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<mods:title id="089333686CE190955CC4AF0A3D848BBD">Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species</mods:title>
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<mods:namePart id="8EC4D8B59F466E19C2ACDE039076DF7A">Van Der Geer, Alexandra A. E.</mods:namePart>
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<taxonomicName id="4C7B4D37FD1D3710FEF7FF22FEDCFF74" box="[151,320,151,177]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="20" pageNumber="14" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B90FEAA6FD1D3710FEF7FF22FEDCFF74" bold="true" box="[151,320,151,177]" italics="true" pageId="20" pageNumber="21">Scontromeryx</emphasis>
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sp. indet.
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<taxonomicName id="4C7B4D37FD1D3710FEF7FF0FFEBEFF15" box="[151,290,186,208]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B90FEAA6FD1D3710FEF7FF0FFEBEFF15" box="[151,290,186,208]" italics="true" pageId="20" pageNumber="21">Hoplitomeryx</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="EFEA4B45FD1D3710FF2CFF0FFE7EFF15" author="Leinders" box="[332,482,186,208]" pageId="20" pageNumber="21" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
<bibRefCitation id="EFEA4B45FD1D3710FF9CFF0FFD6DFF15" author="Mazza" box="[508,753,186,208]" pageId="20" pageNumber="21" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni 2011</bibRefCitation>
: p. 1326.
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD1D3710FEF7FF48FEBFFE63" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BC436B4FD1D3710FEF7FF48FB25FED3" blockId="20.[151,1436,253,746]" box="[151,1209,253,278]" pageId="20" pageNumber="21">
<emphasis id="B90FEAA6FD1D3710FEF7FF48FE9AFED3" bold="true" box="[151,262,253,278]" pageId="20" pageNumber="21">Material.</emphasis>
Horn cores SCT10, SCT28, SCT4, SCT213, SCT11 and an uncatalogued specimen.
</paragraph>
<paragraph id="8BC436B4FD1D3710FEA7FE94FA9FFEFF" blockId="20.[151,1436,253,746]" box="[199,1283,289,314]" pageId="20" pageNumber="21">
<emphasis id="B90FEAA6FD1D3710FEA7FE94FD80FEFF" bold="true" box="[199,540,289,314]" pageId="20" pageNumber="21">Preservation and deposition.</emphasis>
Soprintendenza Archeologica dellAbruzzo (Chieti, central
<collectingCountry id="F36C7624FD1D3710FADFFE94FB6BFEFF" box="[1215,1271,289,314]" name="Italy" pageId="20" pageNumber="21">Italy</collectingCountry>
).
</paragraph>
<paragraph id="8BC436B4FD1D3710FEA7FEF0FEBFFE63" blockId="20.[151,1436,253,746]" pageId="20" pageNumber="21">
<emphasis id="B90FEAA6FD1D3710FEA7FEF0FE57FE9B" bold="true" box="[199,459,325,350]" pageId="20" pageNumber="21">Locality and horizon.</emphasis>
Tortonian Scontrone Member of the Lithothamnium Limestone (
<bibRefCitation id="EFEA4B45FD1D3710FAD8FEF3FA0BFE9B" author="Patacca" box="[1208,1431,325,350]" pageId="20" pageNumber="21" refString="Patacca, E., Scandone, P. &amp; Mazza, P. (2008) Oligocene migration path for Apulia macromammals: the Central-Adriatic bridge. Bollettino della Societa Geologica Italiana, 127, 337 - 355." type="journal article" year="2008">
Patacca
<emphasis id="B90FEAA6FD1D3710FB78FEF3FACFFE98" box="[1304,1363,325,349]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2008
</bibRefCitation>
;
<geoCoordinate id="EE4F5073FD1D3710FEF7FEDCFEA3FE47" box="[151,319,361,386]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="41.75432">41°45'15.54''N</geoCoordinate>
,
<geoCoordinate id="EE4F5073FD1D3710FF2AFEDCFE6DFE44" box="[330,497,361,385]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="14.0369835">14°02'13.14''E</geoCoordinate>
), outskirts of Scontrone, southern border of the National Park of Abruzzi, LAquila, central
<collectingCountry id="F36C7624FD1D3710FE8AFE38FE83FE63" box="[234,287,397,422]" name="Italy" pageId="20" pageNumber="21">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD1D3710FEA7FE04FBF4FD2F" pageId="20" pageNumber="21" type="description">
<paragraph id="8BC436B4FD1D3710FEA7FE04FBF4FD2F" blockId="20.[151,1436,253,746]" pageId="20" pageNumber="21">
<emphasis id="B90FEAA6FD1D3710FEA7FE04FEC5FE0F" bold="true" box="[199,345,433,458]" pageId="20" pageNumber="21">Description.</emphasis>
SCT10 and SCT28 are described (
<bibRefCitation id="EFEA4B45FD1D3710FC80FE04FC6CFE0C" author="Mazza" box="[736,1008,433,458]" pageId="20" pageNumber="21" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni 2011</bibRefCitation>
) as
<typeStatus id="54C08816FD1D3710FA7DFE07FBD1FE0F" box="[1053,1101,434,458]" pageId="20" pageNumber="21">type</typeStatus>
I nasal horn cores, SCT4 and SCT213 as
<typeStatus id="54C08816FD1D3710FF7CFE63FED0FE2B" box="[284,332,470,494]" pageId="20" pageNumber="21">type</typeStatus>
II nasal horn cores and SCT11 and a not-catalogued specimen as
<typeStatus id="54C08816FD1D3710FA59FE63FBF5FE2B" box="[1081,1129,470,494]" pageId="20" pageNumber="21">type</typeStatus>
V orbital horn cores. Only SCT213 and SCT28 are figured (
<figureCitation id="13402A31FD1D3710FC6AFE4CFDD2FDD7" box="[522,590,505,530]" captionStart="FIGURE 6" captionStartId="12.[151,250,1824,1846]" captionTargetBox="[205,1378,205,1796]" captionTargetId="figure@12.[196,1392,193,1803]" captionTargetPageId="12" captionText="FIGURE 6. Postcranial elements of Hoplitomeryx. A, left tibia (RGM 335.882 + 425.328; San Giovannino): medial view; B, right tibia (RGM 425.144; Chiro D 1): anterior view. Astragals of different size in anterior (dorsal) view: C, left astragal (RGM 425.385; Falcone 2 B); D, left astragal (RGM 260.863; San Giovannino); E, right astragal (RGM 425.421; Fina N). F, left astragal (RGM 263.951; Gargano, unknown fissure): lateral view; G, left calcaneum (RGM 260.901; San Giovannino): anterior view; H, same specimen: posterior view; I, same specimen: medial view; J,; K, detail of distal right metatarsal (RGM 178.518; San Giovannino) showing the proximal extension of the dorsal gully into the (fused) cubonavicular." httpUri="https://zenodo.org/record/286821/files/figure.png" pageId="20" pageNumber="21">Fig. 6</figureCitation>
, IK). The nasal horn cores lack the typical continuation of the septum nasi in the basal part of the horn as can be seen in their fig. 6 and can therefore not reliably diagnosed as being nasal horn cores. The isolated orbital horn cores (
<typeStatus id="54C08816FD1D3710FCF2FDF4FD58FD9C" box="[658,708,577,601]" pageId="20" pageNumber="21">type</typeStatus>
V: SCT11 and an uncatalogued specimen) do not fit Leinders (1984) description either. One of them has been depicted in
<bibRefCitation id="EFEA4B45FD1D3710FD20FDD0FBAFFDBB" box="[832,1075,613,638]" pageId="20" pageNumber="21" refString="Rustioni, M., Mazza, P., Azzaroli, A., Boscagli, G., Cozzini, F., Di Vito, E., Masseti, M. &amp; Pisano, A. (1992) Miocene Vertebrate remains from Scontrone, National Park of Abruzzi, Central Italy. Rendiconti della Classe Scienze Fisiche, Matematiche e Naturali dell'Accademia dei Lincei, 9, 227 - 237." type="journal article">
Rustioni
<emphasis id="B90FEAA6FD1D3710FDC7FDD3FC46FDB8" box="[935,986,613,637]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
. (1992)
</bibRefCitation>
and is too large to fit Leinders (1984) horn core
<typeStatus id="54C08816FD1D3710FF39FD3FFE15FD67" box="[345,393,650,674]" pageId="20" pageNumber="21">type</typeStatus>
V and differs morphologically from the other horn core
<typeStatus id="54C08816FD1D3710FD9FFD3FFBA2FD67" box="[1023,1086,650,674]" pageId="20" pageNumber="21">types</typeStatus>
. The isolated orbital horn cores also differ from those in skull SCT17 (
<taxonomicName id="4C7B4D37FD1D3710FC04FD1BFD4EFD00" box="[612,722,686,709]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mazzai">
<emphasis id="B90FEAA6FD1D3710FC04FD1BFD4EFD00" box="[612,722,686,709]" italics="true" pageId="20" pageNumber="21">S. mazzai</emphasis>
</taxonomicName>
). The isolated horn cores cannot be associated to any of the described species and could be distal parts of either horn cores, antlers or proto-antlers.
</paragraph>
</subSubSection>
</treatment>
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