEA2FF1D" blockId="13.[174,280,115,133]" box="[174,280,115,133]" pageId="13" pageNumber="454">
Peigné S.
<emphasis id="B90FEAB8A70AFFEE1BDCFF9FFEA2FF1D" box="[244,280,116,133]" italics="true" pageId="13" pageNumber="454">et al.</emphasis>
</paragraph>
<paragraph id="8BC436AAA70AFFEE1BBFFF15FDF7FD0D" blockId="13.[149,686,254,692]" pageId="13" pageNumber="454">and they are lingually flanked by a small conule. The metaconule is transversely more elongated than the paraconule, and it is located on the distal rim of the tooth, just lingual to the metacone from which it is separated by a small notch. The paraconule is located on the mesial rim of the preprotocrista. The trigon basin is wide and deep, especially distally. The protocone is a large, triangular transverselydeveloped cusp. The cingulum is poorly developed mesially to the paracone-paraconule and around the protocone; it is slightly more developed labially and on the parastylar shelf; it is completely absent at the level of the metacone-metaconule.</paragraph>
<paragraph id="8BC436AAA70AFFEE1B86FD72FDE8FD2C" blockId="13.[149,686,254,692]" box="[174,594,665,692]" pageId="13" pageNumber="454">Measurements: M3 L = 9.5; W = 18.7.</paragraph>
<paragraph id="8BC436AAA70AFFEE1BBFFD31FE50FD69" blockId="13.[149,686,730,1706]" box="[151,490,730,756]" pageId="13" pageNumber="454">COMPARISONS AND DISCUSSION</paragraph>
<paragraph id="8BC436AAA70AFFEE1BBFFD12FE42FA54" blockId="13.[149,686,730,1706]" pageId="13" pageNumber="454">
Although there is a great size difference between NMMTmk-500 and NMMTdg-801 (assigned here to
<taxonomicName id="4C7B4D29A70AFFEE1B9AFCD2FEABFCCA" box="[178,273,824,850]" class="Mammalia" family="Hyaenodontidae" genus="Kyawdawia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="13" pageNumber="454" phylum="Chordata" rank="species" species="lupina">
<emphasis id="B90FEAB8A70AFFEE1B9AFCD2FEABFCCA" box="[178,273,824,850]" italics="true" pageId="13" pageNumber="454">K. lupina</emphasis>
</taxonomicName>
, see above) it is less than to the
<typeStatus id="54C08808A70AFFEE1978FCD3FD17FCCB" box="[592,685,824,851]" pageId="13" pageNumber="454" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7B4D29A70AFFEE1B9AFCB3FEA8FCE9" box="[178,274,855,881]" class="Mammalia" family="Hyaenodontidae" genus="Kyawdawia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="13" pageNumber="454" phylum="Chordata" rank="species" species="lupina">
<emphasis id="B90FEAB8A70AFFEE1B9AFCB3FEA8FCE9" box="[178,274,855,881]" italics="true" pageId="13" pageNumber="454">K. lupina</emphasis>
</taxonomicName>
. The great size of NMM Tmk-500 may therefore be insignificant. In addition to the size, however, proportional and morphological differences indicate that this specimen probably represents a different species. Compared to
<taxonomicName id="4C7B4D29A70AFFEE1AC7FC3CFD16FC69" authority=", NMM" authorityName="NMM" box="[495,684,982,1009]" class="Mammalia" family="Hyaenodontidae" genus="Kyawdawia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="13" pageNumber="454" phylum="Chordata" rank="species" species="lupina">
<emphasis id="B90FEAB8A70AFFEE1AC7FC3CFDEEFC68" box="[495,596,982,1008]" italics="true" pageId="13" pageNumber="454">K. lupina</emphasis>
, NMM
</taxonomicName>
Tmk-500 has a less reduced metacone compared to the paracone and, more or less correlatively, a lesser fusion between the bases of these cusps. It also possesses a shorter parastyle, and a greater length, resulting in part from a less reduced and more distally prominent metacone and from a more open angle (at the parastylar extremity) between the transverse mesial and the distolabial faces. Only the size is a significant difference of NMM Tmk-500 to
<taxonomicName id="4C7B4D29A70AFFEE1977FB18FF5EFAB5" class="Mammalia" family="Hyaenodontidae" genus="Yarshea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="13" pageNumber="454" phylum="Chordata" rank="species" species="cruenta">
<emphasis id="B90FEAB8A70AFFEE1977FB18FF5EFAB5" italics="true" pageId="13" pageNumber="454">Yarshea cruenta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D29A70AFFEE1A36FAF8FE11FAB6" authorityName="Schlosser" authorityYear="1886" box="[286,427,1299,1326]" class="Mammalia" family="Hyaenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="454" phylum="Chordata" rank="subFamily" subFamily="Proviverrinae">Proviverrinae</taxonomicName>
indet. from Pondaung. The distinction is even greater with
<taxonomicName id="4C7B4D29A70AFFEE1AD5FAD8FD1FFAD5" authorityName=", Kumar" authorityYear="1992" box="[509,677,1331,1357]" class="Mammalia" genus="Paratritemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="13" pageNumber="454" phylum="Chordata" rank="genus">
<emphasis id="B90FEAB8A70AFFEE1AD5FAD8FD1FFAD5" box="[509,677,1331,1357]" italics="true" pageId="13" pageNumber="454">Paratritemnodon</emphasis>
</taxonomicName>
. Besides a much larger size, NMM Tmk-500 differs in lacking a distal cingulum and in the retention of a metaconule, well developed metacone and protocone, and a postmetacrista.
</paragraph>
<paragraph id="8BC436AAA70AFFEE1B86FA3AFD16F932" blockId="13.[149,686,730,1706]" pageId="13" pageNumber="454">Because we would need more material and a better knowledge of the morphological variations in these Asian creodonts, we refrain from naming this isolated tooth though it possibly represents a new species.If so, it would reveal an even richer creodont fauna from Pondaung than previously thought and would further support the significance of Pondaung</paragraph>
<paragraph id="8BC436AAA70AFFEE1BBFF944FDE7F952" blockId="13.[151,605,1711,1738]" box="[151,605,1711,1738]" pageId="13" pageNumber="454">
in the evolution of the
<taxonomicName id="4C7B4D29A70AFFEE1AB8F944FDBEF952" authorityName="Cope" authorityYear="1875" box="[400,516,1711,1738]" class="Mammalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="13" pageNumber="454" phylum="Chordata" rank="order">Creodonta</taxonomicName>
in Asia.
</paragraph>
</subSubSection>
</treatment>
</document>