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<document id="EE3FDD4A97CFBED816BD25F1D2F86610" ID-DOI="10.15468/dl.qysyqm" ID-ISSN="0132-8069" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713266371983" checkinUser="felipe" docAuthor="User, GBIF. Org" docDate="2023" docId="03C987AB2238EA7DFC85FED2FC536C24" docLanguage="en" docName="RussEntomolJ.32.1.53-57.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15468/dl.qysyqm" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Exomala orientalis" docType="treatment" docVersion="1" lastPageNumber="55" masterDocId="FFF0FFD32239EA7FFFFEFFA3FF856437" masterDocTitle="Occurrence Download" masterLastPageNumber="57" masterPageNumber="53" pageNumber="54" updateTime="1713266372729" updateUser="felipe">
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<subSubSection id="C37A65362238EA7EFC85FED2FB506591" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="8BDF36BD2238EA7EFC85FED2FAA265BE" blockId="1.[812,1398,369,531]" box="[891,1319,369,393]" pageId="1" pageNumber="54">
<heading id="D09781D12238EA7EFC85FED2FAA265BE" box="[891,1319,369,393]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="54" reason="8">
<taxonomicName id="4C604D3E2238EA7EFC85FED2FAA265BE" ID-CoL="7W6NX" authority="(Waterhouse, 1875)" baseAuthorityName="Waterhouse" baseAuthorityYear="1875" box="[891,1319,369,393]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B914EAAF2238EA7EFC85FED2FBCC65BE" box="[891,1097,369,393]" italics="true" pageId="1" pageNumber="54">Exomala orientalis</emphasis>
(
<bibRefCitation id="EFF14B4C2238EA7EFBA7FED2FAA565BE" author="Waterhouse C. O." box="[1113,1312,369,393]" pageId="1" pageNumber="54" pagination="71 - 116" refId="ref2692" refString="Waterhouse C. O. 1875. On the Lamellicorn Coleoptera of Japan // The Transactions of the Entomological Society of London, P. 71 - 116. Pl. III." type="book chapter" year="1875">Waterhouse, 1875</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDF36BD2238EA7EFC30FE2DFB506591" blockId="1.[812,1398,369,531]" box="[974,1237,398,422]" pageId="1" pageNumber="54">Figs 17, 1113, 1617.</paragraph>
</subSubSection>
<subSubSection id="C37A65362238EA7DFCACFE10FC536C24" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="8BDF36BD2238EA7EFCACFE10FBE96624" blockId="1.[812,1398,369,531]" pageId="1" pageNumber="54">
<materialsCitation id="3B083CE02238EA7EFCACFE10FBED6624" collectionCode="NHMUK" country="Japan" location="Japan" pageId="1" pageNumber="54" specimenCode="NHMUK014400115" specimenCount="1" typeStatus="lectotype">
MATERIAL. “
<typeStatus id="54DB881F2238EA7EFC0BFE14FBDD65FD" box="[1013,1112,439,458]" pageId="1" pageNumber="54" type="lectotype">Lectotypus</typeStatus>
,
<taxonomicName id="4C604D3E2238EA7EFB9AFE14FC2665D4" authority="(Waterhouse, 1875)" baseAuthorityName="Waterhouse" baseAuthorityYear="1875" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="orientalis">
Phyllopertha orientalis (
<bibRefCitation id="EFF14B4C2238EA7EFAC7FE14FC1865D4" author="Waterhouse C. O." pageId="1" pageNumber="54" pagination="71 - 116" refId="ref2692" refString="Waterhouse C. O. 1875. On the Lamellicorn Coleoptera of Japan // The Transactions of the Entomological Society of London, P. 71 - 116. Pl. III." type="book chapter" year="1875">Waterhouse, 1875</bibRefCitation>
)
</taxonomicName>
, des. Kasatkin D., 2022”, “
<taxonomicName id="4C604D3E2238EA7EFB55FE73FC0265CB" authority="O. C. Wat." authorityName="O. C. Wat." class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="orientalis">Phyllopertha orientalis O.C. Wat.</taxonomicName>
, type, var. B.”, “type (circle label with red ring)”, “
<collectingCountry id="F377762D2238EA7EFABAFE4BFAF365CC" box="[1348,1398,488,507]" name="Japan" pageId="1" pageNumber="54">Japan</collectingCountry>
(f.v. 112 4)”,
<specimenCode id="DBC69EC62238EA7EFC57FDA3FBDA6624" box="[937,1119,512,531]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="54" type="Museum">NHMUK014400115</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36BD223BEA7DFF2DFB14FB9E60E6" blockId="2.[211,627,1206,1233]" box="[211,1051,1206,1233]" lastBlockId="2.[1036,1051,1207,1233]" pageId="2" pageNumber="55">
<emphasis id="B914EAAF223BEA7DFF2DFB14FB9E60E6" bold="true" box="[211,1051,1206,1233]" pageId="2" pageNumber="55">5 6 7</emphasis>
</paragraph>
<caption id="DF1F6635223BEA7DFF1CFB53FD936178" pageId="2" pageNumber="55" startId="2.[226,263,1264,1283]" targetBox="[190,1396,315,1196]" targetPageId="2" targetType="figure">
<paragraph id="8BDF36BD223BEA7DFF1CFB53FEFD612C" blockId="2.[189,1400,1264,1359]" pageId="2" pageNumber="55">
Figs 57. Labels of type specimens of
<taxonomicName id="4C604D3E223BEA7DFDCDFB53FD796134" box="[563,764,1264,1283]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B914EAAF223BEA7DFDCDFB53FD796134" box="[563,764,1264,1283]" italics="true" pageId="2" pageNumber="55">Phyllopertha orientalis</emphasis>
</taxonomicName>
: 56 — paralectotypes, 7 — lectotype; a — reverse side of circle labels. Photos by D. Telnov.
</paragraph>
<paragraph id="8BDF36BD223BEA7DFF1CFA87FD936178" blockId="2.[189,1400,1264,1359]" pageId="2" pageNumber="55">
Рис. 57. Этикетки типовых ЭкЗемплЯров
<taxonomicName id="4C604D3E223BEA7DFD97FA87FCB66100" box="[617,819,1316,1335]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B914EAAF223BEA7DFD97FA87FCB66100" box="[617,819,1316,1335]" italics="true" pageId="2" pageNumber="55">Phyllopertha orientalis</emphasis>
</taxonomicName>
: 56 — паралектотипы, 7 — лектотип; a — обратнаЯ сторона круглой Этикетки. Фото Д. Тельнова.
</paragraph>
</caption>
<paragraph id="8BDF36BD223BEA7DFF1CFA23FE0661EC" blockId="2.[189,776,1407,2063]" pageId="2" pageNumber="55">
<materialsCitation id="3B083CE0223BEA7DFF1CFA23FEFA61EC" collectionCode="NHMUK" collectorName="Kasatkin D." country="Japan" location="Japan" pageId="2" pageNumber="55" specimenCode="NHMUK014400117" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="54DB881F223BEA7DFF1CFA23FEE161A4" box="[226,356,1408,1427]" pageId="2" pageNumber="55" type="paralectotype">Paralectotypus</typeStatus>
,
<taxonomicName id="4C604D3E223BEA7DFE95FA23FD5861A5" authority="(Waterhouse, 1875)" baseAuthorityName="Waterhouse" baseAuthorityYear="1875" box="[363,733,1407,1427]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
Phyllopertha orientalis (
<bibRefCitation id="EFF14B4C223BEA7DFDC9FA23FD5261A5" author="Waterhouse C. O." box="[567,727,1407,1427]" pageId="2" pageNumber="55" pagination="71 - 116" refId="ref2692" refString="Waterhouse C. O. 1875. On the Lamellicorn Coleoptera of Japan // The Transactions of the Entomological Society of London, P. 71 - 116. Pl. III." type="book chapter" year="1875">Waterhouse, 1875</bibRefCitation>
)
</taxonomicName>
, des.
<collectorName id="2695536B223BEA7DFF43FA3BFEAD619C" box="[189,296,1432,1451]" pageId="2" pageNumber="55">Kasatkin D.</collectorName>
, 2022”, “
<taxonomicName id="4C604D3E223BEA7DFE83FA3BFD2F619C" authority="O. C. Wat." authorityName="O. C. Wat." box="[381,682,1432,1451]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">Phyllopertha orientalis O.C. Wat.</taxonomicName>
, type, var. A.”, “type (circle label with red ring)”, “
<collectingCountry id="F377762D223BEA7DFDBDFA13FDF061F4" box="[579,629,1456,1475]" name="Japan" pageId="2" pageNumber="55">Japan</collectingCountry>
(f.v. 61 128)”,
<specimenCode id="DBC69EC6223BEA7DFF43FA6BFEF361EC" box="[189,374,1480,1499]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="55" type="Museum">NHMUK014400117</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36BD223BEA7DFF1CFA43FE06620C" blockId="2.[189,776,1407,2063]" pageId="2" pageNumber="55">
<materialsCitation id="3B083CE0223BEA7DFF1CFA43FEFA620C" collectionCode="NHMUK" collectorName="Kasatkin D." country="Japan" location="Japan" pageId="2" pageNumber="55" specimenCode="NHMUK014400116" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="54DB881F223BEA7DFF1CFA43FEE161C4" box="[226,356,1504,1523]" pageId="2" pageNumber="55" type="paralectotype">Paralectotypus</typeStatus>
,
<taxonomicName id="4C604D3E223BEA7DFE95FA43FD5861C5" authority="(Waterhouse, 1875)" baseAuthorityName="Waterhouse" baseAuthorityYear="1875" box="[363,733,1503,1523]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
Phyllopertha orientalis (
<bibRefCitation id="EFF14B4C223BEA7DFDC9FA43FD5261C5" author="Waterhouse C. O." box="[567,727,1503,1523]" pageId="2" pageNumber="55" pagination="71 - 116" refId="ref2692" refString="Waterhouse C. O. 1875. On the Lamellicorn Coleoptera of Japan // The Transactions of the Entomological Society of London, P. 71 - 116. Pl. III." type="book chapter" year="1875">Waterhouse, 1875</bibRefCitation>
)
</taxonomicName>
, des.
<collectorName id="2695536B223BEA7DFF43FA5BFEAD623C" box="[189,296,1528,1547]" pageId="2" pageNumber="55">Kasatkin D.</collectorName>
, 2022”, “
<taxonomicName id="4C604D3E223BEA7DFE83FA5BFD2F623C" authority="O. C. Wat." authorityName="O. C. Wat." box="[381,682,1528,1547]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">Phyllopertha orientalis O.C. Wat.</taxonomicName>
, type, var. C.”, “type (circle label with red ring)”, “
<collectingCountry id="F377762D223BEA7DFDB4F9B3FDF86214" box="[586,637,1552,1571]" name="Japan" pageId="2" pageNumber="55">Japan</collectingCountry>
(f.v. 66 15)”,
<specimenCode id="DBC69EC6223BEA7DFF43F98BFEF3620C" box="[189,374,1576,1595]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="55" type="Museum">NHMUK014400116</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36BD223BEA7DFF1CF99CFE5562DD" blockId="2.[189,776,1407,2063]" pageId="2" pageNumber="55">
<materialsCitation id="3B083CE0223BEA7DFF1CF99CFE4962DD" country="United Kingdom" county="London" location="All" municipality="Natural History Museum" pageId="2" pageNumber="55" specimenCount="2" typeStatus="lectotype">
The
<typeStatus id="54DB881F223BEA7DFEE8F99CFEFD6260" box="[278,376,1599,1623]" pageId="2" pageNumber="55" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C604D3E223BEA7DFE5DF99CFD1B6260" box="[419,670,1599,1623]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B914EAAF223BEA7DFE5DF99CFD1B6260" box="[419,670,1599,1623]" italics="true" pageId="2" pageNumber="55">Phyllopertha orientalis</emphasis>
</taxonomicName>
is designated here. We designated as a
<typeStatus id="54DB881F223BEA7DFDEEF9FEFDF76242" box="[528,626,1629,1653]" pageId="2" pageNumber="55" type="lectotype">lectotype</typeStatus>
the specimen marked as var. B as the most consistent with the typical coloration of this species.
<location id="8EBF6066223BEA7DFE2CF934FE716298" LSID="urn:lsid:plazi:treatment:03C987AB2238EA7DFC85FED2FC536C24:8EBF6066223BEA7DFE2CF934FE716298" box="[466,500,1687,1711]" country="United Kingdom" county="London" municipality="Natural History Museum" name="All" pageId="2" pageNumber="55">All</location>
types are deposited in the collection of
<collectingMunicipality id="6BBBACC7223BEA7DFEABF916FDEA62FB" box="[341,623,1717,1741]" pageId="2" pageNumber="55">Natural History Museum</collectingMunicipality>
,
<collectingCounty id="62BE4E31223BEA7DFD81F916FD5062FA" box="[639,725,1717,1741]" pageId="2" pageNumber="55">London</collectingCounty>
, the
<collectingCountry id="F377762D223BEA7DFF43F971FEF462DD" box="[189,369,1746,1770]" name="United Kingdom" pageId="2" pageNumber="55">United Kingdom</collectingCountry>
(NHM)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36BD223BEA7DFF1CF953FC6863BD" blockId="2.[189,776,1407,2063]" lastBlockId="2.[812,1399,1408,1930]" pageId="2" pageNumber="55">
In the Palearctic catalogue, the nominative subgenus
<taxonomicName id="4C604D3E223BEA7DFF10F8AEFED56312" authorityName="Reitter" authorityYear="1903" box="[238,336,1805,1829]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAAF223BEA7DFF10F8AEFED56312" box="[238,336,1805,1829]" italics="true" pageId="2" pageNumber="55">Exomala</emphasis>
</taxonomicName>
includes also
<taxonomicName id="4C604D3E223BEA7DFE05F8ADFF7A6375" authority="(Sawada, 1941)" baseAuthorityName="Sawada" baseAuthorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="ohdaiensis">
<emphasis id="B914EAAF223BEA7DFE05F8ADFD106312" box="[507,661,1805,1829]" italics="true" pageId="2" pageNumber="55">E. ohdaiensis</emphasis>
(Sawada, 1941)
</taxonomicName>
,
<taxonomicName id="4C604D3E223BEA7DFEEEF888FDFC6375" authority="(Reitter, 1903)" baseAuthorityName="Reitter" baseAuthorityYear="1903" box="[272,633,1834,1858]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="pallidipennis">
<emphasis id="B914EAAF223BEA7DFEEEF888FE406375" box="[272,453,1834,1858]" italics="true" pageId="2" pageNumber="55">E. pallidipennis</emphasis>
(
<bibRefCitation id="EFF14B4C223BEA7DFE26F889FDF76375" author="Reitter E." box="[472,626,1834,1858]" pageId="2" pageNumber="55" pagination="28 - 158" refId="ref2594" refString="Reitter E. 1903. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern enthaltend die Gruppen der Rutelini, Hopliini und Glaphyrini. (Schluss) // Verhandlungen des Naturforschenden Vereins in Brunn. T. 46 [1902]. S. 28 - 158." type="journal article" year="1903">Reitter, 1903</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C604D3E223BEA7DFD46F888FE4C6368" authority="(Ohaus, 1925)" baseAuthorityName="Ohaus" baseAuthorityYear="1925" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="okiwaensis">
<emphasis id="B914EAAF223BEA7DFD46F888FE906368" italics="true" pageId="2" pageNumber="55">E. okiwaensis</emphasis>
(
<bibRefCitation id="EFF14B4C223BEA7DFED5F8E4FE476368" author="Ohaus F. I." box="[299,450,1863,1887]" pageId="2" pageNumber="55" pagination="122 - 131" refId="ref2462" refString="Ohaus F. I. 1925. Nachtrag zur Rutelinenfauna Formosas (Col. Lamell.) // Archiv fur Naturgeschichte. Bd. 91 (A). H. 5. S. 122 - 131." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
)
</taxonomicName>
[
<bibRefCitation id="EFF14B4C223BEA7DFE20F8EBFD3D6368" author="Zorn C. &amp; Bezdek A." box="[478,696,1863,1887]" pageId="2" pageNumber="55" pagination="317 - 358" refId="ref2850" refString="Zorn C., Bezdek A. 2016. Rutelinae // In: Lobl I., Lobl D. (eds.). Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Leiden, Boston: Brill. P. 317 - 358." type="book chapter" year="2016">
Zorn
<emphasis id="B914EAAF223BEA7DFDDBF8EBFDD86368" box="[549,605,1863,1887]" italics="true" pageId="2" pageNumber="55">et al</emphasis>
., 2016
</bibRefCitation>
]. Both editions of the Palaearctic Catalogue indicate that
<taxonomicName id="4C604D3E223BEA7DFD11F8C5FEB263AD" baseAuthorityName="Ohaus" baseAuthorityYear="1925" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="okiwaensis">
<emphasis id="B914EAAF223BEA7DFD11F8C5FEB263AD" italics="true" pageId="2" pageNumber="55">E. okiwaensis</emphasis>
</taxonomicName>
is endemic to Okinawa and has never been found outside the island, but according to the original description of this species [
<bibRefCitation id="EFF14B4C223BEA7DFDB0F81EFD6663E2" author="Ohaus F. I." box="[590,739,1981,2005]" pageId="2" pageNumber="55" pagination="122 - 131" refId="ref2462" refString="Ohaus F. I. 1925. Nachtrag zur Rutelinenfauna Formosas (Col. Lamell.) // Archiv fur Naturgeschichte. Bd. 91 (A). H. 5. S. 122 - 131." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
], it comes from
<collectingCountry id="F377762D223BEA7DFEB4F878FE2E63C5" box="[330,427,2011,2034]" name="Taiwan" pageId="2" pageNumber="55">Formosa</collectingCountry>
(now Taiwan). In the publication about the subgeneric structure of
<taxonomicName id="4C604D3E223BEA7DFD8DF854FD506C38" authorityName="Reitter" authorityYear="1903" box="[627,725,2039,2063]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAAF223BEA7DFD8DF854FD506C38" box="[627,725,2039,2063]" italics="true" pageId="2" pageNumber="55">Exomala</emphasis>
</taxonomicName>
and related genera [
<bibRefCitation id="EFF14B4C223BEA7DFC27FA23FBFD61AF" author="Baraud J." box="[985,1144,1408,1432]" pageId="2" pageNumber="55" pagination="46 - 62" refId="ref1843" refString="Baraud J. 1991. Nouvelle classification proposee pour les especes du genre Blithopertha Reitter, 1903 (Coleoptera, Rutelidae) // Lambillionea. Vol. 91. P. 46 - 62." type="journal article" year="1991">Baraud, 1991</bibRefCitation>
],
<taxonomicName id="4C604D3E223BEA7DFB72FA23FAA361AF" baseAuthorityName="Ohaus" baseAuthorityYear="1925" box="[1164,1318,1408,1432]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="okiwaensis">
<emphasis id="B914EAAF223BEA7DFB72FA23FAA361AF" box="[1164,1318,1408,1432]" italics="true" pageId="2" pageNumber="55">E. okiwaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D3E223BEA7DFAA1FA23FC266182" baseAuthorityName="Sawada" baseAuthorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="ohdaiensis">
<emphasis id="B914EAAF223BEA7DFAA1FA23FC266182" italics="true" pageId="2" pageNumber="55">E. ohdaiensis</emphasis>
</taxonomicName>
were not mentioned. Probably, the author did not have material available for studying the male genitalia (Barauds system is based mainly on the structure of the aedeagus). The study the male genitalia is necessary to clearly understand whether these species really belong to the nominative subgenus
<taxonomicName id="4C604D3E223BEA7DFCD2F9EEFC0B6252" authorityName="Reitter" authorityYear="1903" box="[812,910,1613,1637]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAAF223BEA7DFCD2F9EEFC0B6252" box="[812,910,1613,1637]" italics="true" pageId="2" pageNumber="55">Exomala</emphasis>
</taxonomicName>
(and genus
<taxonomicName id="4C604D3E223BEA7DFBE9F9EEFBFC6252" authorityName="Reitter" authorityYear="1903" box="[1047,1145,1613,1637]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAAF223BEA7DFBE9F9EEFBFC6252" box="[1047,1145,1613,1637]" italics="true" pageId="2" pageNumber="55">Exomala</emphasis>
</taxonomicName>
in generally). The imperfection of the genus system is supported by the fact that
<taxonomicName id="4C604D3E223BEA7DFC71F92BFAC16297" authority="(Sawada, 1941)" baseAuthorityName="Sawada" baseAuthorityYear="1941" box="[911,1348,1672,1696]" class="Insecta" family="Scarabaeidae" genus="Blithopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="tarowana">
<emphasis id="B914EAAF223BEA7DFC71F92BFB0B62A8" box="[911,1166,1672,1696]" italics="true" pageId="2" pageNumber="55">Blithopertha tarowana</emphasis>
(Sawada, 1941)
</taxonomicName>
was assigned to the genus
<taxonomicName id="4C604D3E223BEA7DFBDDF906FB00628A" authorityName="Reitter" authorityYear="1903" box="[1059,1157,1701,1725]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAAF223BEA7DFBDDF906FB00628A" box="[1059,1157,1701,1725]" italics="true" pageId="2" pageNumber="55">Exomala</emphasis>
</taxonomicName>
in the first edition of the Palaearctic catalogue [
<bibRefCitation id="EFF14B4C223BEA7DFBADF960FB5062ED" author="Zorn C." box="[1107,1237,1730,1754]" pageId="2" pageNumber="55" refId="ref2802" refString="Zorn C. 2006. Subfamily Rutelinae // In: Lobl I., Smetana A. (eds). Catalogue of Palaearctic Coleoptera, Vol. 3. Scarabaeoidea - Scirtoidea - Dasciloidea - Buprestoidea - Byrrhoidea. Apollo Books: Stenstrup. 690 pp." type="book" year="2006">Zorn, 2006</bibRefCitation>
] and then was included in
<taxonomicName id="4C604D3E223BEA7DFC55F943FB8B62CF" authorityName="Samouelle" authorityYear="1819" box="[939,1038,1760,1784]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAAF223BEA7DFC55F943FB8B62CF" box="[939,1038,1760,1784]" italics="true" pageId="2" pageNumber="55">Anomala</emphasis>
</taxonomicName>
in the second one [
<bibRefCitation id="EFF14B4C223BEA7DFB26F943FCEC6322" author="Zorn C. &amp; Bezdek A." pageId="2" pageNumber="55" pagination="317 - 358" refId="ref2850" refString="Zorn C., Bezdek A. 2016. Rutelinae // In: Lobl I., Lobl D. (eds.). Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Leiden, Boston: Brill. P. 317 - 358." type="book chapter" year="2016">Zorn, Bezdek, 2016</bibRefCitation>
]. The endophallus of
<taxonomicName id="4C604D3E223BEA7DFB9FF95DFB6B6322" box="[1121,1262,1789,1813]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B914EAAF223BEA7DFB9FF95DFB6B6322" box="[1121,1262,1789,1813]" italics="true" pageId="2" pageNumber="55">E. orientalis</emphasis>
</taxonomicName>
was briefly described and illustrated by
<bibRefCitation id="EFF14B4C223BEA7DFB7BF8B9FABB6305" author="Shokhin I." box="[1157,1342,1818,1842]" pageId="2" pageNumber="55" pagination="107 - 111" refId="ref2644" refString="Shokhin I. 2017. [Morphology of endophalli of the genera Exomala Reitter, 1903 and Micropertha Baraud, 1991 (Coleoptera, Scarabaeidae, Rutelinae)] // Nauka Yugha Rossii. Vol. 13. P. 107 - 111 [in Russian]." type="book chapter" year="2017">Shokhin [2017]</bibRefCitation>
. We publish here comparative images of the endophallus and the aedeagus of
<taxonomicName id="4C604D3E223BEA7DFBF4F8F5FB3F635A" baseAuthorityName="Reitter" baseAuthorityYear="1903" box="[1034,1210,1877,1901]" class="Insecta" family="Scarabaeidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="pallidipennis">
<emphasis id="B914EAAF223BEA7DFBF4F8F5FB3F635A" box="[1034,1210,1877,1901]" italics="true" pageId="2" pageNumber="55">E. pallidipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D3E223BEA7DFB13F8F5FAF3635A" box="[1261,1398,1877,1901]" class="Insecta" family="Scarabaeidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B914EAAF223BEA7DFB13F8F5FAF3635A" box="[1261,1398,1877,1901]" italics="true" pageId="2" pageNumber="55">E. orientalis</emphasis>
</taxonomicName>
here (
<figureCitation id="135B2A38223BEA7DFC92F8D1FC6563BD" box="[876,992,1906,1930]" captionStart-0="Figs 813" captionStart-1="Figs 1417" captionStartId-0="3.[226,263,1219,1238]" captionStartId-1="3.[226,262,2017,2036]" captionTargetBox-0="[190,1399,315,1205]" captionTargetBox-1="[206,1402,1315,2000]" captionTargetId-0="figure-120@3.[189,1399,314,1206]" captionTargetId-1="figure-118@3.[198,1408,1314,2008]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionText-0="Figs 813. Aedeagus of Exomala spp: 810 — E. pallidipennis; 1113 — E. orientalis; 8, 11 — ventral view; 9, 12 — dorsal view; 10, 13 — lateral view. Рис. 813. Эдеагус Exomala spp: 810 — E. pallidipennis; 1113 — E. orientalis; 8, 11 — сниЗу; 9, 12 — сверху; 10, 13 — сбоку." captionText-1="Figs 1417. Endophallus of Exomala spp: 1415 — E. pallidipennis; 1617 — E. orientalis; 14, 16 — lateral view; 15, 17 — dorsal view. Рис. 1417. Эндофаллус Exomala spp: 1415 — E. pallidipennis; 1617 — E. orientalis; 14, 16 — сбоку; 15, 17 — сверху." pageId="2" pageNumber="55">Figs 817</figureCitation>
).
</paragraph>
<paragraph id="8BDF36BD223BEA7DFCACF80EFC536C24" blockId="2.[812,1399,1965,2067]" pageId="2" pageNumber="55">
<emphasis id="B914EAAF223BEA7DFCACF80EFB9063F4" bold="true" box="[850,1045,1965,1987]" pageId="2" pageNumber="55">Acknowledgments.</emphasis>
<materialsCitation id="3B083CE0223BEA7DFBE3F80EFC576C24" collectorName="Dr. Dmitry Telnov &amp; Dr. Keita Matsumoto" country="United Kingdom" location="Natural History Museum" municipality="London" pageId="2" pageNumber="55" specimenCount="1">
We are grateful
<collectorName id="2695536B223BEA7DFB43F80EFAF363F4" box="[1213,1398,1965,1987]" pageId="2" pageNumber="55">Dr. Dmitry Telnov</collectorName>
and
<collectorName id="2695536B223BEA7DFCA7F86BFBAA63EA" box="[857,1071,1992,2014]" pageId="2" pageNumber="55">Dr. Keita Matsumoto</collectorName>
(both
<location id="8EBF6066223BEA7DFB89F86BFAF663EA" LSID="urn:lsid:plazi:treatment:03C987AB2238EA7DFC85FED2FC536C24:8EBF6066223BEA7DFB89F86BFAF663EA" box="[1143,1395,1992,2014]" country="United Kingdom" municipality="London" name="Natural History Museum" pageId="2" pageNumber="55">Natural History Museum</location>
,
<collectingMunicipality id="6BBBACC7223BEA7DFCD2F841FCFF63CF" box="[812,890,2018,2040]" pageId="2" pageNumber="55">London</collectingMunicipality>
,
<collectingCountry id="F377762D223BEA7DFC7AF841FBAD63CF" box="[900,1064,2018,2040]" name="United Kingdom" pageId="2" pageNumber="55">United Kingdom</collectingCountry>
) for provided photos of
<typeStatus id="54DB881F223BEA7DFAEAF840FABB63CF" box="[1300,1342,2019,2040]" pageId="2" pageNumber="55">type</typeStatus>
specimens and labels
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>