236 lines
36 KiB
XML
236 lines
36 KiB
XML
<document id="2CFA5A8D77981C339B2C0045100AC822" ID-DOI="10.1093/zoolinnean/zlad115" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241094" ID-ZooBank="E6D9AF7E-F0AD-4656-B2F2-7FBFAA0312B3R" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1716363659947" checkinUser="plazi" docAuthor="Cherman, Mariana A., Basílio, Daniel S., Clarkson, Bruno, Agostinis, André O., Smith, Andrew B. T., Vaz-De-Mello, Fernando Z. & Almeida, Lúcia M." docDate="2024" docId="03D18442FFC6FF8332EEFB6B5CE2F804" docLanguage="en" docName="zlad115.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad115" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Melolonthidae" docType="key" docVersion="1" lastPageNumber="108" masterDocId="FFE8FC3AFFCCFF893202FFE1597BFF90" masterDocTitle="New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data" masterLastPageNumber="135" masterPageNumber="98" pageNumber="108" updateTime="1716408306816" updateUser="julia">
|
||
<mods:mods id="E7D86BE656A57DDF6B86CA64FC359ED6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="2357964ECCB9BEF2ED5BD27D363C7986">
|
||
<mods:title id="410F2C299AA14DA1E2DE40E1A747DD55">New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="910DA61C2D8A01FCBC77DDD19867D7CF" type="personal">
|
||
<mods:role id="0989E5E24A723A08CD6D44C02B3192DF">
|
||
<mods:roleTerm id="E39A9F2328480BC0E7FD26BE37924309">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2579DB559833A904DE2C54365B287E5A">Cherman, Mariana A.</mods:namePart>
|
||
<mods:affiliation id="2E74EAA625245F28D426DBE73D160EF2">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil & Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Av. Fernando Correia, no. 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil</mods:affiliation>
|
||
<mods:nameIdentifier id="F413AE044142601860238B6DCA551C2C" type="email">marianabioar@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="62FFC8A02CDB8F3C2B37A3D2D7DF15BD" type="personal">
|
||
<mods:role id="41A9F19D3FD9E0962574A2EE2962245B">
|
||
<mods:roleTerm id="05CE50BDA3FA74C16749988D543A03B9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="49468B3F5B3C80D789169CB257D911F3">Basílio, Daniel S.</mods:namePart>
|
||
<mods:affiliation id="EDD0E83AA33A3D52913BA2640E0010B5">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="8AB6BE31637F8B7B984510D2C64E8CAF" type="personal">
|
||
<mods:role id="727D2DA959EFBD01E9DD42CC7A11202F">
|
||
<mods:roleTerm id="7A0906F625E0EDB0B8AA4ADB066ACDE5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="ADE59B972785AB83724F2069C420813C">Clarkson, Bruno</mods:namePart>
|
||
<mods:affiliation id="4D8425C566AE3D74602DA8CDA5BF59B5">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil & Instituto Oswaldo Cruz, Fundação Oswaldo Cruz, Av. Brasil, 4365, Pavilhão Mourisco, Manguinhos, 21040 - 360, Rio de Janeiro, Rio de Janeiro, Brazil & Laboratório de Entomologia e Fitopatologia, Universidade Estadual do Norte Fluminense Darcy Ribeiro, 28013 - 602 Campos dos Goytacazes, Rio de Janeiro, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="B7C563185919B5D08894E537BDD7C740" type="personal">
|
||
<mods:role id="6F08F9656DCB20E4B388ECB5B07AA580">
|
||
<mods:roleTerm id="FE883F34B8268D590D563FE11C517464">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="55EF0B72F2C25F8C208271F178E0849E">Agostinis, André O.</mods:namePart>
|
||
<mods:affiliation id="2337423E573B5DB5C7E2192E40DE6ECC">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="CE5AEEFD756B26B748E5771380A3745E" type="personal">
|
||
<mods:role id="FD67DE0649D351D8FD574E9C0A28799A">
|
||
<mods:roleTerm id="F2F5D299502849CDD329ED72A10D2042">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AC15A7F8343B11CD7008831D6AA32AB2">Smith, Andrew B. T.</mods:namePart>
|
||
<mods:affiliation id="5E2FE6C810B987A6F5CC7F50CEFAB78A">Canadian Museum of Nature, 1740 Chemin Pink, Gatineau, Quebec, J 9 J 3 N 7, Canada</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="D8789A1BE08561F23C0FD6F7C1561639" type="personal">
|
||
<mods:role id="57E379C8E4971DF2D266BDB117ABECF4">
|
||
<mods:roleTerm id="8FC3D1EF45A04CBA461C97C531F128E9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0479E9CD25F63397F569A6CFB36BB9D1">Vaz-De-Mello, Fernando Z.</mods:namePart>
|
||
<mods:affiliation id="58E40E3DAD717FF4317239A379426A19">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Av. Fernando Correia, no. 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="DA79CE965648E31B8F9FF6354B063D4F" type="personal">
|
||
<mods:role id="6B2B6CB7E70DC69412162A78FAF39E91">
|
||
<mods:roleTerm id="E392FB6D1362EE0D3E52EF7FCD165114">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2AF22893B202C7FE503A506505A38522">Almeida, Lúcia M.</mods:namePart>
|
||
<mods:affiliation id="8E183A89D4306279772D94B67025AC0F">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="512C5AF3CFB675DC71D5445C8A51A600">text</mods:typeOfResource>
|
||
<mods:relatedItem id="13EDE2CC79F9BB57ECD1795DBEA3AC9A" type="host">
|
||
<mods:titleInfo id="62F2FE4467D6D171BF7199D7C41AEDE0">
|
||
<mods:title id="BBEBA8F7033F4506811EA316C9AB22DA">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="5E00B576ACDEF171F92EC4B18D4C8DDA">
|
||
<mods:date id="7AB94D2DED2BCEC69A2F28F5DFA4B462">2024</mods:date>
|
||
<mods:detail id="BF24181AEDB3716290F86E2B5ECCD912" type="pubDate">
|
||
<mods:number id="B81405EA1093F236EF08826B63ADB21C">2023-10-09</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="47A884678150B37C1EF34A0C36D41920" type="volume">
|
||
<mods:number id="3D90C955079379DA20FDB6E4A2E77976">201</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="26A5B82039869E0E0516BCECE3FD923F" type="issue">
|
||
<mods:number id="93CED5FD68B7AFA4B6CBDEFBC48AB9D1">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6B56B16049D7BD6776FFE49F71A63042" unit="page">
|
||
<mods:start id="531F1BF064BA9C7A4E60A85B31D53739">98</mods:start>
|
||
<mods:end id="E5579849CF5BF3485AFBD3C4B2A89547">135</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="2CA1674393DB81CB1A61CD4112CBE35B">
|
||
<mods:url id="D7D8B29566BD1CCEB93E117927628332">http://dx.doi.org/10.1093/zoolinnean/zlad115</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="B0D06CF1FFC716200058B0A0BCCB65C0">journal article</mods:classification>
|
||
<mods:identifier id="24CBA20E7D5457EEAE51AE37BC1AD8F1" type="DOI">10.1093/zoolinnean/zlad115</mods:identifier>
|
||
<mods:identifier id="18851B28B70F4DB3ECA38FEF2FF76C53" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="D4FE6EEE4A40D16D15B77EC71CAA9C76" type="Zenodo-Dep">11241094</mods:identifier>
|
||
<mods:identifier id="9574CDF4773EB80F310CFB9100725D1E" type="ZooBank">E6D9AF7E-F0AD-4656-B2F2-7FBFAA0312B3R</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D18442FFC6FF8332EEFB6B5CE2F804" LSID="urn:lsid:plazi:treatment:03D18442FFC6FF8332EEFB6B5CE2F804" httpUri="http://treatment.plazi.org/id/03D18442FFC6FF8332EEFB6B5CE2F804" lastPageNumber="108" pageId="10" pageNumber="108">
|
||
<subSubSection id="C36266DFFFC6FF8332EEFB6B5C42FB35" box="[236,1337,1162,1189]" pageId="10" pageNumber="108" type="nomenclature">
|
||
<paragraph id="8BC73554FFC6FF8332EEFB6B5C42FB35" blockId="10.[140,1433,1162,1941]" box="[236,1337,1162,1189]" pageId="10" pageNumber="108">
|
||
IDENTIFICATION KEY TO NEW WORLD
|
||
<taxonomicName id="4C784ED7FFC6FF8330E1FB6B5AEBFB34" box="[739,912,1162,1188]" family="Melolonthidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" rank="tribe">DIPLOTAXINI</taxonomicName>
|
||
[modified from
|
||
<bibRefCitation id="EFE948A5FFC6FF83363DFB6B5C56FB34" author="Cherman MA & Mise KM & Moron MA" box="[1087,1325,1162,1188]" pageId="10" pageNumber="108" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">
|
||
Cherman
|
||
<emphasis id="B90CE946FFC6FF8336A9FB6A5DA0FB34" box="[1195,1243,1162,1188]" italics="true" pageId="10" pageNumber="108">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
]
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36266DFFFC6FF83328EFB2A5CE2F804" pageId="10" pageNumber="108" type="key">
|
||
<key id="FFE96265FFC6FF83328EFB2A5CE2F804" pageId="10" pageNumber="108">
|
||
<keyStep id="308C2DF1FFC6FF83328EFB2A5CE3FAF1" pageId="10" pageNumber="108">
|
||
<paragraph id="8BC73554FFC6FF83328EFB2A5CE3FAB2" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EFB2A5CE3FAB2" pageId="10" pageNumber="108">
|
||
1. Division between clypeus and frons distinct [see
|
||
<bibRefCitation id="EFE948A5FFC6FF8330B1FB2A5AE2FB73" author="Cherman MA & Moron MA & Almeida LM" box="[691,921,1227,1251]" pageId="10" pageNumber="108" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
|
||
Cherman
|
||
<emphasis id="B90CE946FFC6FF833118FB2D5A31FB73" box="[794,842,1227,1251]" italics="true" pageId="10" pageNumber="108">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
for more details] (
|
||
<figureCitation id="134329D1FFC6FF83365EFB2A5DDEFB73" box="[1116,1189,1227,1251]" captionStart="Figure 4" captionStartId="12.[115,180,1760,1784]" captionTargetBox="[115,1459,144,1730]" captionTargetId="figure-7@12.[115,1459,144,1732]" captionTargetPageId="12" captionText="Figure 4. Diagnostic features and transformations that support Homalochilus. Head in dorsal view (A), head and anterior portion of prothorax in dorsolateral view (B); head in frontal view (C); maxilla (D); labium (E); elytra, white square indicates erect setae (F); head, pronotum, scutellum and proleg (G), thoracic and abdominal ventrites, white lines indicate size of ventrites (H). White arrows in each figure indicate transformations and/or detailed features." figureDoi="http://doi.org/10.5281/zenodo.11241102" httpUri="https://zenodo.org/record/11241102/files/figure.png" pageId="10" pageNumber="108">Fig. 4A</figureCitation>
|
||
); sutural ridge of elytra not narrowed posteriad (
|
||
<figureCitation id="134329D1FFC6FF8333B8FB0A5B79FA92" box="[442,514,1258,1283]" captionStart="Figure 6" captionStartId="17.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-60@17.[180,1423,144,1808]" captionTargetPageId="17" captionText="Figure 6. Homalochilus nigripennis. Male holotype habitus (A); head and pronotum (B); lateral (C); posterior (D) views; metatibial apex (E); parameres in dorsal (F) and lateral (G) views. White arrow indicates the longest spur." figureDoi="http://doi.org/10.5281/zenodo.11241106" httpUri="https://zenodo.org/record/11241106/files/figure.png" pageId="10" pageNumber="108">Fig. 6A</figureCitation>
|
||
); galea of maxilla curved, not forming an angle at the apex of its external margin (
|
||
<figureCitation id="134329D1FFC6FF83373DFB0A5CF1FA93" box="[1343,1418,1259,1283]" captionStart="Figure 4" captionStartId="12.[115,180,1760,1784]" captionTargetBox="[115,1459,144,1730]" captionTargetId="figure-7@12.[115,1459,144,1732]" captionTargetPageId="12" captionText="Figure 4. Diagnostic features and transformations that support Homalochilus. Head in dorsal view (A), head and anterior portion of prothorax in dorsolateral view (B); head in frontal view (C); maxilla (D); labium (E); elytra, white square indicates erect setae (F); head, pronotum, scutellum and proleg (G), thoracic and abdominal ventrites, white lines indicate size of ventrites (H). White arrows in each figure indicate transformations and/or detailed features." figureDoi="http://doi.org/10.5281/zenodo.11241102" httpUri="https://zenodo.org/record/11241102/files/figure.png" pageId="10" pageNumber="108">Fig. 4D</figureCitation>
|
||
. ............................................................................................................................................................................................................................2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC73554FFC6FF83328EFACB5CE3FAF1" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EFACB5CE3FAF1" pageId="10" pageNumber="108">-. Division between clypeus and frons indistinct; sutural ridge of elytra strongly narrowed posteriad; galea of maxilla straight, forming an angle at the apex of its external margin..................................................................................................................................3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="308C2DF1FFC6FF83328EFA895CE3FA6D" pageId="10" pageNumber="108">
|
||
<paragraph id="8BC73554FFC6FF83328EFA895CE3FA0F" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EFA895CE3FA0F" pageId="10" pageNumber="108">
|
||
2. Pronotal anterior corners reduced, evenly continuous with hypomeron (
|
||
<figureCitation id="134329D1FFC6FF833199FA895A9AFA10" box="[923,993,1384,1408]" captionStart="Figure 4" captionStartId="12.[115,180,1760,1784]" captionTargetBox="[115,1459,144,1730]" captionTargetId="figure-7@12.[115,1459,144,1732]" captionTargetPageId="12" captionText="Figure 4. Diagnostic features and transformations that support Homalochilus. Head in dorsal view (A), head and anterior portion of prothorax in dorsolateral view (B); head in frontal view (C); maxilla (D); labium (E); elytra, white square indicates erect setae (F); head, pronotum, scutellum and proleg (G), thoracic and abdominal ventrites, white lines indicate size of ventrites (H). White arrows in each figure indicate transformations and/or detailed features." figureDoi="http://doi.org/10.5281/zenodo.11241102" httpUri="https://zenodo.org/record/11241102/files/figure.png" pageId="10" pageNumber="108">Fig. 4B</figureCitation>
|
||
); propygidium entirely exposed; pygidium subquadrate; elytra flattened, apex of sutural ridge of elytra rounded (
|
||
<figureCitation id="134329D1FFC6FF83317EFA665AA9FA0F" box="[892,978,1415,1439]" captionStart="Figure 14" captionStartId="28.[113,178,1624,1648]" captionTargetBox="[114,1458,144,1596]" captionTargetId="figure-123@28.[114,1458,144,1596]" captionTargetPageId="28" captionText="Figure 14. Posterior margin of elytra, variation in apex of sutural ridge among Diplotaxini.Apex angulate (A, F); apex subangulate (E) or rounded (B–D). Diplotaxis tristis Kirby (A); Pachrodema castanea Blanchard (B); Homalochilus punctatostriatus (C); Pacuvia philippiana Gutiérrez (D); Liogenys suturalis (E); Careocallus tehuelche (F)." figureDoi="http://doi.org/10.5281/zenodo.11241126" httpUri="https://zenodo.org/record/11241126/files/figure.png" pageId="10" pageNumber="108">Fig. 14C</figureCitation>
|
||
) ......................................................
|
||
<taxonomicName id="4C784ED7FFC6FF833710FA665CE3FA0F" authorityName="Blanchard" authorityYear="1851" box="[1298,1432,1415,1439]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CE946FFC6FF833710FA665CE3FA0F" box="[1298,1432,1415,1439]" italics="true" pageId="10" pageNumber="108">Homalochilus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC73554FFC6FF83328EFA465CE3FA6D" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EFA465CE3FA6D" pageId="10" pageNumber="108">
|
||
-. Pronotal anterior corners distinctly elevated, hypomeron not continuous with dorsal surface; pygidium suboval; propygidium mostly hidden by elytra; elytra flattened or convex, apex of sutural ridge of elytra angled (
|
||
<figureCitation id="134329D1FFC6FF8336CFFA275C58FA4E" box="[1229,1315,1478,1502]" captionStart="Figure 14" captionStartId="28.[113,178,1624,1648]" captionTargetBox="[114,1458,144,1596]" captionTargetId="figure-123@28.[114,1458,144,1596]" captionTargetPageId="28" captionText="Figure 14. Posterior margin of elytra, variation in apex of sutural ridge among Diplotaxini.Apex angulate (A, F); apex subangulate (E) or rounded (B–D). Diplotaxis tristis Kirby (A); Pachrodema castanea Blanchard (B); Homalochilus punctatostriatus (C); Pacuvia philippiana Gutiérrez (D); Liogenys suturalis (E); Careocallus tehuelche (F)." figureDoi="http://doi.org/10.5281/zenodo.11241126" httpUri="https://zenodo.org/record/11241126/files/figure.png" pageId="10" pageNumber="108">Fig. 14A</figureCitation>
|
||
) .................. ............................................................................................................................................................................................................
|
||
<taxonomicName id="4C784ED7FFC6FF833730FA045CE3FA6D" authorityName="Kirby" authorityYear="1837" box="[1330,1432,1509,1533]" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CE946FFC6FF833730FA045CE3FA6D" box="[1330,1432,1509,1533]" italics="true" pageId="10" pageNumber="108">Diplotaxis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="308C2DF1FFC6FF83328EF9E45CE3F9AC" pageId="10" pageNumber="108">
|
||
<paragraph id="8BC73554FFC6FF83328EF9E45CE3F98C" blockId="10.[140,1433,1162,1941]" box="[140,1432,1540,1565]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EF9E45CE3F98C" box="[140,1432,1540,1565]" pageId="10" pageNumber="108">
|
||
3. Frons and clypeus coplanar; concavity on disc of mentum and arched transverse carina absent (
|
||
<figureCitation id="134329D1FFC6FF833681F9E55DA3F98C" box="[1155,1240,1540,1564]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15A</figureCitation>
|
||
) .................
|
||
<taxonomicName id="4C784ED7FFC6FF83374EF9E45CE3F98C" authorityName="Curtis" authorityYear="1844" box="[1356,1432,1541,1564]" class="Insecta" family="Scarabaeidae" genus="Pacuvia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CE946FFC6FF83374EF9E45CE3F98C" box="[1356,1432,1541,1564]" italics="true" pageId="10" pageNumber="108">Pacuvia</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC73554FFC6FF83328EF9C55CE3F9AC" blockId="10.[140,1433,1162,1941]" box="[140,1432,1572,1596]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EF9C55CE3F9AC" box="[140,1432,1572,1596]" pageId="10" pageNumber="108">
|
||
-. Frons and clypeus not coplanar, broad concavity on disc of mentum limited by arched transverse carina (
|
||
<figureCitation id="134329D1FFC6FF8336E0F9C55C4DF9AC" box="[1250,1334,1572,1596]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15B</figureCitation>
|
||
).............4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="308C2DF1FFC6FF83328EF9A25CE3F948" pageId="10" pageNumber="108">
|
||
<paragraph id="8BC73554FFC6FF83328EF9A25CE3F90A" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EF9A25CE3F90A" pageId="10" pageNumber="108">
|
||
4. Inner margin of both eyes more-or-less convergent (
|
||
<figureCitation id="134329D1FFC6FF8330DBF9A25AF5F9CB" box="[729,910,1603,1627]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15D, E, G, H</figureCitation>
|
||
): anterior margin of clypeus widely rounded (
|
||
<figureCitation id="134329D1FFC6FF833770F9A25992F9EA" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15D</figureCitation>
|
||
) or subemarginate (
|
||
<figureCitation id="134329D1FFC6FF8333B8F9835B75F9EA" box="[442,526,1634,1658]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15E</figureCitation>
|
||
), never forming tooth-like projections; frons and clypeus medially flattened, not concave ............................................................................................................................................................................................................................5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC73554FFC6FF83328EF9435CE3F948" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EF9435CE3F948" pageId="10" pageNumber="108">
|
||
-. Inner margin of both eyes parallel (
|
||
<figureCitation id="134329D1FFC6FF83301DF9405BF5F929" box="[543,654,1697,1721]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15C, F</figureCitation>
|
||
); anterior margin of clypeus emarginate, forming tooth-like projections (
|
||
<figureCitation id="134329D1FFC6FF833770F9405993F948" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15C</figureCitation>
|
||
); frons and clypeus medially flattened or concave .............................................................................................................
|
||
<taxonomicName id="4C784ED7FFC6FF833746F9205CE3F948" box="[1348,1432,1729,1752]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CE946FFC6FF833746F9205CE3F948" box="[1348,1432,1729,1752]" italics="true" pageId="10" pageNumber="108">Liogenys</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="308C2DF1FFC6FF83328EF9015CE2F804" pageId="10" pageNumber="108">
|
||
<paragraph id="8BC73554FFC6FF83328EF9015CE2F8A6" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EF9015CE2F8A6" pageId="10" pageNumber="108">
|
||
5. Clypeus rounded, sometimes anterior margin narrowly emarginate (
|
||
<figureCitation id="134329D1FFC6FF833184F9015A9AF968" box="[902,993,1760,1784]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">Fig. 15D</figureCitation>
|
||
); apex of both mandibles forming obtuse angle, seen in frontal view of head (
|
||
<figureCitation id="134329D1FFC6FF83302BF91E5B05F887" box="[553,638,1791,1815]" captionStart="Figure 13" captionStartId="27.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-7@27.[180,1423,144,1808]" captionTargetPageId="27" captionText="Figure 13. Head in frontoventral view (A–C) and left mandible (D–G) of some Neotropical Diplotaxini. Careocallus tehuelche (A, D); Pachrodema sp. (B–G); Liogenys suturalis (Blanchard) (C, E); Homalochilus punctatostriatus (F)." figureDoi="http://doi.org/10.5281/zenodo.11241122" httpUri="https://zenodo.org/record/11241122/files/figure.png" pageId="10" pageNumber="108">Fig. 13B</figureCitation>
|
||
); apex of sutural ridge of elytra rounded or subangulate (
|
||
<figureCitation id="134329D1FFC6FF8336CCF91E5C59F887" box="[1230,1314,1791,1815]" captionStart="Figure 14" captionStartId="28.[113,178,1624,1648]" captionTargetBox="[114,1458,144,1596]" captionTargetId="figure-123@28.[114,1458,144,1596]" captionTargetPageId="28" captionText="Figure 14. Posterior margin of elytra, variation in apex of sutural ridge among Diplotaxini.Apex angulate (A, F); apex subangulate (E) or rounded (B–D). Diplotaxis tristis Kirby (A); Pachrodema castanea Blanchard (B); Homalochilus punctatostriatus (C); Pacuvia philippiana Gutiérrez (D); Liogenys suturalis (E); Careocallus tehuelche (F)." figureDoi="http://doi.org/10.5281/zenodo.11241126" httpUri="https://zenodo.org/record/11241126/files/figure.png" pageId="10" pageNumber="108">Fig. 14B</figureCitation>
|
||
), subapical callus of elytra distinct on apex, more-or-less defined (
|
||
<figureCitation id="134329D1FFC6FF8330E1F8FF5A5FF8A6" box="[739,804,1822,1846]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="10" pageNumber="108">Fig. 7I</figureCitation>
|
||
)........................................................................................
|
||
<taxonomicName id="4C784ED7FFC6FF833722F8FF5CE2F8A6" box="[1312,1433,1822,1846]" class="Insecta" family="Scarabaeidae" genus="Pachrodema" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CE946FFC6FF833722F8FF5CE2F8A6" box="[1312,1433,1822,1846]" italics="true" pageId="10" pageNumber="108">Pachrodema</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC73554FFC6FF83328EF8DF5CE2F804" blockId="10.[140,1433,1162,1941]" pageId="10" pageNumber="108">
|
||
<keyLead id="30899661FFC6FF83328EF8DF5CE2F804" pageId="10" pageNumber="108">
|
||
-. Clypeus trapezoidal, subemarginate (
|
||
<figureCitation id="134329D1FFC6FF833039F8DF5BF7F8C6" box="[571,652,1854,1878]" captionStart="Figure 8" captionStartId="22.[113,178,1836,1860]" captionTargetBox="[165,1408,144,1808]" captionTargetId="figure-8@22.[165,1408,144,1808]" captionTargetPageId="22" captionText="Figure 8. Careocallus tehuelche. Male holotype habitus (A), head and pronotum (B), lateral (C), metatibia (D), posterior (E), and parameres in dorsal (F) and lateral (G) views." figureDoi="http://doi.org/10.5281/zenodo.11241110" httpUri="https://zenodo.org/record/11241110/files/figure.png" pageId="10" pageNumber="108">Figs. 8B</figureCitation>
|
||
,
|
||
<figureCitation id="134329D1FFC6FF83309AF8DF5BCEF8C6" box="[664,693,1854,1878]" captionStart="Figure 9" captionStartId="23.[128,193,1836,1860]" captionTargetBox="[141,1461,144,1808]" captionTargetId="figure-7@23.[141,1461,144,1808]" captionTargetPageId="23" captionText="Figure 9. Careocallus tehuelche. Female paratype habitus (A); lateral (B) and posterior (C) views; protarsal claw of female (D) and male (E)." figureDoi="http://doi.org/10.5281/zenodo.11241112" httpUri="https://zenodo.org/record/11241112/files/figure.png" pageId="10" pageNumber="108">9A</figureCitation>
|
||
,
|
||
<figureCitation id="134329D1FFC6FF8330C1F8DF5B90F8C6" box="[707,747,1854,1878]" captionStart="Figure 10" captionStartId="24.[115,180,1836,1860]" captionTargetBox="[166,1409,144,1808]" captionTargetId="figure-7@24.[166,1409,144,1808]" captionTargetPageId="24" captionText="Figure 10. Careocallus densicollis. Male holotype habitus (A); head and pronotum (B); lateral (C) and posterior (F) views; metatibia (D); protibial claw (E); parameres in dorsal (G) and lateral (H) views." figureDoi="http://doi.org/10.5281/zenodo.11241114" httpUri="https://zenodo.org/record/11241114/files/figure.png" pageId="10" pageNumber="108">10B</figureCitation>
|
||
,
|
||
<figureCitation id="134329D1FFC6FF8330FAF8DF5A5BF8C6" box="[760,800,1854,1878]" captionStart="Figure 15" captionStartId="29.[129,194,1721,1745]" captionTargetBox="[129,1473,142,1693]" captionTargetId="figure-77@29.[129,1473,142,1693]" captionTargetPageId="29" captionText="Figure 15. Mentum disc, arched transverse carina and concavity absent (A) or present (B); inner margins of both eyes parallel (C, F) or with degrees of convergence (D–E, G–H). Pacuvia castanea Curtis (A); Liogenys bidenticeps Moser (B); L. fusca Blanchard (C, F); Pachrodema castanea (D, G); Careocallus densicollis (E, H)." figureDoi="http://doi.org/10.5281/zenodo.11241128" httpUri="https://zenodo.org/record/11241128/files/figure.png" pageId="10" pageNumber="108">15E</figureCitation>
|
||
); apex of both mandibles forming acute angle, seen in frontal view of head (
|
||
<figureCitation id="134329D1FFC6FF83334FF8BC58DFF8E5" box="[333,420,1885,1909]" captionStart="Figure 13" captionStartId="27.[129,194,1836,1860]" captionTargetBox="[180,1423,144,1808]" captionTargetId="figure-7@27.[180,1423,144,1808]" captionTargetPageId="27" captionText="Figure 13. Head in frontoventral view (A–C) and left mandible (D–G) of some Neotropical Diplotaxini. Careocallus tehuelche (A, D); Pachrodema sp. (B–G); Liogenys suturalis (Blanchard) (C, E); Homalochilus punctatostriatus (F)." figureDoi="http://doi.org/10.5281/zenodo.11241122" httpUri="https://zenodo.org/record/11241122/files/figure.png" pageId="10" pageNumber="108">Fig. 13A</figureCitation>
|
||
); apex of sutural ridge of elytra sharp, right or obtuse-angulate (
|
||
<figureCitation id="134329D1FFC6FF833637F8BC5DF3F8E5" box="[1077,1160,1885,1909]" captionStart="Figure 14" captionStartId="28.[113,178,1624,1648]" captionTargetBox="[114,1458,144,1596]" captionTargetId="figure-123@28.[114,1458,144,1596]" captionTargetPageId="28" captionText="Figure 14. Posterior margin of elytra, variation in apex of sutural ridge among Diplotaxini.Apex angulate (A, F); apex subangulate (E) or rounded (B–D). Diplotaxis tristis Kirby (A); Pachrodema castanea Blanchard (B); Homalochilus punctatostriatus (C); Pacuvia philippiana Gutiérrez (D); Liogenys suturalis (E); Careocallus tehuelche (F)." figureDoi="http://doi.org/10.5281/zenodo.11241126" httpUri="https://zenodo.org/record/11241126/files/figure.png" pageId="10" pageNumber="108">Fig. 14F</figureCitation>
|
||
), subapical callus of elytra absent (
|
||
<figureCitation id="134329D1FFC6FF83330CF89D5823F804" box="[270,344,1916,1940]" captionStart="Figure 9" captionStartId="23.[128,193,1836,1860]" captionTargetBox="[141,1461,144,1808]" captionTargetId="figure-7@23.[141,1461,144,1808]" captionTargetPageId="23" captionText="Figure 9. Careocallus tehuelche. Female paratype habitus (A); lateral (B) and posterior (C) views; protarsal claw of female (D) and male (E)." figureDoi="http://doi.org/10.5281/zenodo.11241112" httpUri="https://zenodo.org/record/11241112/files/figure.png" pageId="10" pageNumber="108">Fig. 9C</figureCitation>
|
||
) ............................................................................................................................................................................
|
||
<taxonomicName id="4C784ED7FFC6FF83372AF89D5CE2F804" box="[1320,1433,1916,1940]" class="Insecta" family="Scarabaeidae" genus="Careocallus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CE946FFC6FF83372AF89D5CE2F804" box="[1320,1433,1916,1940]" italics="true" pageId="10" pageNumber="108">Careocallus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |