treatments-xml/data/03/D1/84/03D18442FFDCFF9B317CFF6E5C70FF7D.xml
2024-06-21 12:22:17 +02:00

403 lines
46 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D9BD3C8A9C3003330DE05F1D3E8C2220" ID-DOI="10.1093/zoolinnean/zlad115" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241094" ID-ZooBank="E6D9AF7E-F0AD-4656-B2F2-7FBFAA0312B3R" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716363659947" checkinUser="plazi" docAuthor="Cherman, Mariana A., Basílio, Daniel S., Clarkson, Bruno, Agostinis, André O., Smith, Andrew B. T., Vaz-De-Mello, Fernando Z. &amp; Almeida, Lúcia M." docDate="2024" docId="03D18442FFDCFF9B317CFF6E5C70FF7D" docLanguage="en" docName="zlad115.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad115" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Homalochilus nigripennis Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="116" masterDocId="FFE8FC3AFFCCFF893202FFE1597BFF90" masterDocTitle="New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data" masterLastPageNumber="135" masterPageNumber="98" pageNumber="114" updateTime="1716365025324" updateUser="ExternalLinkService">
<mods:mods id="71F0813672B16F3AB02C964B50EB3391" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E07E676E49DD30CFD3015CABE4D00EB8">
<mods:title id="682F7A65C560DB6CC5C4CED48284FBEF">New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data</mods:title>
</mods:titleInfo>
<mods:name id="78DDF440915231C4D8548BA15EC2789C" type="personal">
<mods:role id="62102AC0A28E609271388F8303199E77">
<mods:roleTerm id="D5F38993AC6C08664852E2A5743651AD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DADF2E2F4F2B1223DCEF49F212E9FA90">Cherman, Mariana A.</mods:namePart>
<mods:affiliation id="A4D01367EF764E188D37692575FA980C">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil &amp; Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Av. Fernando Correia, no. 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil</mods:affiliation>
<mods:nameIdentifier id="D37F9D06FCC729F8AC3F8A7D6433C819" type="email">marianabioar@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="BCF93E25A9995E10935D8ECFE731FBC5" type="personal">
<mods:role id="BFF7E712BDAF4E76F2755EED28D86CA6">
<mods:roleTerm id="1446E1F637DD0125924995B56E8391C8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32F7C788EA16F636532400450E4FC26C">Basílio, Daniel S.</mods:namePart>
<mods:affiliation id="BB433F8A17B8E39F71640009AC176D1E">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
</mods:name>
<mods:name id="AC4552D7634E523C009BD0BB64F5CA02" type="personal">
<mods:role id="E6FC2F3A063C5E061D783286D1703766">
<mods:roleTerm id="2F315F60622067AC8D93FECFEFA67FDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D43A39178CE5A0363D4BC79180F2199">Clarkson, Bruno</mods:namePart>
<mods:affiliation id="85EACA0FBD2E4200C7145E8518B44E98">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil &amp; Instituto Oswaldo Cruz, Fundação Oswaldo Cruz, Av. Brasil, 4365, Pavilhão Mourisco, Manguinhos, 21040 - 360, Rio de Janeiro, Rio de Janeiro, Brazil &amp; Laboratório de Entomologia e Fitopatologia, Universidade Estadual do Norte Fluminense Darcy Ribeiro, 28013 - 602 Campos dos Goytacazes, Rio de Janeiro, Brazil</mods:affiliation>
</mods:name>
<mods:name id="5867203AFC765C2F2829F67173F3C60A" type="personal">
<mods:role id="56D157DED46D66CA0B8749C4892BD6EE">
<mods:roleTerm id="F75B065148C6FBFE0A19E88E1EE9D4B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70C435368EB6F3F2D40798E05F6F0691">Agostinis, André O.</mods:namePart>
<mods:affiliation id="0F5DD7181351529BCF80A97BD9B3A2BC">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
</mods:name>
<mods:name id="F34E54331662117D64D7043C8228B6B4" type="personal">
<mods:role id="D401CE0FF0B329D42865A702FFFCF652">
<mods:roleTerm id="37DD2AD442069FD8DA1B8018CC874B5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8ABF7373D035F404A1AD8953CC2AA7B7">Smith, Andrew B. T.</mods:namePart>
<mods:affiliation id="A0DC9209D6394BA205BD638210D90C7C">Canadian Museum of Nature, 1740 Chemin Pink, Gatineau, Quebec, J 9 J 3 N 7, Canada</mods:affiliation>
</mods:name>
<mods:name id="7EEB081DDF01D600DB78A1DB67FBF09B" type="personal">
<mods:role id="B3CFC6A20D727002D3846B6B2E7731C6">
<mods:roleTerm id="2B24EBCADFE64C0F583516D6A6FD82BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2357CD2B12FD70792864EADBB800DA53">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:affiliation id="DD3A5EBF44B632E0796B3C6E87D5BEC1">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Av. Fernando Correia, no. 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:name id="AE33B9A65DFD733C7205B951E6F26902" type="personal">
<mods:role id="59813FDB96393D837F37CC90D959A465">
<mods:roleTerm id="F82E9C281987A5155B71B040D988FE9C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="42A2A808F7A9216D9F7BEE5DA08754D8">Almeida, Lúcia M.</mods:namePart>
<mods:affiliation id="C207135C06D16FBF5C04C8B857FC0AC3">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="4AD5840AD2D3017889C13E3F1C464074">text</mods:typeOfResource>
<mods:relatedItem id="2549C5EF86714BD015882A269E152C9F" type="host">
<mods:titleInfo id="73988DC1F9BBE0EBECD447A283FF58A7">
<mods:title id="B91A43D3B5B81722B68ABB49E606DC4E">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E0D345BFCAE12788D30E2154DE0ADF29">
<mods:date id="C4F91076CD1618BED5DE063CB1A7961D">2024</mods:date>
<mods:detail id="0476CE4A25147F99EA571D878AF0AB79" type="pubDate">
<mods:number id="E677D6D12C9C1B4F7C5780AD0F028085">2023-10-09</mods:number>
</mods:detail>
<mods:detail id="2788C8C737D497060CD8BE75627CFF92" type="volume">
<mods:number id="CEDD085DC38A471951BC8932552EC5E2">201</mods:number>
</mods:detail>
<mods:detail id="95AA967A7D598C689F6760AD7FAB44F2" type="issue">
<mods:number id="FDCB67F3BFF8A47221E4AAC816851B00">1</mods:number>
</mods:detail>
<mods:extent id="F00D9DBB6B17112A9AA04C0018614ADC" unit="page">
<mods:start id="9DCC4189B3916FBC2F0EF77345282B57">98</mods:start>
<mods:end id="6CDE7482A5EB89A653B51C10FE00DEDF">135</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E7D64FD49A5CA7EC066F8B01C6DCE41C">
<mods:url id="0827115FC6CDB44859F5B18CA3C7D8A5">http://dx.doi.org/10.1093/zoolinnean/zlad115</mods:url>
</mods:location>
<mods:classification id="B2C590D070A0808DCBEB86F1F537D711">journal article</mods:classification>
<mods:identifier id="EB2263D0FD1EA0B6FE041D04C810F205" type="DOI">10.1093/zoolinnean/zlad115</mods:identifier>
<mods:identifier id="13CB8A2944DCA04832091882C7252E0F" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="8EFBF5021B87FFFD57C3346C1D5B0452" type="Zenodo-Dep">11241094</mods:identifier>
<mods:identifier id="34D0BE90F2CF54F4931C59E0A0BDE867" type="ZooBank">E6D9AF7E-F0AD-4656-B2F2-7FBFAA0312B3R</mods:identifier>
</mods:mods>
<treatment id="03D18442FFDCFF9B317CFF6E5C70FF7D" LSID="urn:lsid:plazi:treatment:03D18442FFDCFF9B317CFF6E5C70FF7D" httpUri="http://treatment.plazi.org/id/03D18442FFDCFF9B317CFF6E5C70FF7D" lastPageId="18" lastPageNumber="116" pageId="16" pageNumber="114">
<subSubSection id="C36266DFFFDCFF99317CFF6E5C24FF3A" box="[894,1375,143,170]" pageId="16" pageNumber="114" type="nomenclature">
<paragraph id="8BC73554FFDCFF99317CFF6E5C24FF3A" blockId="16.[894,1375,143,170]" box="[894,1375,143,170]" pageId="16" pageNumber="114">
<heading id="D08F8238FFDCFF99317CFF6E5C24FF3A" bold="true" box="[894,1375,143,170]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="114" reason="2">
<emphasis id="B90CE946FFDCFF99317CFF6E5C24FF3A" bold="true" box="[894,1375,143,170]" pageId="16" pageNumber="114">
<taxonomicName id="4C784ED7FFDCFF99317CFF6E5C72FF3A" ID-CoL="BV5Y9" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" box="[894,1289,143,170]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripennis" status="sp. nov.">
<emphasis id="B90CE946FFDCFF99317CFF6E5DE4FF3A" bold="true" box="[894,1183,143,170]" italics="true" pageId="16" pageNumber="114">Homalochilus nigripennis</emphasis>
Cherman
</taxonomicName>
,
<taxonomicNameLabel id="A23F543DFFDCFF993711FF715C24FF3A" box="[1299,1375,144,170]" pageId="16" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36266DFFFDCFF9B362CFF5E5B56F801" lastPageId="18" lastPageNumber="116" pageId="16" pageNumber="114" type="description">
<paragraph id="8BC73554FFDCFF99362CFF5E5DD4FF49" blockId="16.[1070,1199,191,217]" box="[1070,1199,191,217]" pageId="16" pageNumber="114">
<emphasis id="B90CE946FFDCFF99362CFF5E5DD4FF49" bold="true" box="[1070,1199,191,217]" pageId="16" pageNumber="114">
(
<figureCitation id="134329D1FFDCFF99363BFF5E5DDFFF49" box="[1081,1188,191,217]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="16" pageNumber="114">Fig. 6AG</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BC73554FFDCFF99362CFF0B5DD4FE93" blockId="16.[1070,1199,233,259]" box="[1070,1199,233,259]" pageId="16" pageNumber="114">
<emphasis id="B90CE946FFDCFF99362CFF0B5DD4FE93" box="[1070,1199,233,259]" italics="true" pageId="16" pageNumber="114">Type material</emphasis>
</paragraph>
<paragraph id="8BC73554FFDCFF993128FEFC5D6AFEC4" blockId="16.[810,1441,285,340]" pageId="16" pageNumber="114">
ZooBank:
<uri id="FFE93956FFDCFF993195FEFC5D6AFEC4" pageId="16" pageNumber="114">
urn:lsid:zoobank.org:act:
<uuid id="FFDE0F81FFDCFF993695FEFC5D6AFEC4" pageId="16" pageNumber="114">51B1BCAE-E8E3-4ED7- B406-D5691B40352A</uuid>
</uri>
</paragraph>
<paragraph id="8BC73554FFDCFF993128FE8D5CE4F976" blockId="16.[810,1460,364,1860]" pageId="16" pageNumber="114">
<taxonomicName id="4C784ED7FFDCFF993128FE8D5D4BFE13" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" box="[810,1072,364,388]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripennis">
<emphasis id="B90CE946FFDCFF993128FE8D5D4BFE13" box="[810,1072,364,388]" italics="true" pageId="16" pageNumber="114">Homalochilus nigripennis</emphasis>
</taxonomicName>
<typeStatus id="54C38BF6FFDCFF993646FE8D5DD9FE14" box="[1092,1186,364,388]" pageId="16" pageNumber="114" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D7EFEDDFFDCFF9936B9FE8E5DAEFE13" box="[1211,1237,367,387]" count="1" pageId="16" pageNumber="114" type="male"></specimenCount>
, (NHMB): [white typeset] Humahuaca/
<collectingCountry id="F36F75C4FFDCFF993616FE6A5DFBFE33" box="[1044,1152,395,419]" name="Argentina" pageId="16" pageNumber="114">Argentina</collectingCountry>
/
<quantity id="4C8098B1FFDCFF993693FE6A5D9FFE32" box="[1169,1252,395,419]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="16" pageNumber="114" unit="m" value="2950.0">2950 m</quantity>
, I.56, [white handwritten]
<taxonomicName id="4C784ED7FFDCFF993189FE4B5D64FE52" authorityName="Blanchard" authorityYear="1851" box="[907,1055,426,450]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">Homalochilus</taxonomicName>
/niger Blanch, [white, outlined black, typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF993643FE2B5DACFE72" authorityName="Blanchard" authorityYear="1851" box="[1089,1239,458,482]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">Homalochilus</taxonomicName>
/puntatostriatus/ Bl/ det. Cherman M. A. 2018, [red typeset]
<taxonomicName id="4C784ED7FFDCFF9936D0FE085CD0FD91" authorityName="Blanchard" authorityYear="1851" box="[1234,1451,489,513]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">HOMALOCHILUS</taxonomicName>
/ NIGRIPENNIS/
<typeStatus id="54C38BF6FFDCFF9931DEFDE95D10FDB0" box="[988,1131,520,544]" pageId="16" pageNumber="114" type="holotype">HOLOTYPE</typeStatus>
/Cherman M. A., genitalia mounted.
<typeStatus id="54C38BF6FFDCFF993196FDC95A8CFDD0" box="[916,1015,552,576]" pageId="16" pageNumber="114" type="paratype">Paratypes</typeStatus>
(117), all bearing the label [yellow typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF99361DFDA65D83FDCF" authorityName="Blanchard" authorityYear="1851" box="[1055,1272,583,607]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">HOMALOCHILUS</taxonomicName>
/NIGRIPENNIS/
<typeStatus id="54C38BF6FFDCFF993128FD875ACBFDEE" box="[810,944,614,638]" pageId="16" pageNumber="114" type="paratype">PARATYPE</typeStatus>
/Cherman M. A..
<specimenCount id="9D7EFEDDFFDCFF99367CFD875C53FDED" box="[1150,1320,614,638]" count="24" pageId="16" pageNumber="114" type="female">Twenty-four ♀</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF99376DFD875CC8FDED" box="[1391,1459,614,638]" count="56" pageId="16" pageNumber="114" type="male">56 ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF993128FD675AF6FD0E" box="[810,909,646,670]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
(NHMB) with the same data as the
<typeStatus id="54C38BF6FFDCFF993702FD675C25FD0E" box="[1280,1374,646,670]" pageId="16" pageNumber="114" type="holotype">holotype</typeStatus>
. Among those,
<specimenCount id="9D7EFEDDFFDCFF993172FD445AC2FD2C" box="[880,953,677,700]" count="1" pageId="16" pageNumber="114" type="female">one ♀</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF9931F3FD445D3DFD2C" box="[1009,1094,677,701]" count="8" pageId="16" pageNumber="114" type="male">eight ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF99364DFD445DC9FD2D" box="[1103,1202,677,701]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
with an additional label [white, outlined black, typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF993722FD255A18FD6B" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="species" species="niger">Homalochilus niger</taxonomicName>
/det. 1964/A. Martínez.
<specimenCount id="9D7EFEDDFFDCFF99366CFD055DBBFD6B" box="[1134,1216,740,764]" count="1" pageId="16" pageNumber="114" type="female">One ♀</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF9936FEFD025C3EFD6B" box="[1276,1349,739,763]" count="5" pageId="16" pageNumber="114" type="male">five ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF993752FD055CC8FD6C" box="[1360,1459,740,764]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
(NHMB): [white typeset] Cienaguilla, Arg./
<collectingRegion id="49BCFBB6FFDCFF993711FCE25C36FC8B" box="[1299,1357,771,795]" country="Argentina" name="Jujuy" pageId="16" pageNumber="114">Jujuy</collectingRegion>
,
<quantity id="4C8098B1FFDCFF99375CFCE25CD0FC8B" box="[1374,1451,771,795]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="114" unit="m" value="3500.0">3500m</quantity>
/ Pena, 9.2.70.
<specimenCount id="9D7EFEDDFFDCFF9931BAFCC35D7CFCA9" box="[952,1031,802,826]" count="1" pageId="16" pageNumber="114" type="male">One ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF99360DFCC25D13FCAB" box="[1039,1128,803,827]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(MZUC): [white handwritten] 1340, [white handwritten]
<collectingCountry id="F36F75C4FFDCFF993686FCA05D97FCC9" box="[1156,1260,833,857]" name="Argentina" pageId="16" pageNumber="114">Argentina</collectingCountry>
/Humahuaca/
<collectingRegion id="49BCFBB6FFDCFF99377BFCA05CC8FCC9" box="[1401,1459,833,857]" country="Argentina" name="Jujuy" pageId="16" pageNumber="114">Jujuy</collectingRegion>
Bosq/II.44, [white typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF993716FC805CD0FCE9" authorityName="Blanchard" authorityYear="1851" box="[1300,1451,865,889]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">Homalochilus</taxonomicName>
/ punctatostriatus/M.A. Cherman det. 2016.
<specimenCount id="9D7EFEDDFFDCFF9936FFFC615C34FC07" box="[1277,1359,896,920]" count="1" pageId="16" pageNumber="114" type="male">One ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF993758FC605CC8FC09" box="[1370,1459,897,921]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(MZUC): [white handwritten] 1341, [white handwritten]
<collectingCountry id="F36F75C4FFDCFF99312CFC5E5AEDFC47" box="[814,918,959,983]" name="Argentina" pageId="16" pageNumber="114">Argentina</collectingCountry>
/Humahuaca/
<collectingRegion id="49BCFBB6FFDCFF993621FC5E5D26FC47" box="[1059,1117,959,983]" country="Argentina" name="Jujuy" pageId="16" pageNumber="114">Jujuy</collectingRegion>
Bosq/II.44, [white typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF9931F0FC3F5DFDFC66" authorityName="Blanchard" authorityYear="1851" box="[1010,1158,990,1014]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">Homalochilus</taxonomicName>
/
<taxonomicName id="4C784ED7FFDCFF99368DFC3F5C64FC66" authority="Blanch" authorityName="Blanchard" authorityYear="1851" box="[1167,1311,990,1014]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="species" species="niger">niger Blanch</taxonomicName>
/R. Gutierrez- Det
<specimenCount id="9D7EFEDDFFDCFF993160FBE05A07FB85" box="[866,892,1025,1045]" count="1" pageId="16" pageNumber="114" type="male"></specimenCount>
/52.
<specimenCount id="9D7EFEDDFFDCFF9931B8FC1F5D69FB85" box="[954,1042,1022,1046]" count="1" pageId="16" pageNumber="114" type="male">One ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF993626FC1F5D06FB86" box="[1060,1149,1022,1046]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(CMNC): [white typeset]
<collectingCountry id="F36F75C4FFDCFF99312CFBFC5AB0FBA5" box="[814,971,1053,1077]" name="Argentina" pageId="16" pageNumber="114">ARGENTINA</collectingCountry>
/Humahuaca/
<date id="FFC61394FFDCFF993658FBFC5C72FBA5" box="[1114,1289,1053,1077]" pageId="16" pageNumber="114" value="1956-01-06">6 January 1956</date>
/
<quantity id="4C8098B1FFDCFF993713FBFC5C0BFBA5" box="[1297,1392,1053,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="16" pageNumber="114" unit="m" value="2950.0">2950 m</quantity>
.s.n.m..
<specimenCount id="9D7EFEDDFFDCFF993128FBDD5A01FBC3" box="[810,890,1084,1108]" count="1" pageId="16" pageNumber="114" type="male">One ♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF9931B3FBDC5A81FBC3" box="[945,1018,1085,1108]" count="1" pageId="16" pageNumber="114" type="female">one ♀</specimenCount>
<typeStatus id="54C38BF6FFDCFF993600FBDC5D1EFBC5" box="[1026,1125,1085,1109]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
(CMNC): [white handwritten]
<collectingCountry id="F36F75C4FFDCFF99312CFBBD5AB1FBE4" box="[814,970,1116,1140]" name="Argentina" pageId="16" pageNumber="114">ARGENTINA</collectingCountry>
/Do Cochinoca/
<collectingRegion id="49BCFBB6FFDCFF993691FBBA5DB6FBE3" box="[1171,1229,1115,1139]" country="Argentina" name="Jujuy" pageId="16" pageNumber="114">Jujuy</collectingRegion>
/Tres Cruces/Coll. Martínez/Dic.-944, [white typeset] H. &amp; A. HOWDEN/ COLLECTION/
<emphasis id="B90CE946FFDCFF9931DCFB7A5A81FB22" box="[990,1018,1179,1202]" italics="true" pageId="16" pageNumber="114">ex.</emphasis>
A.Martinez coll..
<specimenCount id="9D7EFEDDFFDCFF9936E1FB7B5C39FB21" box="[1251,1346,1178,1202]" count="1" pageId="16" pageNumber="114" type="male">One ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF993758FB7A5CC8FB23" box="[1370,1459,1179,1203]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(CMNC): [white typeset and handwritten]
<collectingCountry id="F36F75C4FFDCFF993712FB5B5CD0FB42" box="[1296,1451,1210,1234]" name="Argentina" pageId="16" pageNumber="114">ARGENTINA</collectingCountry>
/
<collectingRegion id="49BCFBB6FFDCFF993128FB385A18FB61" box="[810,867,1241,1265]" country="Argentina" name="Jujuy" pageId="16" pageNumber="114">Jujuy</collectingRegion>
/La Quiaca/COLL. DURET/1944/Cipolletti leg., [white typeset] H. &amp; A. HOWDEN/COLLECTION/
<emphasis id="B90CE946FFDCFF99371EFB185C43FA80" box="[1308,1336,1273,1296]" italics="true" pageId="16" pageNumber="114">ex.</emphasis>
A.Martinez coll., [white typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF993716FAF65CD0FABF" authorityName="Blanchard" authorityYear="1851" box="[1300,1451,1303,1327]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">Homalochilus</taxonomicName>
/ niger/Blanch./A.MARTINEZ-DET.1956.
<specimenCount id="9D7EFEDDFFDCFF9936EFFAD65C3CFADE" box="[1261,1351,1335,1359]" count="1" pageId="16" pageNumber="114" type="female">One ♀</specimenCount>
<typeStatus id="54C38BF6FFDCFF993758FAD65CC8FADF" box="[1370,1459,1335,1359]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(CMNC): [white typeset] Sgo. de ESTERO/RIO SALADO/ WAGNER COL, [white typeset] H. &amp; A. HOWDEN/ COLLECTION/
<emphasis id="B90CE946FFDCFF9931DCFA745A81FA3C" box="[990,1018,1429,1452]" italics="true" pageId="16" pageNumber="114">ex.</emphasis>
A.Martinez coll..
<specimenCount id="9D7EFEDDFFDCFF9936C8FA745C5BFA3C" box="[1226,1312,1429,1453]" count="1" pageId="16" pageNumber="114" type="male">One ♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF993761FA745CC8FA3C" box="[1379,1459,1429,1452]" count="1" pageId="16" pageNumber="114" type="female">one ♀</specimenCount>
<typeStatus id="54C38BF6FFDCFF993128FA555AF6FA5C" box="[810,909,1460,1484]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
mounted on the same pin (CMNC): [white handwritten]
<collectingCountry id="F36F75C4FFDCFF993198FA325D48FA7B" box="[922,1075,1491,1515]" name="Argentina" pageId="16" pageNumber="114">ARGENTINA</collectingCountry>
/
<collectingRegion id="49BCFBB6FFDCFF993639FA325DF3FA7B" box="[1083,1160,1491,1515]" country="Argentina" name="Jujuy" pageId="16" pageNumber="114">JUJUY</collectingRegion>
/La Quiaca/Cipolletti-leg./ Coll. Martínez/Dic.-943, [white typeset] H. &amp; A. HOWDEN/ COLLECTION/
<emphasis id="B90CE946FFDCFF9931DCF9F25A8FF9BA" box="[990,1012,1555,1578]" italics="true" pageId="16" pageNumber="114">ex</emphasis>
. A.Martinez coll., [white typeset and handwritten]
<taxonomicName id="4C784ED7FFDCFF993196F9D05D51F9D9" authorityName="Blanchard" authorityYear="1851" box="[916,1066,1585,1609]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="114" phylum="Arthropoda" rank="genus">Homalochilus</taxonomicName>
/niger/Bl./A.MARTINEZ-DET.1956. Ten
<specimenCount id="9D7EFEDDFFDCFF99315AF9B55A09F9F8" box="[856,882,1620,1640]" count="1" pageId="16" pageNumber="114" type="male"></specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF9931A5F9B05A87F9F8" box="[935,1020,1617,1641]" count="3" pageId="16" pageNumber="114" type="female">three ♀</specimenCount>
<typeStatus id="54C38BF6FFDCFF993601F9B05D1DF9F9" box="[1027,1126,1617,1641]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
(UMRM),
<specimenCount id="9D7EFEDDFFDCFF9936E3F9B05C4CF9F8" box="[1249,1335,1617,1641]" count="3" pageId="16" pageNumber="114" type="male">three ♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF993769F9B05CC8F9F8" box="[1387,1459,1617,1640]" count="1" pageId="16" pageNumber="114" type="female">one ♀</specimenCount>
<typeStatus id="54C38BF6FFDCFF993128F9915AF8F918" box="[810,899,1648,1672]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(CMNC), and
<specimenCount id="9D7EFEDDFFDCFF993621F9915D0CF917" box="[1059,1143,1648,1672]" count="3" pageId="16" pageNumber="114" type="male">three ♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDCFF9936AAF9915D95F917" box="[1192,1262,1648,1671]" count="1" pageId="16" pageNumber="114" type="female">one ♀</specimenCount>
<typeStatus id="54C38BF6FFDCFF9936F1F9915C37F918" box="[1267,1356,1648,1672]" pageId="16" pageNumber="114" type="paratype">paratype</typeStatus>
(DZUP): [white typeset]
<collectingCountry id="F36F75C4FFDCFF993608F96E5DDCF937" box="[1034,1191,1679,1703]" name="Argentina" pageId="16" pageNumber="114">ARGENTINA</collectingCountry>
/Humahuaca/
<date id="FFC61394FFDCFF993735F96E5A24F956" pageId="16" pageNumber="114" value="1956-01-06">6 January 1956</date>
/
<quantity id="4C8098B1FFDCFF993165F94F5ABBF956" box="[871,960,1710,1734]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="16" pageNumber="114" unit="m" value="2950.0">2950 m</quantity>
. s. n. m..
<specimenCount id="9D7EFEDDFFDCFF99362EF94E5D05F956" box="[1068,1150,1711,1734]" count="2" pageId="16" pageNumber="114" type="male">Two ♂</specimenCount>
<typeStatus id="54C38BF6FFDCFF99368BF94E5D97F957" box="[1161,1260,1711,1735]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
(UMRM): [white typeset]
<collectingCountry id="F36F75C4FFDCFF993185F92F5D5AF976" box="[903,1057,1742,1766]" name="Argentina" pageId="16" pageNumber="114">ARGENTINA</collectingCountry>
/Tres Cruces/
<date id="FFC61394FFDCFF9936B3F92F5C40F976" box="[1201,1339,1742,1766]" pageId="16" pageNumber="114" value="1956-01-06">6 Enero 1956</date>
/
<quantity id="4C8098B1FFDCFF993741F92F5CEFF976" box="[1347,1428,1742,1766]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="114" unit="m" value="3400.0">3400 m</quantity>
..
</paragraph>
<paragraph id="8BC73554FFDCFF993147F90C5CCFF8D4" blockId="16.[810,1460,364,1860]" pageId="16" pageNumber="114">
<materialsCitation id="3B103F09FFDCFF993147F90C5CCFF8D4" collectionCode="NHMB, CMNC, CEMT, DZUP, UMRM, MZUC" pageId="16" pageNumber="114" specimenCount="107" typeStatus="holotype">
<typeStatus id="54C38BF6FFDCFF993147F90C5AD3F895" box="[837,936,1773,1797]" pageId="16" pageNumber="114" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9D7EFEDDFFDCFF9931F2F90C5D06F896" box="[1008,1149,1773,1798]" count="86" pageId="16" pageNumber="114" type="generic" typeStatus="paratype">86 paratypes</specimenCount>
deposited at
<collectionCode id="ED69AD91FFDCFF993724F90F5C0CF895" box="[1318,1399,1774,1797]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="114">NHMB</collectionCode>
. Ten
<typeStatus id="54C38BF6FFDCFF993128F8EC5AF6F8B5" box="[810,909,1805,1829]" pageId="16" pageNumber="114" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED69AD91FFDCFF9931A9F8EC5A86F8B5" box="[939,1021,1805,1829]" pageId="16" pageNumber="114">CMNC</collectionCode>
.
<specimenCount id="9D7EFEDDFFDCFF993605F8ED5DD0F8B5" box="[1031,1195,1804,1829]" count="3" pageId="16" pageNumber="114" type="generic" typeStatus="paratype">Three paratypes</specimenCount>
at
<collectionCode id="ED69AD91FFDCFF9936CBF8EC5C68F8B5" box="[1225,1299,1805,1829]" pageId="16" pageNumber="114">CEMT</collectionCode>
.
<specimenCount id="9D7EFEDDFFDCFF99371EF8EC5CC8F8B5" box="[1308,1459,1805,1829]" count="2" pageId="16" pageNumber="114" type="generic" typeStatus="paratype">Two paratypes</specimenCount>
at
<collectionCode id="ED69AD91FFDCFF993140F8CD5AFCF8D3" box="[834,903,1836,1859]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="16" pageNumber="114" type="University or college">DZUP</collectionCode>
.
<specimenCount id="9D7EFEDDFFDCFF99318DF8CD5D45F8D4" box="[911,1086,1836,1860]" count="15" pageId="16" pageNumber="114" type="generic" typeStatus="paratype">Fifteen paratypes</specimenCount>
at
<collectionCode id="ED69AD91FFDCFF993659F8CD5DCAF8D3" box="[1115,1201,1836,1859]" country="USA" httpUri="http://grbio.org/cool/w4qt-0wzz" name="W.R. Enns Entomology Museum" pageId="16" pageNumber="114">UMRM</collectionCode>
.
<specimenCount id="9D7EFEDDFFDCFF9936BAF8CD5C3FF8D4" box="[1208,1348,1836,1860]" count="1" pageId="16" pageNumber="114" type="generic" typeStatus="paratype">One paratype</specimenCount>
at
<collectionCode id="ED69AD91FFDCFF993763F8CD5CCBF8D4" box="[1377,1456,1836,1860]" country="Chile" httpUri="http://grbio.org/cool/70mh-t5r1" name="Museo de Zoologia, Universidad de Concepcion" pageId="16" pageNumber="114">MZUC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC73554FFDCFF993642F8825DE6F8EC" blockId="16.[810,1459,1891,1984]" box="[1088,1181,1891,1916]" pageId="16" pageNumber="114">
<emphasis id="B90CE946FFDCFF993642F8825DE6F8EC" box="[1088,1181,1891,1916]" italics="true" pageId="16" pageNumber="114">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BC73554FFDCFF9B3128F869585FFE33" blockId="16.[810,1459,1891,1984]" lastBlockId="18.[113,762,144,419]" lastPageId="18" lastPageNumber="116" pageId="16" pageNumber="114">
Body, head, and pronotum entirely black, semiopaque; elytra entirely black, semiopaque, unicolorous with body (
<figureCitation id="134329D1FFDCFF993756F8495CDBF850" box="[1364,1440,1960,1984]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="16" pageNumber="114">Fig. 6A</figureCitation>
); clypeus slightly lobed, head as long as pronotum (
<figureCitation id="134329D1FFDDFF9830A8F89A5B8DF803" box="[682,758,1915,1939]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="17" pageNumber="115">Fig. 6B</figureCitation>
); fovea of distal maxillary palpomere reaching the midline; club of antenna slightly shorter than the funicle; anterior margin of pronotum slightly constricted; disc coarsely and roughly punctate; pronotal posterior corners subangulate (
<figureCitation id="134329D1FFDDFF983146F8585AE3F841" box="[836,920,1977,2001]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="17" pageNumber="115">Fig. 6B</figureCitation>
); metacoxae slightly shorter than the distance between mesocoxae and metacoxae; elytra near two times longer than the pronotum; four elytral ridges, both outermost less noticeable, sparse long bristles along the sutural ridge, scattered bristles on disc, erect long setae on apex of elytra present; protarsomere I longer than tarsomere II in both sexes; disc of ventrites with long bristles (
<figureCitation id="134329D1FFDEFF9B3096FECC5B93FED5" box="[660,744,301,325]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="18" pageNumber="116">Fig. 6C</figureCitation>
); pygidium subtrapezoidal (
<figureCitation id="134329D1FFDEFF9B3399FEAD588AFEF4" box="[411,497,332,356]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="18" pageNumber="116">Fig. 6D</figureCitation>
); parameres with apex curved downwards, forming right angle with the longitudinal axis (
<figureCitation id="134329D1FFDEFF9B32AEFE6A5868FE33" box="[172,275,394,419]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="18" pageNumber="116">Fig. 6F, G</figureCitation>
).
</paragraph>
<caption id="DF0765DCFFDDFF983283F8CD5AD9F8F0" ID-DOI="http://doi.org/10.5281/zenodo.11241106" ID-Zenodo-Dep="11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="17" pageNumber="115" startId="17.[129,194,1836,1860]" targetBox="[180,1423,144,1808]" targetPageId="17" targetType="figure">
<paragraph id="8BC73554FFDDFF983283F8CD5AD9F8F0" blockId="17.[129,1458,1836,1888]" pageId="17" pageNumber="115">
<emphasis id="B90CE946FFDDFF983283F8CD59A3F8D4" bold="true" box="[129,216,1836,1860]" pageId="17" pageNumber="115">Figure 6.</emphasis>
<taxonomicName id="4C784ED7FFDDFF9832DFF8CD58BAF8D4" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" box="[221,449,1836,1860]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="115" phylum="Arthropoda" rank="species" species="nigripennis">
<emphasis id="B90CE946FFDDFF9832DFF8CD58BAF8D4" box="[221,449,1836,1860]" italics="true" pageId="17" pageNumber="115">Homalochilus nigripennis</emphasis>
</taxonomicName>
. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur.
</paragraph>
</caption>
<paragraph id="8BC73554FFDEFF9B3357FE325B6DFE7D" blockId="18.[113,763,467,1500]" box="[341,534,467,493]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B3357FE325B6DFE7D" box="[341,534,467,493]" italics="true" pageId="18" pageNumber="116">
<typeStatus id="54C38BF6FFDEFF9B3357FE3258D7FE7D" box="[341,428,467,493]" pageId="18" pageNumber="116" type="holotype">Holotype</typeStatus>
description
</emphasis>
</paragraph>
<paragraph id="8BC73554FFDEFF9B3273FE185851FA4C" blockId="18.[113,763,467,1500]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B3273FE1859BAFD81" box="[113,193,505,529]" italics="true" pageId="18" pageNumber="116">Length:</emphasis>
8.0 mm; width:
<quantity id="4C8098B1FFDEFF9B3381FE1858A0FD81" box="[387,475,505,529]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="18" pageNumber="116" unit="mm" value="4.9">4.9 mm</quantity>
. Body short, semiopaque, sides parallel, body, head, and pronotum black, elytra on disc red and on margins black; glabrous; eyes almost entirely placed laterally, frons almost as wide as the anterior margin of pronotum; frons and clypeus coplanar; clypeus truncate; galea of maxilla with two teeth, the apical stronger; distal maxillary palpomere acuminate, fovea deep, reaching the midline of the palpomere; labium widened at the transverse midline, slightly wider than it is long; sloped from the transverse carina towards apex; apical emargination wide and rounded; antenna black, with nine antennomeres, club and as long as the funicle; pronotal anterior corners scaly, fused with hypomeron; pronotal disc coarsely punctate, simple punctures sparsely disposed; posterior corners obtuse angulate; mesocoxae and metacoxae almost equal in length; scutellum long; elytra flattened, near 1.5 longer than the pronotum; four elytral ridges, the outermost less noticeable; subapical callus strong, callus and apex of the sutural ridge at the same level; three protibial teeth, the basal tooth shorter than the other two (angle obtuse in relation to axis); mesotibia cylindrical, two transverse carina, the apical complete; metatibial spurs strongly uneven (
<figureCitation id="134329D1FFDEFF9B3368FB6A58C8FB33" box="[362,435,1163,1187]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="18" pageNumber="116">Fig. 6E</figureCitation>
), the longest almost two times the diameter of the tibial apex; all tarsi cylindrical (in both sexes), protarsal claws simple, mesotarsal and metatarsal claws bifid (in females all claws bifid); protarsomere I slightly shorter than tarsomere II (in both sexes); propygidium exposed, scaly in basal region, mostly glabrous; pygidium flat, as wide as it is long; subtrapezoidal; pygidial disc glabrous, scaly basally; basal region strongly sulcate, basally widened and slightly swollen; parameres narrow, as long as the basal region, apex curved downwards, forming right angle with the longitudinal axis.
</paragraph>
<paragraph id="8BC73554FFDEFF9B338BF9EA5899F9B4" blockId="18.[113,704,1547,1609]" box="[393,482,1547,1572]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B338BF9EA5899F9B4" box="[393,482,1547,1572]" italics="true" pageId="18" pageNumber="116">Variation</emphasis>
</paragraph>
<paragraph id="8BC73554FFDEFF9B3273F9D05BBBF9D8" blockId="18.[113,704,1547,1609]" box="[113,704,1584,1609]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B3273F9D05875F9D8" box="[113,270,1585,1609]" italics="true" pageId="18" pageNumber="116">
Male
<typeStatus id="54C38BF6FFDEFF9B32ABF9D05872F9D8" box="[169,265,1585,1608]" pageId="18" pageNumber="116" type="paratype">paratypes</typeStatus>
:
</emphasis>
Length: 8.0
<quantity id="4C8098B1FFDEFF9B339EF9D05896F9D8" box="[412,493,1585,1609]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="18" pageNumber="116" unit="mm" value="9.4">9.4 mm</quantity>
; width: 4.25.0 mm.
</paragraph>
<paragraph id="8BC73554FFDEFF9B3273F98B59CFF970" blockId="18.[113,763,1642,1760]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B3273F98B5851F912" box="[113,298,1642,1666]" italics="true" pageId="18" pageNumber="116">
Female
<typeStatus id="54C38BF6FFDEFF9B32C3F98A585EF912" box="[193,293,1643,1666]" pageId="18" pageNumber="116" type="paratype">paratypes</typeStatus>
:
</emphasis>
Length: 8.0
<quantity id="4C8098B1FFDEFF9B33C6F98B5B61F911" box="[452,538,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="18" pageNumber="116" unit="mm" value="9.6">9.6 mm</quantity>
; width: 4.25.0 mm. The females also have the apical carina on mesotibia complete, while the metatibial spurs are strongly uneven especially in males.
</paragraph>
<paragraph id="8BC73554FFDEFF9B3379F8ED5B80F8D9" blockId="18.[113,763,1803,1865]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B3379F8ED588AF8B5" box="[379,497,1803,1829]" italics="true" pageId="18" pageNumber="116">Type locality</emphasis>
<collectingCountry id="F36F75C4FFDEFF9B3273F8D05873F8D9" box="[113,264,1841,1865]" name="Argentina" pageId="18" pageNumber="116">ARGENTINA</collectingCountry>
.
<collectingRegion id="49BCFBB6FFDEFF9B3312F8D0583FF8D9" box="[272,324,1841,1865]" country="Argentina" name="Jujuy" pageId="18" pageNumber="116">Jujuy</collectingRegion>
, Humahuaca [23°12ʹ45.4ʹʹS 65°20ʹ54.0ʹʹW].
</paragraph>
<paragraph id="8BC73554FFDEFF9B333DF8965B56F801" blockId="18.[113,557,1911,1973]" box="[319,557,1911,1937]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B333DF8965B56F801" box="[319,557,1911,1937]" italics="true" pageId="18" pageNumber="116">Geographical distribution</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36266DFFFDEFF9B3273F87C5C70FF7D" pageId="18" pageNumber="116" type="materials_examined">
<paragraph id="8BC73554FFDEFF9B3273F87C5B56F825" blockId="18.[113,557,1911,1973]" box="[113,557,1949,1973]" pageId="18" pageNumber="116">
<materialsCitation id="3B103F09FFDEFF9B3273F87C5B52F825" box="[113,553,1949,1973]" country="Argentina" location="Jujuy" pageId="18" pageNumber="116" specimenCount="1" stateProvince="Jujuy">
<collectingCountry id="F36F75C4FFDEFF9B3273F87C5873F825" box="[113,264,1949,1973]" name="Argentina" pageId="18" pageNumber="116">ARGENTINA</collectingCountry>
(
<collectingRegion id="49BCFBB6FFDEFF9B331EF87C5829F825" box="[284,338,1949,1973]" country="Argentina" name="Jujuy" pageId="18" pageNumber="116">Jujuy</collectingRegion>
,
<collectingRegion id="49BCFBB6FFDEFF9B3358F87C5B59F825" box="[346,546,1949,1973]" country="Argentina" name="Santiago del Estero" pageId="18" pageNumber="116">Santiago del Estero</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC73554FFDEFF9B3646FF715DE3FF3A" blockId="18.[810,1459,144,238]" box="[1092,1176,144,170]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B3646FF715DE3FF3A" box="[1092,1176,144,170]" italics="true" pageId="18" pageNumber="116">Remarks</emphasis>
</paragraph>
<paragraph id="8BC73554FFDEFF9B3128FF575C70FF7D" blockId="18.[810,1459,144,238]" pageId="18" pageNumber="116">
<bibRefCitation id="EFE948A5FFDEFF9B3128FF575D77FF5E" author="Gutierrez R" box="[810,1036,182,206]" pageId="18" pageNumber="116" pagination="207 - 27" refId="ref27805" refString="Gutierrez R. Notas sobre Scarabaeidae Neotropicos. Revista Chilena de Entomologia 1952; 2: 207 - 27." type="journal article" year="1952">Gutiérrez (1952: 213</bibRefCitation>
, 214) erroneously used part of the
<typeStatus id="54C38BF6FFDEFF9B3787FF565CC8FF5F" box="[1413,1459,183,207]" pageId="18" pageNumber="116">type</typeStatus>
series in his redescription of
<emphasis id="B90CE946FFDEFF9B364EFF375C70FF7D" box="[1100,1291,214,238]" italics="true" pageId="18" pageNumber="116">
<taxonomicName id="4C784ED7FFDEFF9B364EFF375C7CFF7D" authorityName="Blanchard" authorityYear="1851" box="[1100,1287,214,238]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="niger">Homalochilus niger</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>