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<document id="D343C7D7A837B8FD4DC199EECABF65E5" ID-CLB-Dataset="2380" ID-DOI="10.5281/zenodo.5739643" ID-GBIF-Dataset="f9439543-76ca-43e9-aca5-0a8d3899a2b1" ID-ISSN="0374-1036" ID-Zenodo-Dep="5739643" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638252251656" checkinUser="marcus" docAuthor="Makranczy, György" docDate="2013" docId="03CEDF09FFFFF34BD15F92E9D9FDF9D5" docLanguage="en" docName="ActaEntMusNatPra.2013.53.1.177-208.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinodromus nigerius Gildenkov 2000" docType="treatment" docVersion="8" lastPageNumber="188" masterDocId="FFF7A771FFF4F340D0219333DB72FFF2" masterDocTitle="Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="208" masterPageNumber="177" pageNumber="188" updateTime="1699471184067" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E64B52199B40CF1A752019FCA9F442B5">Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:roleTerm id="0016F026A827A8BB56839C5089CDAA01">Author</mods:roleTerm>
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<mods:namePart id="2500F9C381D61F6456E4F3A4DFF77990">Makranczy, György</mods:namePart>
<mods:affiliation id="D41E0CAC4AAD75B737D8460FDA8E81C9">Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross utca 13, Hungary</mods:affiliation>
<mods:nameIdentifier id="0E5F61FB47504F5EDD50BFC4C99DC35A" type="email">makranczy@zoo.nhmus.hu</mods:nameIdentifier>
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<mods:title id="175486A7C239A92A065800D9D96D1081">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="ECEB790DD1E21DC90624A3345789F282">2013</mods:date>
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<mods:number id="B348EF16FC5BEF29139A1CF0530D51A6">2013-07-15</mods:number>
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<mods:number id="9637908458428B7C6CFFE662C668DE2B">53</mods:number>
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<mods:identifier id="D160EDA8B8AB26BC98FA989353F8AF5E" type="DOI">10.5281/zenodo.5739643</mods:identifier>
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<treatment id="03CEDF09FFFFF34BD15F92E9D9FDF9D5" ID-DOI="http://doi.org/10.5281/zenodo.5911273" ID-GBIF-Taxon="192662469" ID-Zenodo-Dep="5911273" LSID="urn:lsid:plazi:treatment:03CEDF09FFFFF34BD15F92E9D9FDF9D5" httpUri="http://treatment.plazi.org/id/03CEDF09FFFFF34BD15F92E9D9FDF9D5" lastPageNumber="188" pageId="11" pageNumber="188">
<subSubSection id="C37D3D94FFFFF34BD15F92E9D841FE01" box="[382,819,474,499]" pageId="11" pageNumber="188" type="nomenclature">
<paragraph id="8BD86E1FFFFFF34BD15F92E9D841FE01" blockId="11.[382,819,474,499]" box="[382,819,474,499]" pageId="11" pageNumber="188">
<heading id="D090D973FFFFF34BD15F92E9D841FE01" bold="true" box="[382,819,474,499]" centered="true" fontSize="8" level="2" pageId="11" pageNumber="188" reason="3">
<taxonomicName id="4C67159CFFFFF34BD15F92E9D841FE01" ID-CoL="56H3L" authority="Gildenkov, 2000" authorityName="Gildenkov" authorityYear="2000" box="[382,819,474,499]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD15F92E9D841FE01" bold="true" box="[382,819,474,499]" pageId="11" pageNumber="188">
<emphasis id="B913B20DFFFFF34BD15F92E9D907FE01" bold="true" box="[382,629,474,499]" italics="true" pageId="11" pageNumber="188">Thinodromus nigerius</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD25D92E8D841FE01" author="GILDENKOV M. &amp; YU" box="[636,819,475,499]" pageId="11" pageNumber="188" pagination="43 - 56" refId="ref14247" refString="GILDENKOV M. YU. 2000: Novyy podrod i sem' novykh dlya nauki vidov roda Thinodromus iz efiopskoy zoogeograficheskoy oblast'i (Coleoptera, Staphylinidae). [New subgenus and seven species, new to science, of the genus Thinodromus from Ethiopian zoogeographic region (Coleoptera, Staphylinidae)]. Izvestiya Khar ' kovskogo Entomologicheskogo Obshchestva [Kharkov Entomological Society Gazette] 8 (1): 43 - 56 (in Russian)." type="journal article" year="2000">Gildenkov, 2000</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFFF34BD1D392C9D9CDFDFF" box="[498,703,506,525]" pageId="11" pageNumber="188" type="description">
<paragraph id="8BD86E1FFFFFF34BD1D392C9D9CDFDFF" blockId="11.[113,703,506,613]" box="[498,703,506,525]" pageId="11" pageNumber="188">
(
<figureCitation id="135C729AFFFFF34BD1D992C9D929FDFF" box="[504,603,506,525]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="11" pageNumber="188">Figs 13–16</figureCitation>
,
<figureCitation id="135C729AFFFFF34BD24292C9D9E9FDFF" box="[611,667,506,525]" captionStart="Figs 70–76. 70–74" captionStartId="23.[113,150,1492,1511]" captionTargetBox="[115,1063,221,1431]" captionTargetId="figure-10@23.[113,1065,219,1457]" captionTargetPageId="23" captionText="Figs 70–76. 70–74 – Thinodromus capensis (Bernhauer, 1934), male (70 – sternite VIII; 71 – tergite X; 72–73 – aedeagus; 74 – paramere); 75–76 – T. nigerius Gildenkov, 2000, male (75 – sclerotized subapical plate of aedeagus; 76 – apical copulatory sclerite).Abbreviations:AC – apical copulatory sclerite; BA – basal sclerites, BM – basomedial sclerites, MA – medioapical sclerites ML – medial lamellae. Scale bar: 0.05 mm for 76, 0.08 mm for 75, 0.11 mm for 72–74, 0.12 mm for 71, 0.14 mm for 70." figureDoi="http://doi.org/10.5281/zenodo.5739677" httpUri="https://zenodo.org/record/5739677/files/figure.png" pageId="11" pageNumber="188">75–76</figureCitation>
,
<figureCitation id="135C729AFFFFF34BD28292C9D9CBFDFF" box="[675,697,506,525]" captionStart="Figs 81–85. 81–83" captionStartId="25.[113,150,1532,1551]" captionTargetBox="[128,1073,222,1462]" captionTargetId="figure-10@25.[124,1076,219,1466]" captionTargetPageId="25" captionText="Figs 81–85. 81–83 – female ringstructures.81 – Thinodromus tibialis (Fauvel, 1907); 82 – T. dasys Gildenkov, 2000; 83 – T. nigerius Gildenkov, 2000. 84–85 – female ringstructures with spermathecae. 84 – T. facilis Gildenkov, 2000; 85 – T. gildenkovi sp. nov. Scale bar: 0.10 mm for 85, 0.12 mm for 81 and 83–84, 0.14 mm for 82." figureDoi="http://doi.org/10.5281/zenodo.5739681" httpUri="https://zenodo.org/record/5739681/files/figure.png" pageId="11" pageNumber="188">83</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFFF34BD050912ED976FD96" pageId="11" pageNumber="188" type="reference_group">
<treatmentCitationGroup id="AB774931FFFFF34BD050912ED976FD96" pageId="11" pageNumber="188">
<paragraph id="8BD86E1FFFFFF34BD050912ED91FFDDD" blockId="11.[113,703,506,613]" box="[113,621,540,559]" pageId="11" pageNumber="188">
<treatmentCitation id="0AC6480EFFFFF34BD050912ED91BFDDD" author="GILDENKOV M. &amp; YU" box="[113,617,540,559]" page="54" pageId="11" pageNumber="188" year="2000">
<taxonomicName id="4C67159CFFFFF34BD050912ED91BFDDD" ID-CoL="56H3L" authority="Gildenkov, 2000: 54" authorityName="Gildenkov" authorityPageNumber="54" authorityYear="2000" box="[113,617,540,559]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius" subGenus="Thinodromus">
<emphasis id="B913B20DFFFFF34BD050912EDB90FDDD" box="[113,226,541,559]" italics="true" pageId="11" pageNumber="188">Thinodromus</emphasis>
(
<emphasis id="B913B20DFFFFF34BD0CE912EDA12FDDD" box="[239,352,541,559]" italics="true" pageId="11" pageNumber="188">Thinodromus</emphasis>
)
<emphasis id="B913B20DFFFFF34BD14D912EDAC0FDDD" box="[364,434,541,559]" italics="true" pageId="11" pageNumber="188">nigerius</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD196912FD91BFDDD" author="GILDENKOV M. &amp; YU" box="[439,617,540,559]" pageId="11" pageNumber="188" pagination="43 - 56" refId="ref14247" refString="GILDENKOV M. YU. 2000: Novyy podrod i sem' novykh dlya nauki vidov roda Thinodromus iz efiopskoy zoogeograficheskoy oblast'i (Coleoptera, Staphylinidae). [New subgenus and seven species, new to science, of the genus Thinodromus from Ethiopian zoogeographic region (Coleoptera, Staphylinidae)]. Izvestiya Khar ' kovskogo Entomologicheskogo Obshchestva [Kharkov Entomological Society Gazette] 8 (1): 43 - 56 (in Russian)." type="journal article" year="2000">Gildenkov, 2000: 54</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</paragraph>
<paragraph id="8BD86E1FFFFFF34BD050910BD90BFDB8" blockId="11.[113,703,506,613]" box="[113,633,567,586]" pageId="11" pageNumber="188">
<treatmentCitation id="0AC6480EFFFFF34BD050910BD953FDB8" author="MAKRANCZY GY" box="[113,545,567,586]" page="44" pageId="11" pageNumber="188" year="2009">
<taxonomicName id="4C67159CFFFFF34BD050910BD953FDB8" ID-CoL="8YBSL" authority="Makranczy, 2009: 44" authorityName="Makranczy" authorityPageNumber="44" authorityYear="2009" box="[113,545,567,586]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="kedougouensis">
<emphasis id="B913B20DFFFFF34BD050910BDA14FDB8" box="[113,358,568,586]" italics="true" pageId="11" pageNumber="188">Thinodromus kedougouensis</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD14A9104D953FDB8" author="MAKRANCZY GY" box="[363,545,567,586]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">Makranczy, 2009: 44</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B913B20DFFFFF34BD20B9104D90BFDB8" bold="true" box="[554,633,567,586]" pageId="11" pageNumber="188">
<taxonomicNameLabel id="A2200F76FFFFF34BD20B9104D90BFDB8" box="[554,633,567,586]" pageId="11" pageNumber="188">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD86E1FFFFFF34BD0509161D976FD96" blockId="11.[113,703,506,613]" box="[113,516,593,613]" pageId="11" pageNumber="188">
<treatmentCitation id="0AC6480EFFFFF34BD0509161D972FD96" author="MAKRANCZY GY" box="[113,512,593,613]" page=")" pageId="11" pageNumber="188" year="2009">
<taxonomicName id="4C67159CFFFFF34BD0509161D972FD96" ID-CoL="56H3L" authority=": MAKRANCZY (2009: 53)" authorityName="MAKRANCZY" authorityPageNumber="53" authorityYear="2009" box="[113,512,593,613]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD0509161DA5CFD96" box="[113,302,594,612]" italics="true" pageId="11" pageNumber="188">Thinodromus nigerius</emphasis>
:
<bibRefCitation id="EFF613EEFFFFF34BD1189161D972FD96" author="MAKRANCZY GY" box="[313,512,593,613]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">MAKRANCZY (2009: 53)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection id="C37D3D94FFFFF34BD0509149D9FDF9D5" pageId="11" pageNumber="188" type="materials_examined">
<paragraph id="8BD86E1FFFFFF34BD0509149DAE3FC8C" blockId="11.[113,1089,634,1576]" pageId="11" pageNumber="188">
<emphasis id="B913B20DFFFFF34BD0509149DA56FD7C" bold="true" box="[113,292,634,654]" pageId="11" pageNumber="188">Material examined.</emphasis>
<materialsCitation id="3B0F6442FFFFF34BD10B9149DB97FD5B" ID-GBIF-Occurrence="3458217328" collectingDate="1988-04-28" collectionCode="SMNS" collectorName="F. - T. Krell" country="Ivory Coast" latitude="9.583333" location="Kafolo" longLatPrecision="1299" longitude="-4.3" pageId="11" pageNumber="188" specimenCount="1" specimenCount-female="1" stateProvince="Comoe">
<emphasis id="B913B20DFFFFF34BD10B9149DAAAFD7C" bold="true" box="[298,472,634,654]" pageId="11" pageNumber="188">
<collectingCountry id="F3702E8FFFFFF34BD10B9149DAA6FD7C" box="[298,468,634,654]" name="Ivory Coast" pageId="11" pageNumber="188">CÔTE D’IVOIRE</collectingCountry>
:
</emphasis>
<location id="8EB838C4FFFFF34BD1FF9148D969FD7C" LSID="urn:lsid:plazi:treatment:03CEDF09FFFFF34BD15F92E9D9FDF9D5:8EB838C4FFFFF34BD1FF9148D969FD7C" box="[478,539,635,654]" country="Ivory Coast" latitude="9.583333" longLatPrecision="1299" longitude="-4.3" name="Kafolo" pageId="11" pageNumber="188" stateProvince="Comoe">Kafolo</location>
,
<collectingRegion id="49A3A0FDFFFFF34BD2059148D910FD7C" box="[548,610,635,654]" country="Ivory Coast" name="Comoe" pageId="11" pageNumber="188">Comoé</collectingRegion>
(river) [
<geoCoordinate id="EE5308D8FFFFF34BD28B9148D986FD7D" box="[682,756,634,655]" degrees="09" direction="north" minutes="35" orientation="latitude" pageId="11" pageNumber="188" precision="925" value="9.583333">09°35′N</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD2DF9148D83CFD7D" box="[766,846,634,655]" degrees="04" direction="west" minutes="18" orientation="longitude" pageId="11" pageNumber="188" precision="925" value="-4.3">04°18′W</geoCoordinate>
],
<date id="FFD948DFFFFFF34BD37C9148D8CBFD7C" box="[861,953,635,654]" pageId="11" pageNumber="188" value="1988-04-28">
<collectingDate id="EF9DB137FFFFF34BD37C9148D8CBFD7C" box="[861,953,635,654]" pageId="11" pageNumber="188" value="1988-04-28">28.iv.1988</collectingDate>
</date>
, leg.
<collectorName id="26920BC9FFFFF34BD3C6914FDF33FD7C" box="[999,1089,635,655]" pageId="11" pageNumber="188">F.-T. Krell</collectorName>
(
<specimenCount id="9D61A596FFFFF34BD05B91A5DBEAFD5B" box="[122,152,656,681]" pageId="11" pageNumber="188" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD08091A5DBADFD5B" box="[161,223,662,681]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="11" pageNumber="188">SMNS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0F6442FFFFF34BD0CD91A6D881FD5B" ID-GBIF-Occurrence="3458217301" box="[236,1011,656,681]" collectingDate="1967-07-06" collectionCode="MHNG" collectorName="S. Endrody-Younga" country="Ghana" latitude="6.6833334" location="Kumasi" longLatPrecision="1304" longitude="-1.6166667" municipality="Kumasi" pageId="11" pageNumber="188" specimenCount="1" specimenCount-female="1">
<emphasis id="B913B20DFFFFF34BD0CD91A6DA31FD5B" bold="true" box="[236,323,661,681]" pageId="11" pageNumber="188">
<collectingCountry id="F3702E8FFFFFF34BD0CD91A6DA4DFD5B" box="[236,319,661,681]" name="Ghana" pageId="11" pageNumber="188">GHANA</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BBCF465FFFFF34BD16691A5DAFBFD5B" box="[327,393,662,681]" country="Ghana" latitude="6.6833334" longLatPrecision="1304" longitude="-1.6166667" name="Kumasi" pageId="11" pageNumber="188">Kumasi</collectingMunicipality>
[
<geoCoordinate id="EE5308D8FFFFF34BD1B291A5DAACFD5B" box="[403,478,661,681]" degrees="06" direction="north" minutes="41" orientation="latitude" pageId="11" pageNumber="188" precision="925" value="6.6833334">06°41′N</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD1C791A5D944FD5B" box="[486,566,661,681]" degrees="01" direction="west" minutes="37" orientation="longitude" pageId="11" pageNumber="188" precision="925" value="-1.6166667">01°37′W</geoCoordinate>
],
<date id="FFD948DFFFFFF34BD26291A5D9EEFD5B" box="[579,668,662,681]" pageId="11" pageNumber="188" value="1967-07-06">
<collectingDate id="EF9DB137FFFFF34BD26291A5D9EEFD5B" box="[579,668,662,681]" pageId="11" pageNumber="188" value="1967-07-06">6.vii.1967</collectingDate>
</date>
, leg.
<collectorName id="26920BC9FFFFF34BD2E991A5D802FD5B" box="[712,880,661,681]" pageId="11" pageNumber="188">S. Endrődy-Younga</collectorName>
(
<specimenCount id="9D61A596FFFFF34BD35D91A5D8EBFD5B" box="[892,921,656,681]" pageId="11" pageNumber="188" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD38591A5D89BFD5B" box="[932,1001,662,681]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="188" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B0F6442FFFFF34BD3D791A6D911FD2C" ID-GBIF-Occurrence="3458217326" collectingDate="2006-02-07" collectionCode="NHMW" collectorName="Goergen &amp; Komarek &amp; Houngue" country="Benin" latitude="7.2012496" location="Abomey - Cove" longLatPrecision="1" longitude="2.2888334" pageId="11" pageNumber="188" specimenCount="1" specimenCount-female="1" stateProvince="Zou">
<emphasis id="B913B20DFFFFF34BD3D791A6DF33FD5B" bold="true" box="[1014,1089,661,681]" pageId="11" pageNumber="188">
<collectingCountry id="F3702E8FFFFFF34BD3D791A6DF4FFD5B" box="[1014,1085,661,681]" name="Benin" pageId="11" pageNumber="188">BÉNIN</collectingCountry>
:
</emphasis>
<collectingRegion id="49A3A0FDFFFFF34BD0509182DA74FD36" box="[113,262,689,708]" country="Benin" name="Zou" pageId="11" pageNumber="188">Departement Zou</collectingRegion>
,
<collectorName id="26920BC9FFFFF34BD12C9182DA2CFD36" box="[269,350,689,708]" pageId="11" pageNumber="188">Za-Kpota</collectorName>
&amp;
<collectorName id="26920BC9FFFFF34BD1579182DAD7FD36" box="[374,421,689,708]" pageId="11" pageNumber="188">Covè</collectorName>
,
<collectingRegion id="49A3A0FDFFFFF34BD18D9182DABCFD36" box="[428,462,689,708]" country="Benin" name="Zou" pageId="11" pageNumber="188">Zou</collectingRegion>
riv., bridge at road
<location id="8EB838C4FFFFF34BD24A9183D980FD36" LSID="urn:lsid:plazi:treatment:03CEDF09FFFFF34BD15F92E9D9FDF9D5:8EB838C4FFFFF34BD24A9183D980FD36" box="[619,754,688,708]" country="Benin" latitude="7.2012496" longLatPrecision="1" longitude="2.2888334" name="Abomey - Cove" pageId="11" pageNumber="188" stateProvince="Zou">Abomey - Covè</location>
,
<geoCoordinate id="EE5308D8FFFFF34BD2D99182D81CFD36" box="[760,878,688,708]" degrees="07" direction="north" minutes="12" orientation="latitude" pageId="11" pageNumber="188" precision="1" seconds="04.5" value="7.2012496">07°12′04.5″N</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD3579182D89BFD36" box="[886,1001,688,708]" degrees="02" direction="east" minutes="17" orientation="longitude" pageId="11" pageNumber="188" precision="1" seconds="19.8" value="2.2888334">02°17′19.8″E</geoCoordinate>
,
<date id="FFD948DFFFFFF34BD3D19183DF4FFD31" box="[1008,1085,688,707]" pageId="11" pageNumber="188" value="2006-02-07">
<collectingDate id="EF9DB137FFFFF34BD3D19183DF4FFD31" box="[1008,1085,688,707]" pageId="11" pageNumber="188" value="2006-02-07">7.ii.2006</collectingDate>
</date>
, leg.
<collectorName id="26920BC9FFFFF34BD0B691F8DB90FD2C" box="[151,226,715,734]" pageId="11" pageNumber="188">Goergen</collectorName>
,
<collectorName id="26920BC9FFFFF34BD0CA91F8DA48FD2C" box="[235,314,715,734]" pageId="11" pageNumber="188">Komarek</collectorName>
&amp;
<collectorName id="26920BC9FFFFF34BD17691F8DAD7FD2C" box="[343,421,715,734]" pageId="11" pageNumber="188">Hounguè</collectorName>
(3 ♂♂
<specimenCount id="9D61A596FFFFF34BD1C691F8D97AFD2C" box="[487,520,709,734]" pageId="11" pageNumber="188" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD23291F8D92FFD2C" box="[531,605,715,734]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="188" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0F6442FFFFF34BD24A91F9D925FD0B" ID-GBIF-Occurrence="3458217314" collectingDate="1998-10-01" collectingDateMax="1998-11-02" collectingDateMin="1998-10-01" collectionCode="CMNH" collectorName="R. D. Ward" country="Nigeria" latitude="9.066667" location="Abuja" longLatPrecision="1300" longitude="7.483333" pageId="11" pageNumber="188" specimenCount="1" specimenCount-female="1" stateProvince="Federal Capital Territory">
<emphasis id="B913B20DFFFFF34BD24A91F9D9A2FD2C" bold="true" box="[619,720,714,734]" pageId="11" pageNumber="188">
<collectingCountry id="F3702E8FFFFFF34BD24A91F9D9BEFD2C" box="[619,716,714,734]" name="Nigeria" pageId="11" pageNumber="188">NIGERIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49A3A0FDFFFFF34BD2F791F8D8DEFD2C" box="[726,940,715,734]" country="Nigeria" name="Federal Capital Territory" pageId="11" pageNumber="188">Federal Capital Territory</collectingRegion>
,
<location id="8EB838C4FFFFF34BD39291F8D895FD2C" LSID="urn:lsid:plazi:treatment:03CEDF09FFFFF34BD15F92E9D9FDF9D5:8EB838C4FFFFF34BD39291F8D895FD2C" box="[947,999,715,734]" country="Nigeria" latitude="9.066667" longLatPrecision="1300" longitude="7.483333" name="Abuja" pageId="11" pageNumber="188" stateProvince="Federal Capital Territory">Abuja</location>
[
<geoCoordinate id="EE5308D8FFFFF34BD3D391F8DF4FFD2D" box="[1010,1085,714,735]" degrees="09" direction="north" minutes="04" orientation="latitude" pageId="11" pageNumber="188" precision="925" value="9.066667">09°04′N</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD05091D5DBCBFD0B" box="[113,185,741,761]" degrees="07" direction="east" minutes="29" orientation="longitude" pageId="11" pageNumber="188" precision="925" value="7.483333">07°29′E</geoCoordinate>
],
<date id="FFD948DFFFFFF34BD0E891D5DA14FD0B" box="[201,358,742,761]" pageId="11" pageNumber="188" value="1998-11-02">
<collectingDate id="EF9DB137FFFFF34BD0E891D5DA14FD0B" box="[201,358,742,761]" pageId="11" pageNumber="188" value="1998-11-02">2 November 1998</collectingDate>
</date>
,
<collectorName id="26920BC9FFFFF34BD14E91D5DAA3FD0B" box="[367,465,742,761]" pageId="11" pageNumber="188">R. D. Ward</collectorName>
(1 ♂,
<collectionCode id="ED76F6DAFFFFF34BD22B91D5D93FFD0B" box="[522,589,742,761]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="11" pageNumber="188" type="Museum">CMNH</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B0F6442FFFFF34BD27D91D5D8B8FD0B" ID-GBIF-Occurrence="3458217332" box="[604,970,736,761]" collectingDate="1998-10-01" collectingDateMax="1998-11-02" collectingDateMin="1998-10-01" collectionCode="CMNH" collectorName="R. D. Ward" country="Nigeria" latitude="9.066667" location="Abuja" longLatPrecision="1300" longitude="7.483333" pageId="11" pageNumber="188" specimenCount="1" specimenCount-female="1" stateProvince="Federal Capital Territory">
same but
<date id="FFD948DFFFFFF34BD28E91D5D835FD0B" box="[687,839,742,761]" pageId="11" pageNumber="188" value="1998-10-01" valueMax="1998-10-07" valueMin="1998-10-01">
<collectingDate id="EF9DB137FFFFF34BD28E91D5D835FD0B" box="[687,839,742,761]" pageId="11" pageNumber="188" value="1998-10-01" valueMax="1998-10-07" valueMin="1998-10-01">1-7 October 1998</collectingDate>
</date>
(
<specimenCount id="9D61A596FFFFF34BD37791D5D807FD0B" box="[854,885,736,761]" pageId="11" pageNumber="188" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD3A191D5D8B6FD0B" box="[896,964,742,761]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="11" pageNumber="188" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0F6442FFFFF34BD3F391D6DF33FCDC" ID-GBIF-Occurrence="3458217337" collectingDate="1985-12-12" collectingDateMax="1996-03-07" collectingDateMin="1985-12-12" collectionCode="TMSA, SMNS, FMNH" collectorName="S. Endrody-Younga " country="South Africa" elevation="270" latitude="-24.983334" location="Skukuza" longLatPrecision="1248" longitude="31.6" pageId="11" pageNumber="188" specimenCount="4" specimenCount-female="4" stateProvince="Limpopo">
<emphasis id="B913B20DFFFFF34BD3F391D6DA4FFCE6" bold="true" pageId="11" pageNumber="188">
<collectingCountry id="F3702E8FFFFFF34BD3F391D6DA4BFCE6" name="South Africa" pageId="11" pageNumber="188">REPUBLIC OF SOUTH AFRICA</collectingCountry>
:
</emphasis>
<collectorName id="26920BC9FFFFF34BD1629032DAD8FCE1" box="[323,426,768,788]" pageId="11" pageNumber="188">N Transvaal</collectorName>
[=
<collectingRegion id="49A3A0FDFFFFF34BD1E09033D93AFCE6" box="[449,584,768,788]" country="South Africa" name="Limpopo" pageId="11" pageNumber="188">Limpopo prov.</collectingRegion>
],
<collectorName id="26920BC9FFFFF34BD2749032D9CFFCE6" box="[597,701,769,788]" pageId="11" pageNumber="188">Kruger N.P.</collectorName>
,
<location id="8EB838C4FFFFF34BD2E49033D87CFCE1" LSID="urn:lsid:plazi:treatment:03CEDF09FFFFF34BD15F92E9D9FDF9D5:8EB838C4FFFFF34BD2E49033D87CFCE1" box="[709,782,768,787]" country="South Africa" latitude="-24.983334" longLatPrecision="1248" longitude="31.6" name="Skukuza" pageId="11" pageNumber="188" stateProvince="Limpopo">Skukuza</location>
research camp [Sabie river,
<quantity id="4C9FC3FAFFFFF34BD4259032DF4AFCE6" box="[1028,1080,769,788]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="188" unit="m" value="270.0">
<elevation id="004A892CFFFFF34BD4259032DF4AFCE6" box="[1028,1080,769,788]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="188" unit="m" value="270.0">270m</elevation>
</quantity>
],
<geoCoordinate id="EE5308D8FFFFF34BD0509028DBC4FCDC" box="[113,182,794,814]" degrees="24" direction="south" minutes="59" orientation="latitude" pageId="11" pageNumber="188" precision="925" value="-24.983334">24°59′S</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD0E19028DA75FCDD" box="[192,263,794,815]" degrees="31" direction="east" minutes="35" orientation="longitude" pageId="11" pageNumber="188" precision="925" value="31.583334">31°35′E</geoCoordinate>
,
<date id="FFD948DFFFFFF34BD1319028DA11FCDC" box="[272,355,795,814]" pageId="11" pageNumber="188" value="1996-03-07">
<collectingDate id="EF9DB137FFFFF34BD1319028DA11FCDC" box="[272,355,795,814]" pageId="11" pageNumber="188" value="1996-03-07">7.iii.1996</collectingDate>
</date>
, leg.
<collectorName id="26920BC9FFFFF34BD1B09028D948FCDC" box="[401,570,794,814]" pageId="11" pageNumber="188">S. Endrődy-Younga</collectorName>
(E-Y 3220), UV light (1 ♂
<specimenCount id="9D61A596FFFFF34BD3089028D82BFCDC" box="[809,857,789,814]" pageId="11" pageNumber="188" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD3429028D8D3FCDC" box="[867,929,795,814]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="11" pageNumber="188">TMSA</collectionCode>
; 1 ♂
<specimenCount id="9D61A596FFFFF34BD3EE9028D89DFCDC" box="[975,1007,789,814]" pageId="11" pageNumber="188" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD3DB9028DF45FCDC" box="[1018,1079,795,814]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="11" pageNumber="188">SMNS</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B0F6442FFFFF34BD0509005D833FCBB" ID-GBIF-Occurrence="3458217318" box="[113,833,817,842]" collectingDate="1985-12-12" collectingDateMax="1996-03-07" collectingDateMin="1985-12-12" collectionCode="TMSA, SMNS, FMNH" country="South Africa" elevation="270" latitude="-24.983334" location="Skukuza" longLatPrecision="1248" longitude="31.6" pageId="11" pageNumber="188" specimenCount="4" specimenCount-female="4" stateProvince="Limpopo">
same but
<geoCoordinate id="EE5308D8FFFFF34BD0E79005DA79FCBB" box="[198,267,821,841]" degrees="25" direction="south" minutes="00" orientation="latitude" pageId="11" pageNumber="188" precision="925" value="-25.0">25°00′S</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD1379005DA2FFCBB" box="[278,349,821,841]" degrees="31" direction="east" minutes="35" orientation="longitude" pageId="11" pageNumber="188" precision="925" value="31.583334">31°35′E</geoCoordinate>
,
<date id="FFD948DFFFFFF34BD1469005DACFFCBB" box="[359,445,822,841]" pageId="11" pageNumber="188" value="1995-02-19">
<collectingDate id="EF9DB137FFFFF34BD1469005DACFFCBB" box="[359,445,822,841]" pageId="11" pageNumber="188" value="1995-02-19">19.ii.1995</collectingDate>
</date>
(E-Y 3102), UV light &amp; trap (1 ♂,
<collectionCode id="ED76F6DAFFFFF34BD2D99005D845FCBB" box="[760,823,822,841]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="11" pageNumber="188">TMSA</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B0F6442FFFFF34BD3669005DAFFFC8C" ID-GBIF-Occurrence="3458217306" collectingDate="1985-12-12" collectingDateMax="1996-03-07" collectingDateMin="1985-12-12" collectionCode="TMSA, SMNS, FMNH" collectorName="S. Peck" country="South Africa" elevation="270" latitude="-24.983334" location="Skukuza" longLatPrecision="1248" longitude="31.6" pageId="11" pageNumber="188" specimenCount="4" specimenCount-female="4" stateProvince="Limpopo">
<collectorName id="26920BC9FFFFF34BD3669005D8B4FCBB" box="[839,966,822,841]" pageId="11" pageNumber="188">Rep. S. Africa</collectorName>
:
<collectorName id="26920BC9FFFFF34BD3EE9005DF4CFCBB" box="[975,1086,822,841]" pageId="11" pageNumber="188">E. Transvaal</collectorName>
,
<collectorName id="26920BC9FFFFF34BD0509062DBA0FC91" box="[113,210,848,868]" pageId="11" pageNumber="188">Kruger Pk.</collectorName>
,
<collectorName id="26920BC9FFFFF34BD0F89063DA50FC91" box="[217,290,848,867]" pageId="11" pageNumber="188">Skukuza</collectorName>
[
<geoCoordinate id="EE5308D8FFFFF34BD10D9063DA01FC96" box="[300,371,848,868]" degrees="24" direction="south" minutes="59" orientation="latitude" pageId="11" pageNumber="188" precision="925" value="-24.983334">24°59′S</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFFF34BD15D9063DAB6FC96" box="[380,452,848,868]" degrees="31" direction="east" minutes="36" orientation="longitude" pageId="11" pageNumber="188" precision="925" value="31.6">31°36′E</geoCoordinate>
],
<date id="FFD948DFFFFFF34BD1F39063D927FC91" box="[466,597,848,867]" pageId="11" pageNumber="188" value="1985-12-12" valueMax="1985-12-14" valueMin="1985-12-12">
<collectingDate id="EF9DB137FFFFF34BD1F39063D927FC91" box="[466,597,848,867]" pageId="11" pageNumber="188" value="1985-12-12" valueMax="1985-12-14" valueMin="1985-12-12">12–14.xii.1985</collectingDate>
</date>
, FMHD #85-848, leg.
<collectorName id="26920BC9FFFFF34BD3379062D827FC91" box="[790,853,848,868]" pageId="11" pageNumber="188">S. Peck</collectorName>
(P# 85-281), thorn scrub &amp;
<collectingMethod id="52261608FFFFF34BD0509058DA7CFC8D" box="[113,270,875,895]" pageId="11" pageNumber="188">riverine light traps</collectingMethod>
(
<specimenCount id="9D61A596FFFFF34BD13C9058DA49FC8C" box="[285,315,869,894]" pageId="11" pageNumber="188" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFFF34BD1679058DAFAFC8C" box="[326,392,875,894]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="188" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD86E1FFFFFF34BD05090BCDA1FFA35" blockId="11.[113,1089,634,1576]" pageId="11" pageNumber="188">
<emphasis id="B913B20DFFFFF34BD05090BCDA40FC55" bold="true" box="[113,306,911,935]" pageId="11" pageNumber="188">Taxonomic note.</emphasis>
In 2009, I attempted to interpret the identity of
<taxonomicName id="4C67159CFFFFF34BD36F90A3DA52FC35" authority="Gildenkov, 2000" authorityName="Gildenkov" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD36F90A3DF32FC55" box="[846,1088,912,935]" italics="true" pageId="11" pageNumber="188">Thinodromus nigerius</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD050909CDA52FC35" author="GILDENKOV M. &amp; YU" box="[113,288,943,967]" pageId="11" pageNumber="188" pagination="43 - 56" refId="ref14247" refString="GILDENKOV M. YU. 2000: Novyy podrod i sem' novykh dlya nauki vidov roda Thinodromus iz efiopskoy zoogeograficheskoy oblast'i (Coleoptera, Staphylinidae). [New subgenus and seven species, new to science, of the genus Thinodromus from Ethiopian zoogeographic region (Coleoptera, Staphylinidae)]. Izvestiya Khar ' kovskogo Entomologicheskogo Obshchestva [Kharkov Entomological Society Gazette] 8 (1): 43 - 56 (in Russian)." type="journal article" year="2000">Gildenkov, 2000</bibRefCitation>
</taxonomicName>
based on
<specimenCount id="9D61A596FFFFF34BD1A79083DA94FC35" box="[390,486,943,967]" pageId="11" pageNumber="188" type="male">two male</specimenCount>
specimens from
<collectingCountry id="F3702E8FFFFFF34BD2B7909CD850FC35" box="[662,802,943,967]" name="Ivory Coast" pageId="11" pageNumber="188">Côte d’Ivoire</collectingCountry>
and
<collectingCountry id="F3702E8FFFFFF34BD372909CD8D1FC35" box="[851,931,943,967]" name="Nigeria" pageId="11" pageNumber="188">Nigeria</collectingCountry>
(see redescription in
<bibRefCitation id="EFF613EEFFFFF34BD09A90E3DAD0FC15" author="MAKRANCZY GY" box="[187,418,975,999]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">MAKRANCZY 2009: 53</bibRefCitation>
). Immediately after the publication of the paper, I received two more
<taxonomicName id="4C67159CFFFFF34BD08E90C3DA4DFBF5" authorityName="Kraatz" authorityYear="1857" box="[175,319,1008,1031]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B913B20DFFFFF34BD08E90C3DA4DFBF5" box="[175,319,1008,1031]" italics="true" pageId="11" pageNumber="188">Thinodromus</emphasis>
</taxonomicName>
specimens from
<collectingCountry id="F3702E8FFFFFF34BD1D490DCD938FBF5" box="[501,586,1007,1031]" name="Nigeria" pageId="11" pageNumber="188">Nigeria</collectingCountry>
, a male (
<figureCitation id="135C729AFFFFF34BD28690DCD83BFBF5" box="[679,841,1007,1031]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="11" pageNumber="188">Figs 13, 15–16</figureCitation>
) and a female (
<figureCitation id="135C729AFFFFF34BD3CC90DCDF4EFBF5" box="[1005,1084,1007,1031]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="11" pageNumber="188">Fig. 14</figureCitation>
, without elytra). These specimens doubtlessly belong to the same taxon that I discussed as
<taxonomicName id="4C67159CFFFFF34BD0509703DB97FBB5" authorityName="Gildenkov" authorityYear="2000" box="[113,229,1072,1095]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD0509703DB97FBB5" box="[113,229,1072,1095]" italics="true" pageId="11" pageNumber="188">T. nigerius</emphasis>
</taxonomicName>
. The study of the previously unavailable female ringstructure (
<figureCitation id="135C729AFFFFF34BD3B2971CD896FBB5" box="[915,996,1071,1095]" captionStart="Figs 81–85. 81–83" captionStartId="25.[113,150,1532,1551]" captionTargetBox="[128,1073,222,1462]" captionTargetId="figure-10@25.[124,1076,219,1466]" captionTargetPageId="25" captionText="Figs 81–85. 81–83 – female ringstructures.81 – Thinodromus tibialis (Fauvel, 1907); 82 – T. dasys Gildenkov, 2000; 83 – T. nigerius Gildenkov, 2000. 84–85 – female ringstructures with spermathecae. 84 – T. facilis Gildenkov, 2000; 85 – T. gildenkovi sp. nov. Scale bar: 0.10 mm for 85, 0.12 mm for 81 and 83–84, 0.14 mm for 82." figureDoi="http://doi.org/10.5281/zenodo.5739681" httpUri="https://zenodo.org/record/5739681/files/figure.png" pageId="11" pageNumber="188">Fig. 83</figureCitation>
), i.e. the only diagnostic female genitalic feature in this group of species, revealed that it is closely similar to that of
<taxonomicName id="4C67159CFFFFF34BD1139743D9C6FB75" authority="Makranczy, 2009" authorityName="Makranczy" authorityYear="2009" box="[306,692,1135,1159]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="kedougouensis">
<emphasis id="B913B20DFFFFF34BD1139743DA9CFB75" box="[306,494,1136,1159]" italics="true" pageId="11" pageNumber="188">T. kedougouensis</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD1D6975CD9C6FB75" author="MAKRANCZY GY" box="[503,692,1135,1159]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">Makranczy, 2009</bibRefCitation>
</taxonomicName>
described from
<collectingCountry id="F3702E8FFFFFF34BD34B975CD8B6FB75" box="[874,964,1135,1159]" name="Senegal" pageId="11" pageNumber="188">Senegal</collectingCountry>
. Therefore, it seems that we are dealing with a single externally variable species and the known specimens are just parts of its morphological continuum. Specimens previously mentioned by me as
<taxonomicName id="4C67159CFFFFF34BD09697E3DA59FB15" authorityName="Gildenkov" authorityYear="2000" box="[183,299,1232,1255]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD09697E3DA59FB15" box="[183,299,1232,1255]" italics="true" pageId="11" pageNumber="188">T. nigerius</emphasis>
</taxonomicName>
(see
<figureCitation id="135C729AFFFFF34BD14497FCD97BFB15" box="[357,521,1231,1255]" captionStart="Figs 19–27. 19" captionStartId="15.[113,150,1493,1512]" captionTargetBox="[120,1082,219,1480]" captionTargetId="figure-10@15.[119,1082,219,1481]" captionTargetPageId="15" captionText="Figs 19–27. 19 – Thinodromus facilis Gildenkov, 2000, female protarsus; 20 – T. gildenkovi sp. nov., male protarsus; 21 – T. gildenkovi sp. nov., male, fringe of tergite VII. Antennae. 22 – T. sudanensis (Scheerpeltz, 1974), female; 23 – T. meridionalis sp. nov., female; 24–25 – T. gildenkovi sp. nov. (24 – female; 25 – male); 26–27 – T. dasys Gildenkov, 2000 (26 – female; 27 – male) Scale bar: 0.065 mm for 19, 0.08 mm for 20, 0.12 mm for 21, 0.24 mm for 24–25, 0.3 mm for 22 and 26–27, 0.33 mm for 23." figureDoi="http://doi.org/10.5281/zenodo.5739655" httpUri="https://zenodo.org/record/5739655/files/figure.png" pageId="11" pageNumber="188">Figs 19, 20, 21</figureCitation>
,
<figureCitation id="135C729AFFFFF34BD23497FCD929FB15" box="[533,603,1231,1255]" captionStart="Figs 28–33. 28–31" captionStartId="16.[151,188,1545,1564]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@16.[151,1126,219,1497]" captionTargetPageId="16" captionText="Figs 28–33. 28–31 – Thinodromus tibialis (Fauvel, 1907) (28 – head and pronotum; 29 – female antenna; 30 – male antenna; 31 – elytron and abdominal base); 32–33 – T. gildenkovi sp. nov. (32 – head and pronotum; 33 – elytron and abdominal base). Scale bar: 0.45 mm for 28–31, 0.55 mm for 32–33." figureDoi="http://doi.org/10.5281/zenodo.5739659" httpUri="https://zenodo.org/record/5739659/files/figure.png" pageId="11" pageNumber="188">31, 32</figureCitation>
,
<figureCitation id="135C729AFFFFF34BD24697FCD9F0FB15" box="[615,642,1231,1255]" captionStart="Figs 64–69. 64–66" captionStartId="22.[151,188,1519,1538]" captionTargetBox="[154,1101,221,1465]" captionTargetId="figure-9@22.[151,1103,219,1466]" captionTargetPageId="22" captionText="Figs 64–69. 64–66 – Thinodromus meridionalis sp. nov., male (64–65 – aedeagus; 66 – paramere); 67–69 – T. facilis Gildenkov, 2000, male (67–68 – aedeagus; 69 – paramere). Abbreviations: AC – apical copulatory sclerite; BA – basal sclerites, BM – basomedial sclerites, MA – medioapical sclerites ML – medial lamellae. Scale bar: 0.10 mm for 64–66, 0.12 mm for 67–69." figureDoi="http://doi.org/10.5281/zenodo.5739675" httpUri="https://zenodo.org/record/5739675/files/figure.png" pageId="11" pageNumber="188">68</figureCitation>
in
<bibRefCitation id="EFF613EEFFFFF34BD28497E3D805FB15" author="MAKRANCZY GY" box="[677,887,1231,1255]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">MAKRANCZY (2009)</bibRefCitation>
for terminalia and genitalia) and
<taxonomicName id="4C67159CFFFFF34BD13197C3DABFFAF5" authorityName="Makranczy" authorityYear="2009" box="[272,461,1264,1287]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="kedougouensis">
<emphasis id="B913B20DFFFFF34BD13197C3DABFFAF5" box="[272,461,1264,1287]" italics="true" pageId="11" pageNumber="188">T. kedougouensis</emphasis>
</taxonomicName>
(
<figureCitation id="135C729AFFFFF34BD1FC97DCD9FBFAF5" box="[477,649,1263,1287]" captionStart="Figs 46–51. 46–47" captionStartId="19.[113,150,1545,1564]" captionTargetBox="[115,1063,223,1457]" captionTargetId="figure-9@19.[113,1065,219,1466]" captionTargetPageId="19" captionText="Figs 46–51. 46–47 – Thinodromus dasys Gildenkov, 2000, male (46 – sternite VIII; 47 – tergite X); 48–49 – T. gildenkovi sp. nov., male (48 – sternite VIII; 49 – tergite X); 50–51 – T. rhodesianus (Scheerpeltz, 1974), male (50 – sternite VIII; 51 – tergite X). Scale bar: 0.10 mm for 49 and 51, 0.13 mm for 47–48 and 50, 0.15 mm for 46." figureDoi="http://doi.org/10.5281/zenodo.5739667" httpUri="https://zenodo.org/record/5739667/files/figure.png" pageId="11" pageNumber="188">Figs 47, 48, 49</figureCitation>
,
<figureCitation id="135C729AFFFFF34BD2B497DCD9A8FAF5" box="[661,730,1263,1287]" captionStart="Figs 52–57. 52–54" captionStartId="20.[151,188,1520,1539]" captionTargetBox="[154,1102,220,1466]" captionTargetId="figure-10@20.[151,1103,220,1467]" captionTargetPageId="20" captionText="Figs 52–57. 52–54 – Thinodromus rhodesianus (Scheerpeltz, 1974), male (52–53 – aedeagus; 54 – paramere); 55–57 – T. tibialis (Fauvel, 1907), male (55–56 – aedeagus; 57 – paramere).Abbreviations:AC – apical copulatory sclerite; BA – basal sclerites, BM – basomedial sclerites, MA – medioapical sclerites ML – medial lamellae. Scale bar: 0.10 mm for 55–57, 0.12 mm for 52–54." figureDoi="http://doi.org/10.5281/zenodo.5739671" httpUri="https://zenodo.org/record/5739671/files/figure.png" pageId="11" pageNumber="188">
55,
<quantity id="4C9FC3FAFFFFF34BD29E97DCD9A8FAF5" box="[703,730,1263,1287]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="11" pageNumber="188" unit="in" value="56.0">56</quantity>
</figureCitation>
in
<bibRefCitation id="EFF613EEFFFFF34BD32197C3D8A4FAF5" author="MAKRANCZY GY" box="[768,982,1263,1287]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">MAKRANCZY (2009)</bibRefCitation>
, and
<figureCitation id="135C729AFFFFF34BD43397DCDBC7FAD5" captionStart="Figs 70–76. 70–74" captionStartId="23.[113,150,1492,1511]" captionTargetBox="[115,1063,221,1431]" captionTargetId="figure-10@23.[113,1065,219,1457]" captionTargetPageId="23" captionText="Figs 70–76. 70–74 – Thinodromus capensis (Bernhauer, 1934), male (70 – sternite VIII; 71 – tergite X; 72–73 – aedeagus; 74 – paramere); 75–76 – T. nigerius Gildenkov, 2000, male (75 – sclerotized subapical plate of aedeagus; 76 – apical copulatory sclerite).Abbreviations:AC – apical copulatory sclerite; BA – basal sclerites, BM – basomedial sclerites, MA – medioapical sclerites ML – medial lamellae. Scale bar: 0.05 mm for 76, 0.08 mm for 75, 0.11 mm for 72–74, 0.12 mm for 71, 0.14 mm for 70." figureDoi="http://doi.org/10.5281/zenodo.5739677" httpUri="https://zenodo.org/record/5739677/files/figure.png" pageId="11" pageNumber="188">Figs 75–76</figureCitation>
and
<figureCitation id="135C729AFFFFF34BD0CF963CDA7BFAD5" box="[238,265,1295,1319]" captionStart="Figs 81–85. 81–83" captionStartId="25.[113,150,1532,1551]" captionTargetBox="[128,1073,222,1462]" captionTargetId="figure-10@25.[124,1076,219,1466]" captionTargetPageId="25" captionText="Figs 81–85. 81–83 – female ringstructures.81 – Thinodromus tibialis (Fauvel, 1907); 82 – T. dasys Gildenkov, 2000; 83 – T. nigerius Gildenkov, 2000. 84–85 – female ringstructures with spermathecae. 84 – T. facilis Gildenkov, 2000; 85 – T. gildenkovi sp. nov. Scale bar: 0.10 mm for 85, 0.12 mm for 81 and 83–84, 0.14 mm for 82." figureDoi="http://doi.org/10.5281/zenodo.5739681" httpUri="https://zenodo.org/record/5739681/files/figure.png" pageId="11" pageNumber="188">83</figureCitation>
here) seem therefore to be conspecific. This is also supported by the examined specimens from
<collectingCountry id="F3702E8FFFFFF34BD14F961CDADDFAB5" box="[366,431,1327,1351]" name="Benin" pageId="11" pageNumber="188">Bénin</collectingCountry>
(all from the same collecting event and likely from the same microhabitat) which contain two conspecific males (based on morphology of the genitalia), with entirely different antennae: one has shorter antenna much similar to that illustrated in the original description of
<taxonomicName id="4C67159CFFFFF34BD1B796A3D979FA55" authorityName="Gildenkov" authorityYear="2000" box="[406,523,1424,1447]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD1B796A3D979FA55" box="[406,523,1424,1447]" italics="true" pageId="11" pageNumber="188">T. nigerius</emphasis>
</taxonomicName>
, the other has much more elongate antenna, as that in
<figureCitation id="135C729AFFFFF34BD0AF969CDBA9FA35" box="[142,219,1455,1479]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="11" pageNumber="188">
Fig.
<quantity id="4C9FC3FAFFFFF34BD0E1969CDBA9FA35" box="[192,219,1455,1479]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="11" pageNumber="188" unit="in" value="15.0">15</quantity>
</figureCitation>
in this study.
</paragraph>
<paragraph id="8BD86E1FFFFFF34BD0B196E3D9FDF9D5" blockId="11.[113,1089,634,1576]" pageId="11" pageNumber="188">
For the reasons given above,
<taxonomicName id="4C67159CFFFFF34BD1F696E3D825FA15" authority="Makranczy, 2009" authorityName="Makranczy" authorityYear="2009" box="[471,855,1487,1511]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="kedougouensis">
<emphasis id="B913B20DFFFFF34BD1F696E3D9E3FA15" box="[471,657,1488,1511]" italics="true" pageId="11" pageNumber="188">T. kedougouensis</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD2BA96FCD825FA15" author="MAKRANCZY GY" box="[667,855,1487,1511]" pageId="11" pageNumber="188" pagination="33 - 61" refId="ref14404" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">Makranczy, 2009</bibRefCitation>
</taxonomicName>
is here conditionally synonymized with
<taxonomicName id="4C67159CFFFFF34BD16096C3D907F9F5" authority="Gildenkov, 2000" authorityName="Gildenkov" authorityYear="2000" box="[321,629,1519,1543]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD16096C3DAC6F9F5" box="[321,436,1520,1543]" italics="true" pageId="11" pageNumber="188">T. nigerius</emphasis>
<bibRefCitation id="EFF613EEFFFFF34BD19F96DCD907F9F5" author="GILDENKOV M. &amp; YU" box="[446,629,1519,1543]" pageId="11" pageNumber="188" pagination="43 - 56" refId="ref14247" refString="GILDENKOV M. YU. 2000: Novyy podrod i sem' novykh dlya nauki vidov roda Thinodromus iz efiopskoy zoogeograficheskoy oblast'i (Coleoptera, Staphylinidae). [New subgenus and seven species, new to science, of the genus Thinodromus from Ethiopian zoogeographic region (Coleoptera, Staphylinidae)]. Izvestiya Khar ' kovskogo Entomologicheskogo Obshchestva [Kharkov Entomological Society Gazette] 8 (1): 43 - 56 (in Russian)." type="journal article" year="2000">Gildenkov, 2000</bibRefCitation>
</taxonomicName>
. The synonymy needs to be confirmed by more material from the
<typeStatus id="54DCD0BDFFFFF34BD14E9523DAEFF9DA" box="[367,413,1552,1576]" pageId="11" pageNumber="188">type</typeStatus>
locality of
<taxonomicName id="4C67159CFFFFF34BD2369523D9FAF9D5" authorityName="Gildenkov" authorityYear="2000" box="[535,648,1552,1575]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="nigerius">
<emphasis id="B913B20DFFFFF34BD2369523D9FAF9D5" box="[535,648,1552,1575]" italics="true" pageId="11" pageNumber="188">T. nigerius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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