325 lines
43 KiB
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325 lines
43 KiB
XML
<document id="9B2D1690E8315888C891721CB16D536E" ID-DOI="10.11646/zootaxa.3847.1.1" ID-GBIF-Dataset="774be4c8-f95f-4d06-a555-dbe3f1aa2f21" ID-ISSN="1175-5326" ID-Zenodo-Dep="286815" ID-ZooBank="C7BDD136-2686-4049-B395-684797B26406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460469621688" checkinUser="plazi" docAuthor="Van Der Geer, Alexandra A. E." docDate="2014" docId="03D287A2FD1F371CFEF7FF22FD33FE15" docLanguage="en" docName="zt03847p032.pdf" docOrigin="Zootaxa 3847 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoplitomeryx devosi Geer, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="FFEBFFDAFD093704FE60FFB5FF9CFFC5" masterDocTitle="Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="23" updateTime="1698305463322" updateUser="plazi">
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<mods:title id="E81E30461493E1056D7F7DA2C39AA795">Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species</mods:title>
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<mods:namePart id="A59F8DDC582E6F110D1EA04C0A1B9568">Van Der Geer, Alexandra A. E.</mods:namePart>
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<mods:title id="6DB1832F3538A1A03A61ED88A1D83837">Zootaxa</mods:title>
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<mods:date id="14604922402FC48C987B94486C2E6BC3">2014</mods:date>
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<mods:number id="BCB0AEB800491BBBA681FE2CAABC5202">3847</mods:number>
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<treatment id="03D287A2FD1F371CFEF7FF22FD33FE15" ID-DOI="http://doi.org/10.5281/zenodo.5630050" ID-GBIF-Taxon="127668898" ID-Zenodo-Dep="5630050" LSID="urn:lsid:plazi:treatment:03D287A2FD1F371CFEF7FF22FD33FE15" httpUri="http://treatment.plazi.org/id/03D287A2FD1F371CFEF7FF22FD33FE15" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
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<subSubSection id="C361653FFD1F3712FEF7FF22FE97FF16" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BC436B4FD1F3712FEF7FF22FE68FF77" blockId="22.[151,500,151,211]" box="[151,500,151,178]" pageId="22" pageNumber="23">
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<heading id="D08C81D8FD1F3712FEF7FF22FE68FF77" bold="true" box="[151,500,151,178]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="B90FEAA6FD1F3712FEF7FF22FE68FF77" bold="true" box="[151,500,151,178]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FF22FE13FF74" box="[151,399,151,177]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="devosi" status="sp. nov.">
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<emphasis id="B90FEAA6FD1F3712FEF7FF22FE13FF74" bold="true" box="[151,399,151,177]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx devosi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23C57DDFD1F3712FFF6FF2DFE68FF77" box="[406,500,152,178]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FF0FFE97FF16" blockId="22.[151,500,151,211]" box="[151,267,186,211]" pageId="22" pageNumber="23">
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<figureCitation id="13402A31FD1F3712FEF7FF0FFF4BFF16" box="[151,215,186,211]" captionStart="FIGURE 7" captionStartId="23.[151,250,1887,1909]" captionTargetBox="[239,1344,201,1864]" captionTargetId="figure@23.[220,1375,193,1881]" captionTargetPageId="23" captionText="FIGURE 7. Holotypes of three new Hoplitomeryx species from Gargano, South Italy (Late Miocene; Middle or Late Turolian, MN 12 - 13). A – E Hoplitomeryx devosi sp. n., left metacarpal RGM 178.517; F – I Hoplitomeryx macpheei sp. n., left metacarpal RGM 260.918; J – M Hoplitomeryx kriegsmani sp. n., left metacarpal RGM 178.516. A, F, J proximal view. B, G, K dorsal view. C, H, L palmar view. D, I, M medial view. E distal view. Note that the sideward and backward bending of RGM 178.516 is a post-mortem defect." httpUri="https://zenodo.org/record/286822/files/figure.png" pageId="22" pageNumber="23">Fig. 7</figureCitation>
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A–E
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</paragraph>
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</subSubSection>
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<subSubSection id="C361653FFD1F3712FEF7FEB7FB79FD8E" pageId="22" pageNumber="23" type="reference_group">
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<paragraph id="8BC436B4FD1F3712FEF7FEB7FE5FFEF2" blockId="22.[151,1437,258,587]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FEB7FD8EFEDD" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,530,258,280]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="matthei">
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<emphasis id="B90FEAA6FD1F3712FEF7FEB7FEE9FEDD" box="[151,373,258,280]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx matthei</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FF1BFEB7FD8EFEDD" author="Leinders" box="[379,530,258,280]" pageId="22" pageNumber="23" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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</taxonomicName>
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—
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<bibRefCitation id="EFEA4B45FD1F3712FC4CFEB7FD23FEDD" author="Leinders" box="[556,703,258,280]" pageId="22" pageNumber="23" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders 1984</bibRefCitation>
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(partim): p. 3, fig. 1 [RGM 261.101];
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<bibRefCitation id="EFEA4B45FD1F3712FA28FEB6FAA7FEDD" author="Mazza" box="[1096,1339,258,280]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
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(partim): p. 1320 [RGM 261.102].
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FEF5FBE4FE93" blockId="22.[151,1437,258,587]" box="[151,1144,320,342]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FEF5FE5EFE93" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,450,320,342]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="B90FEAA6FD1F3712FEF7FEF5FEBEFE93" box="[151,290,320,342]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FF48FEF5FE5EFE93" author="Leinders" box="[296,450,320,342]" pageId="22" pageNumber="23" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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</taxonomicName>
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, ear region
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<typeStatus id="54C08816FD1F3712FC59FEF4FDF9FE93" box="[569,613,321,342]" pageId="22" pageNumber="23">type</typeStatus>
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V—Leinders 1984: p. 34, 35, pl. 7 [RGM 261.102].
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FEEBFB1DFEB1" blockId="22.[151,1437,258,587]" box="[151,1153,350,372]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FEEBFE5EFEB1" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,450,350,372]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="B90FEAA6FD1F3712FEF7FEEBFEBEFEB1" box="[151,290,350,372]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FF48FEEBFE5EFEB1" author="Leinders" box="[296,450,350,372]" pageId="22" pageNumber="23" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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</taxonomicName>
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, ear region
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<typeStatus id="54C08816FD1F3712FC59FEEAFDF9FEB1" box="[569,613,351,372]" pageId="22" pageNumber="23">type</typeStatus>
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IV—Leinders 1984: p. 31, 35, pl. 6 [RGM 261.096].
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FEC8FAF8FE56" blockId="22.[151,1437,258,587]" box="[151,1380,381,403]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FEC8FE5EFE56" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,450,381,403]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="B90FEAA6FD1F3712FEF7FEC8FEBEFE56" box="[151,290,381,403]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FF48FEC8FE5EFE56" author="Leinders" box="[296,450,381,403]" pageId="22" pageNumber="23" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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</taxonomicName>
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, size 1—Van der Geer 2005: p. 330, 333; Van der Geer 2008 (partim): p. 153, 154, fig. 4, 5.
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FE29FC52FE77" blockId="22.[151,1437,258,587]" box="[151,974,412,434]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FE29FE5EFE77" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,450,412,434]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="B90FEAA6FD1F3712FEF7FE29FEBEFE77" box="[151,290,412,434]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FF48FE29FE5EFE77" author="Leinders" box="[296,450,412,434]" pageId="22" pageNumber="23" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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</taxonomicName>
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, size 2—Van der Geer 2008 (partim): p. 153, fig. 4.
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FE0FFCD9FE2A" blockId="22.[151,1437,258,587]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FE0FFD1BFE15" authority="Mazza & Rustioni, 2011" authorityName="Mazza & Rustioni" authorityYear="2011" box="[151,647,442,464]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="apulicus">
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<emphasis id="B90FEAA6FD1F3712FEF7FE0FFE1DFE15" box="[151,385,442,464]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx apulicus</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FFE9FE0EFD1BFE15" author="Mazza" box="[393,647,442,464]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni, 2011</bibRefCitation>
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</taxonomicName>
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—
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<bibRefCitation id="EFEA4B45FD1F3712FCC1FE0FFC06FE15" author="Mazza" box="[673,922,442,464]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
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(partim): p. 1318, fig. 3, table 2 [RGM 178.445, RGM 178.656, RGM 260.940, RGM 260.966, RGM 425.234].
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FE4DFCD9FDE9" blockId="22.[151,1437,258,587]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FE4DFD17FDCB" authority="Mazza & Rustioni, 2011" authorityName="Mazza & Rustioni" authorityYear="2011" box="[151,651,504,526]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="falcidens">
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<emphasis id="B90FEAA6FD1F3712FEF7FE4DFE1AFDCB" box="[151,390,504,526]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx falcidens</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FFEEFE4CFD17FDCB" author="Mazza" box="[398,651,504,526]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni, 2011</bibRefCitation>
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||
</taxonomicName>
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||
—
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||
<bibRefCitation id="EFEA4B45FD1F3712FCC5FE4DFC01FDCB" author="Mazza" box="[677,925,504,526]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
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(partim): p. 1312, fig. 1, table 2 [RGM 260.941, RGM 261.132, RGM 261.133, RGM 261.447, RGM 425.201].
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</paragraph>
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<paragraph id="8BC436B4FD1F3712FEF7FD80FB79FD8E" blockId="22.[151,1437,258,587]" box="[151,1253,565,587]" pageId="22" pageNumber="23">
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<taxonomicName id="4C7B4D37FD1F3712FEF7FD80FDEAFD8E" authority="Mazza & Rustioni, 2011" authorityName="Mazza & Rustioni" authorityYear="2011" box="[151,630,565,587]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="minutus">
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<emphasis id="B90FEAA6FD1F3712FEF7FD80FEE4FD8E" box="[151,376,565,587]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx minutus</emphasis>
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<bibRefCitation id="EFEA4B45FD1F3712FF1FFD83FDEAFD8E" author="Mazza" box="[383,630,565,587]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni, 2011</bibRefCitation>
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</taxonomicName>
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—
|
||
<bibRefCitation id="EFEA4B45FD1F3712FCF0FD80FC1EFD8E" author="Mazza" box="[656,898,565,587]" pageId="22" pageNumber="23" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
|
||
(partim): p. 1306 [RGM 261.147].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD1F3712FEF7FDC2FCF2FCAD" pageId="22" pageNumber="23" type="materials_examined">
|
||
<paragraph id="8BC436B4FD1F3712FEF7FDC2FDEFFD55" blockId="22.[151,1437,631,2025]" box="[151,627,631,656]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEF7FDC2FE97FD55" bold="true" box="[151,267,631,656]" pageId="22" pageNumber="23">
|
||
<typeStatus id="54C08816FD1F3712FEF7FDC2FE9AFD55" box="[151,262,631,656]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Left metacarpal
|
||
<collectionCode id="ED6AAE71FD1F3712FFA9FDCDFD95FD55" box="[457,521,632,656]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
178.517.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7FD29FF66FC80" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FD29FEDAFD70" bold="true" box="[199,326,668,693]" pageId="22" pageNumber="23">
|
||
<typeStatus id="54C08816FD1F3712FEA7FD29FEDDFD70" box="[199,321,668,693]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Distal humerus fragment
|
||
<collectionCode id="ED6AAE71FD1F3712FC16FD29FD2AFD71" box="[630,694,668,692]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
443.118, radius
|
||
<collectionCode id="ED6AAE71FD1F3712FD19FD29FC25FD71" box="[889,953,668,692]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
260.860, femur
|
||
<collectionCode id="ED6AAE71FD1F3712FA1BFD29FB27FD71" box="[1147,1211,668,692]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
425.314 (
|
||
<figureCitation id="13402A31FD1F3712FB52FD29FAE4FD70" box="[1330,1400,668,693]" captionStart="FIGURE 5" captionStartId="11.[151,250,1725,1747]" captionTargetBox="[290,1308,202,1695]" captionTargetId="figure@11.[276,1315,193,1705]" captionTargetPageId="11" captionText="FIGURE 5. Postcranial elements of Hoplitomeryx. A, first posterior phalanx (RGM 425.269; San Giovannino) showing a deerlike morphology: abaxial view; B, same specimen, showing triangular sesamoid facets but no ligament attachment areas: volar view; C, first anterior phalanx (RGM 178.503; San Giovannino): abaxial view; D, same specimen, showing pronounced ligament attachments in the form of ridges but no sesamoid facet: volar view; E, posterior second phalanx (RGM 261.251; San Giovannino): abaxial view; F, anterior second phalanx (RGM 260.937; San Giovannino): axial view; G, third phalanx (RGM 178.683; Trefossi 2 A): abaxial view; H, third phalanx (RGM 260.877; San Giovannino): axial view; I, third phalanx (RGM 261.531; Nazario 4): abaxial view; J, third phalanx (RGM 425.386; Falcone 2 B); K, right femur (RGM 425.314; San Giovannino): anterior view; L, left femur (RGM 425.207; Nazario 4): anterior view; M, right femur (RGM 425.245; San Giovannino): anterior view; N, left patella (RGM 425.246; San Giovannino): anterior (dorsal) view; O, same specimen: lateral view; P, same specimen, showing the hook-like extension at the medial side: inner view." httpUri="https://zenodo.org/record/286820/files/figure.png" pageId="22" pageNumber="23">Fig. 5</figureCitation>
|
||
K), tibia
|
||
<collectionCode id="ED6AAE71FD1F3712FEB0FD75FE8CFD1D" box="[208,272,704,728]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
425.285, astragal
|
||
<collectionCode id="ED6AAE71FD1F3712FF80FD75FDBCFD1D" box="[480,544,704,728]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
215.234, calcaneum
|
||
<collectionCode id="ED6AAE71FD1F3712FD71FD75FCCDFD1D" box="[785,849,704,728]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
425.253, posterior second phalanges
|
||
<collectionCode id="ED6AAE71FD1F3712FA9AFD75FAA6FD1D" box="[1274,1338,704,728]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
261.251 (
|
||
<figureCitation id="13402A31FD1F3712FEFFFD51FF7AFD38" box="[159,230,740,765]" captionStart="FIGURE 5" captionStartId="11.[151,250,1725,1747]" captionTargetBox="[290,1308,202,1695]" captionTargetId="figure@11.[276,1315,193,1705]" captionTargetPageId="11" captionText="FIGURE 5. Postcranial elements of Hoplitomeryx. A, first posterior phalanx (RGM 425.269; San Giovannino) showing a deerlike morphology: abaxial view; B, same specimen, showing triangular sesamoid facets but no ligament attachment areas: volar view; C, first anterior phalanx (RGM 178.503; San Giovannino): abaxial view; D, same specimen, showing pronounced ligament attachments in the form of ridges but no sesamoid facet: volar view; E, posterior second phalanx (RGM 261.251; San Giovannino): abaxial view; F, anterior second phalanx (RGM 260.937; San Giovannino): axial view; G, third phalanx (RGM 178.683; Trefossi 2 A): abaxial view; H, third phalanx (RGM 260.877; San Giovannino): axial view; I, third phalanx (RGM 261.531; Nazario 4): abaxial view; J, third phalanx (RGM 425.386; Falcone 2 B); K, right femur (RGM 425.314; San Giovannino): anterior view; L, left femur (RGM 425.207; Nazario 4): anterior view; M, right femur (RGM 425.245; San Giovannino): anterior view; N, left patella (RGM 425.246; San Giovannino): anterior (dorsal) view; O, same specimen: lateral view; P, same specimen, showing the hook-like extension at the medial side: inner view." httpUri="https://zenodo.org/record/286820/files/figure.png" pageId="22" pageNumber="23">Fig. 5</figureCitation>
|
||
E) and
|
||
<collectionCode id="ED6AAE71FD1F3712FF22FD51FE1EFD39" box="[322,386,740,764]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
261.254, pelvis fragment (acetabulum)
|
||
<collectionCode id="ED6AAE71FD1F3712FD3DFD51FC01FD39" box="[861,925,740,764]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
261.221, scapula fragment (glenoid)
|
||
<collectionCode id="ED6AAE71FD1F3712FB3CFD51FA00FD39" box="[1372,1436,740,764]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
179.317, mandible
|
||
<collectionCode id="ED6AAE71FD1F3712FF14FCBDFE28FCE5" box="[372,436,776,800]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
260.940, maxilla fragments
|
||
<collectionCode id="ED6AAE71FD1F3712FC9FFCBDFCA3FCE5" box="[767,831,776,800]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
260.941 and
|
||
<collectionCode id="ED6AAE71FD1F3712FDBFFCBDFB83FCE5" box="[991,1055,776,800]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
425.201, orbital horn core
|
||
<collectionCode id="ED6AAE71FD1F3712FB3CFCBDFA00FCE5" box="[1372,1436,776,800]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="23">RGM</collectionCode>
|
||
260.926.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7FCFAFCF2FCAD" blockId="22.[151,1437,631,2025]" box="[199,878,847,872]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FCFAFE25FCAD" bold="true" box="[199,441,847,872]" pageId="22" pageNumber="23">Referred specimens.</emphasis>
|
||
See Appendix III, fig.
|
||
<date id="FFC51074FD1F3712FCDCFCE5FD49FCAD" box="[700,725,848,872]" pageId="22" pageNumber="23">4I</date>
|
||
, J and fig.
|
||
<date id="FFC51074FD1F3712FD2FFCFAFCF2FCAD" box="[847,878,847,872]" pageId="22" pageNumber="23">5I.</date>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD1F3712FEA7FCC1FDD5FB31" pageId="22" pageNumber="23" type="diagnosis">
|
||
<paragraph id="8BC436B4FD1F3712FEA7FCC1FD05FBD8" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FCC1FEA6FC48" bold="true" box="[199,314,884,909]" pageId="22" pageNumber="23">Diagnosis</emphasis>
|
||
. A small-sized, very robust hoplitomerycid. Estimated body mass 21.0 kg. The trochlea of the astragal is either non-parallel sided or almost parallel. The tibia is spirally winded. The fusion between ulna and radius is extensive and includes the part distally of the spatium interosseum. The ulna leaves a trace on the radius in the form of a ridge. The radial facet on the distal radius extends further palmar than the intermedial facet and is deeper and broader than the intermedial facet.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7FB92FA90FB85" blockId="22.[151,1437,631,2025]" box="[199,1292,1063,1088]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FB92FE52FB85" bold="true" box="[199,462,1063,1088]" pageId="22" pageNumber="23">Differential diagnosis.</emphasis>
|
||
About half the body size of
|
||
<taxonomicName id="4C7B4D37FD1F3712FD6DFB9DFB9EFB85" box="[781,1026,1064,1088]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="matthei">
|
||
<emphasis id="B90FEAA6FD1F3712FD6DFB9DFB9EFB85" box="[781,1026,1064,1088]" italics="true" pageId="22" pageNumber="23">Hoplitomeryx matthei</emphasis>
|
||
</taxonomicName>
|
||
and much more robust.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7FBF9FE0AFB68" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FBF9FE28FBA0" bold="true" box="[199,436,1100,1125]" pageId="22" pageNumber="23">Derivation of name.</emphasis>
|
||
Named after John de Vos, former curator of the Dubois collection, Naturalis Biodiversity Center, Leiden for his contribution to the knowledge of fossil insular mammals, in particular insular deer from the Mediterranean islands.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7FB02FDD5FB31" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FB02FD81FB15" bold="true" box="[199,541,1207,1232]" pageId="22" pageNumber="23">Preservation and deposition.</emphasis>
|
||
Naturalis Biodiversity Center, Leiden,
|
||
<collectingCountry id="F36C7624FD1F3712FDB7FB02FB14FB15" box="[983,1160,1207,1232]" name="Netherlands" pageId="22" pageNumber="23">the Netherlands</collectingCountry>
|
||
(formerly Rijksmuseum van Geologie en Mineralogie (RGM)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD1F3712FEA7FB4AFA0FF9D0" pageId="22" pageNumber="23" type="materials_examined">
|
||
<paragraph id="8BC436B4FD1F3712FEA7FB4AFC71FAA5" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FB4AFD9EFADD" bold="true" box="[199,514,1279,1304]" pageId="22" pageNumber="23">
|
||
<typeStatus id="54C08816FD1F3712FEA7FB4AFE9EFADD" box="[199,258,1279,1304]" pageId="22" pageNumber="23">Type</typeStatus>
|
||
locality and horizon.
|
||
</emphasis>
|
||
Late Miocene (Middle or Late Turolian, MN12-13) fissure filling with code San Giovannino in an abandoned limestone quarry near the farm of San Giovannino south of the provincial road between Poggio Imperiale and Apricena (Province of Foggia, Apulia,
|
||
<collectingCountry id="F36C7624FD1F3712FDC9FAF2FC7DFAA5" box="[937,993,1351,1376]" name="Italy" pageId="22" pageNumber="23">Italy</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7FAD9FA0FF9D0" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7FAD9FE04FA40" bold="true" box="[199,408,1388,1413]" pageId="22" pageNumber="23">Studied localities.</emphasis>
|
||
Fissure fillings with codes Biancone 2, Chirò 2, 2N, 2S, 3, 5, 5A, 7, 10A, 10B, 13, 14, 14B, 18, 26, 27, 28A, 29, 30, D1 and D3, Fina D, H, K and N, Gervasio, Nazario 2A, 3 and 4, Pizzicoli 12, Posticchia 1B, San Giovannino and Trefossi 2A. All localities are located in the north-western portion of the Gargano Peninsula, Apulia, south-eastern
|
||
<collectingCountry id="F36C7624FD1F3712FC6CFA62FDDDFA35" box="[524,577,1495,1520]" name="Italy" pageId="22" pageNumber="23">Italy</collectingCountry>
|
||
. Probable other locality. Fissure filling with code Chirò
|
||
<quantity id="4C839B51FD1F3712FAA2FA6DFB61FA35" box="[1218,1277,1495,1520]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="22" pageNumber="23" unit="in" value="24.0">24 in</quantity>
|
||
the limestone quarry Chirò along the provincial road between Poggio Imperiale and Lessina (Province of Foggia, Apulia,
|
||
<collectingCountry id="F36C7624FD1F3712FB2FFA49FA1BF9D0" box="[1359,1415,1532,1557]" name="Italy" pageId="22" pageNumber="23">Italy</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD1F3712FEA7F9AAFB68F82C" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="8BC436B4FD1F3712FEA7F9AAFAA4F8B8" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7F9AAFD8BF9FD" bold="true" box="[199,535,1567,1592]" pageId="22" pageNumber="23">
|
||
Description of the
|
||
<typeStatus id="54C08816FD1F3712FFC9F9AAFD8EF9FD" box="[425,530,1567,1592]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
RGM 178.517 is a complete, left metacarpal with some splinters of the dorsal surface missing. Basically, it corresponds in morphology with that of
|
||
<taxonomicName id="4C7B4D37FD1F3712FDD0F9F1FB9CF99E" box="[944,1024,1604,1627]" class="Mammalia" family="Cervidae" genus="Cervus" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90FEAA6FD1F3712FDD0F9F1FB9CF99E" box="[944,1024,1604,1627]" italics="true" pageId="22" pageNumber="23">Cervus</emphasis>
|
||
</taxonomicName>
|
||
and accordingly, the medial facet is larger than the lateral facet on the proximal articulation. It is extremely massive and robust. The crest separating the medial and lateral proximal facet ends in the central fossa and runs more or less parasagitally. The crest forms a pronounced ridge between the two facets. The fossa is situated centrally and the contact area between the lateral and medial facet is minimal. The central fossa is in contact with the palmar surface at which point it is wide. The proximal tubercle for attachment of the carpal ligaments on the dorsal surface is pronounced. The scars for the lateral metacarpals are very pronounced. The palmar groove for the M. interosseus palmaris is clearly present up to about one-third above the distal end of the shaft. The distal end of the metacarpal is straight with the lateral and medial trochlea extending equally far. The dorsal profile of both epicondyls in distal view is cone-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD1F3712FEA7F832FB68F82C" blockId="22.[151,1437,631,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90FEAA6FD1F3712FEA7F832FEE1F865" bold="true" box="[199,381,1927,1952]" pageId="22" pageNumber="23">Measurements.</emphasis>
|
||
<typeStatus id="54C08816FD1F3712FFE6F832FE6FF865" box="[390,499,1927,1952]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||
: maximal length =
|
||
<quantity id="4C839B51FD1F3712FCB1F832FCA5F865" box="[721,825,1927,1952]" metricMagnitude="-2" metricUnit="m" metricValue="9.45" pageId="22" pageNumber="23" unit="mm" value="94.5">94.5 mm</quantity>
|
||
, proximal depth (DAPP) =
|
||
<quantity id="4C839B51FD1F3712FA19F832FB7CF865" box="[1145,1248,1927,1952]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="22" pageNumber="23" unit="mm" value="14.6">14.6 mm</quantity>
|
||
, proximal width (DTP) =
|
||
<quantity id="4C839B51FD1F3712FE9DF819FEFFF801" box="[253,355,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="22" pageNumber="23" unit="mm" value="23.2">23.2 mm</quantity>
|
||
, distal depth (DAPD) = 12.0 mm, distal width (DTD) =
|
||
<quantity id="4C839B51FD1F3712FDBDF819FBD8F801" box="[989,1092,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="22" pageNumber="23" unit="mm" value="21.8">21.8 mm</quantity>
|
||
. For measurements of referred specimens, see Appendix III (linear measurements) and Appendix I and II (body mass estimations).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF04663CFD1E3713FEF7F8EAFEF8F835" httpUri="https://zenodo.org/record/286822/files/figure.png" pageId="23" pageNumber="24" targetBox="[239,1344,201,1864]" targetPageId="23">
|
||
<paragraph id="8BC436B4FD1E3713FEF7F8EAFEF8F835" blockId="23.[151,1437,1887,2032]" pageId="23" pageNumber="24">
|
||
<emphasis id="B90FEAA6FD1E3713FEF7F8EAFE91F8B1" bold="true" box="[151,269,1887,1909]" pageId="23" pageNumber="24">FIGURE 7</emphasis>
|
||
. Holotypes of three new
|
||
<taxonomicName id="4C7B4D37FD1E3713FC6CF8EAFD0BF8B0" box="[524,663,1887,1909]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90FEAA6FD1E3713FC6CF8EAFD0BF8B0" box="[524,663,1887,1909]" italics="true" pageId="23" pageNumber="24">Hoplitomeryx</emphasis>
|
||
</taxonomicName>
|
||
species from Gargano, South Italy (Late Miocene; Middle or Late Turolian, MN12-13). A–E
|
||
<taxonomicName id="4C7B4D37FD1E3713FF24F8C8FD8AF856" box="[324,534,1917,1939]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="devosi" status="sp. nov.">
|
||
<emphasis id="B90FEAA6FD1E3713FF24F8C8FD8AF856" box="[324,534,1917,1939]" italics="true" pageId="23" pageNumber="24">Hoplitomeryx devosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAA6FD1E3713FC7CF8CBFDC9F856" bold="true" box="[540,597,1918,1939]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="A23C57DDFD1E3713FC7CF8CBFDC9F856" box="[540,597,1918,1939]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left metacarpal RGM 178.517; F–I
|
||
<taxonomicName id="4C7B4D37FD1E3713FDA7F8C8FB26F856" box="[967,1210,1917,1939]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="macpheei" status="sp. nov.">
|
||
<emphasis id="B90FEAA6FD1E3713FDA7F8C8FB26F856" box="[967,1210,1917,1939]" italics="true" pageId="23" pageNumber="24">Hoplitomeryx macpheei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAA6FD1E3713FAA0F8CBFB64F856" bold="true" box="[1216,1272,1918,1939]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="A23C57DDFD1E3713FAA0F8CBFB64F856" box="[1216,1272,1918,1939]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left metacarpal RGM 260.918; J–M
|
||
<taxonomicName id="4C7B4D37FD1E3713FF08F829FDF5F877" box="[360,617,1948,1970]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="kriegsmani" status="sp. nov.">
|
||
<emphasis id="B90FEAA6FD1E3713FF08F829FDF5F877" box="[360,617,1948,1970]" italics="true" pageId="23" pageNumber="24">Hoplitomeryx kriegsmani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAA6FD1E3713FC0FF828FD3BF877" bold="true" box="[623,679,1949,1970]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="A23C57DDFD1E3713FC0FF828FD3BF877" box="[623,679,1949,1970]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left metacarpal RGM 178.516. A, F, J proximal view. B, G, K dorsal view. C, H, L palmar view. D, I, M medial view. E distal view. Note that the sideward and backward bending of RGM 178.516 is a post-mortem defect.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C361653FFD11371CFEA7FF22FD33FE15" pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph id="8BC436B4FD11371CFEA7FF22FD33FE15" blockId="24.[151,1436,151,464]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FF22FEA7FF6A" bold="true" box="[199,315,151,176]" pageId="24" pageNumber="25">Remarks.</emphasis>
|
||
San Giovannino falls within the younger faunal complex (no hamsters; see Introduction) and is one of the youngest localities. Skull fragment RGM 261.102 (ear region
|
||
<typeStatus id="54C08816FD11371CFDD0FF08FC7CFF10" box="[944,992,189,213]" pageId="24" pageNumber="25">type</typeStatus>
|
||
V in
|
||
<bibRefCitation id="EFEA4B45FD11371CFA47FF09FB41FF11" author="Leinders" box="[1063,1245,188,213]" pageId="24" pageNumber="25" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders (1984)</bibRefCitation>
|
||
from the quarry Gervasio) is smaller than the
|
||
<typeStatus id="54C08816FD11371CFF81FF55FD8DFF3D" box="[481,529,224,248]" pageId="24" pageNumber="25">type</typeStatus>
|
||
skull and differs from it by a circular foramen magnum and larger bullae. Based on occipital width, this skull fragment belongs to the smallest species. The configuration of the proximal articulation of the
|
||
<typeStatus id="54C08816FD11371CFF0EFE92FE4CFE85" box="[366,464,295,320]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
metacarpal was earlier described as morphotype 2 (Van der Geer 2005) and is also observed in the Tokunoshima-type of
|
||
<taxonomicName id="4C7B4D37FD11371CFC2DFEF9FACDFEA1" authority="(Matsumoto, 1926) (Matsumoto & Otsuka 2000)" authorityName="(Matsumoto, 1926) (Matsumoto & Otsuka" authorityYear="2000" box="[589,1361,332,357]" class="Mammalia" family="Cervidae" genus="Cervus" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="astylodon">
|
||
<emphasis id="B90FEAA6FD11371CFC2DFEF9FC88FEA1" box="[589,788,332,356]" italics="true" pageId="24" pageNumber="25">Cervus astylodon</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B45FD11371CFD45FEF9FC6DFEA0" author="Matsumoto" box="[805,1009,332,357]" pageId="24" pageNumber="25" refString="Matsumoto, H. (1926) On some new fossil Cervicorns from Kazusa and Liukiu. Scientific Report of Tohoku Imperial University, Series 2 (Geology), 10, 21 - 23." type="journal article" year="1926">Matsumoto, 1926</bibRefCitation>
|
||
) (
|
||
<bibRefCitation id="EFEA4B45FD11371CFA69FEF9FAD5FEA1" author="Matsumoto" box="[1033,1353,332,357]" pageId="24" pageNumber="25" refString="Matsumoto, Y. & Otsuka, H. (2000) Morphotypes of fossil deer (Cervus astylodon) and its miniaturization process. Tropics, 10, 145 - 154." type="journal article" year="2000">Matsumoto & Otsuka 2000</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B4D37FD11371CFB3FFEF9FE04FE4D" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cervidae" genus="Alces" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="alces">
|
||
<emphasis id="B90FEAA6FD11371CFB3FFEF9FF4DFE4D" italics="true" pageId="24" pageNumber="25">Alces alces</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B45FD11371CFE80FEDAFE0CFE4D" author="Linnaeus" box="[224,400,367,392]" pageId="24" pageNumber="25" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurence Salvius, Holmiae (= Stockholm), 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D37FD11371CFFB0FEC5FD57FE4D" authority="Scott, 1885" authorityName="Scott" authorityYear="1885" box="[464,715,367,392]" class="Mammalia" family="Cervidae" genus="Cervalces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90FEAA6FD11371CFFB0FEC5FDDCFE4D" box="[464,576,368,392]" italics="true" pageId="24" pageNumber="25">Cervalces</emphasis>
|
||
<bibRefCitation id="EFEA4B45FD11371CFC28FEC5FD57FE4D" author="Scott" box="[584,715,367,392]" pageId="24" pageNumber="25" refString="Scott, W. B. (1885) Cervalces americanus, a fossil moose, or elk, from the Quaternary of New Jersey. Proceedings of the Academy of Natural Sciences of Philadelphia, 37, 181 - 202." type="journal article" year="1885">Scott, 1885</bibRefCitation>
|
||
</taxonomicName>
|
||
. Maxillaries RGM 260.941 and RGM 425.201 do not belong to the same individual (contra
|
||
<bibRefCitation id="EFEA4B45FD11371CFFB5FE20FD74FE69" author="Mazza" box="[469,744,404,429]" pageId="24" pageNumber="25" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
|
||
), based on their different eruption patterns (this is confirmed by their different hypsodonty index, see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |