243 lines
34 KiB
XML
243 lines
34 KiB
XML
<document id="35E018C182B92E74EF90AEDD0A436DC3" ID-DOI="10.11646/zootaxa.5004.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5120713" ID-ZooBank="E5AB5F1C-CA71-48D7-95AE-C982C6825AB1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626901326317" checkinUser="plazi" docAuthor="Medrano, Miguel, Kury, Adriano B. & Martinez, Leonel" docDate="2021" docId="03C787D4D642FF9EFF27FC03FC6EFED5" docLanguage="en" docName="zootaxa.5004.3.2.pdf" docOrigin="Zootaxa 5004 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmetus villarreali Medrano & Kury & Martinez 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="441" masterDocId="FFFEFFACD64BFF95FFB0FF9AFF90FFC8" masterDocTitle="A fresh look at Cosmetus Perty, 1833 (Opiliones: Cosmetidae), with new synonymies and description of two new species from Colombia" masterLastPageNumber="446" masterPageNumber="430" pageNumber="439" updateTime="1699012855954" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5698A99F4C34A1E3869865BF316C151A">A fresh look at Cosmetus Perty, 1833 (Opiliones: Cosmetidae), with new synonymies and description of two new species from Colombia</mods:title>
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<mods:name id="54943E02038BAE17311366831160F2EE" type="personal">
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<mods:namePart id="78C0FF3612FA4A499013ECAC96CE8478">Medrano, Miguel</mods:namePart>
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<mods:nameIdentifier id="F738D7C97BBB5A48D3A80D4386828D50" type="ORCID">0000-0002-3215-3592</mods:nameIdentifier>
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<mods:affiliation id="915F59C7FE28D76F6B4BC4261A58E1B9">Departamento de Invertebrados, Museu Nacional / UFRJ, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro - RJ, Brazil Grupo de Investigación Biodiversidad del Caribe Colombiano, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia & miangelmed @ gmail. com; https: // orcid. org / 0000 - 0002 - 3215 - 3592</mods:affiliation>
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<mods:nameIdentifier id="4A368291E20A8BC3F566B65F8AC58438" type="email">miangelmed@gmail.com</mods:nameIdentifier>
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<mods:name id="EE55A1F1DC85E8E5046E330A8C5267BC" type="personal">
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<mods:roleTerm id="4AF9329B7FD092B707D4594674B0CB8E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D75F98278809FFF65A4618DC05A38F9">Kury, Adriano B.</mods:namePart>
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<mods:nameIdentifier id="39E439A7422BB85D4D2899E1A4E32163" type="ORCID">0000-0002-8334-6204</mods:nameIdentifier>
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<mods:affiliation id="65B51C60D56BF7096C1F77E80606F15E">Departamento de Invertebrados, Museu Nacional / UFRJ, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro - RJ, Brazil Grupo de Investigación Biodiversidad del Caribe Colombiano, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia & adrianok @ gmail. com; https: // orcid. org / 0000 - 0002 - 8334 - 6204</mods:affiliation>
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<mods:nameIdentifier id="DF7A28BDEFF2F5BE4E4B99F2B29101B9" type="email">adrianok@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="0988618E48D13DE9A5DAA50981620C93">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0FD4DDFFA8C9D737E1E6A5908E57F146">Martinez, Leonel</mods:namePart>
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<mods:nameIdentifier id="04E06D741133B4B15ABF8E996C89B468" type="ORCID">0000-0002-4166-0561</mods:nameIdentifier>
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<mods:affiliation id="4E0C171F5B03E1791952E9AA5952EB67">leonelmarbio @ gmail. com; https: // orcid. org / 0000 - 0002 - 4166 - 0561</mods:affiliation>
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<mods:nameIdentifier id="6BD148FF7616A89FE82E9903A2ABB0F5" type="email">leonelmarbio@gmail.com</mods:nameIdentifier>
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<mods:title id="0861D888284FEEFCC334996247E600C1">Zootaxa</mods:title>
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<mods:part id="C6ABCBFDD4BCC06E36A248D12E18E8E5">
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<mods:date id="C86792054B1F0759A58ACF93FA2AA47D">2021</mods:date>
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<mods:number id="E1FD35E286F21FC4053B7EB5768C04AF">2021-07-21</mods:number>
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<mods:number id="31E6936A647CCE6DB2D0C71D74329A68">5004</mods:number>
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<mods:classification id="77E1014E9F5F04414C172F4D3B4194FD">journal article</mods:classification>
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<mods:identifier id="D14561663320289ACEFEDCD6ABB716FF" type="DOI">10.11646/zootaxa.5004.3.2</mods:identifier>
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<mods:identifier id="040E213A0B61D588E941245A27DB4447" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="90F6A9538764B07805A4C087C508970F" type="Zenodo-Dep">5120713</mods:identifier>
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<treatment id="03C787D4D642FF9EFF27FC03FC6EFED5" ID-DOI="http://doi.org/10.5281/zenodo.5120708" ID-Zenodo-Dep="5120708" LSID="urn:lsid:plazi:treatment:03C787D4D642FF9EFF27FC03FC6EFED5" httpUri="http://treatment.plazi.org/id/03C787D4D642FF9EFF27FC03FC6EFED5" lastPageId="11" lastPageNumber="441" pageId="9" pageNumber="439">
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<subSubSection id="C3746549D642FF9CFF27FC03FE60FC7B" box="[151,496,921,948]" pageId="9" pageNumber="439" type="nomenclature">
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<paragraph id="8BD136C2D642FF9CFF27FC03FE60FC7B" blockId="9.[151,541,921,983]" box="[151,496,921,948]" pageId="9" pageNumber="439">
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<heading id="D09981AED642FF9CFF27FC03FE60FC7B" bold="true" box="[151,496,921,948]" fontSize="11" level="1" pageId="9" pageNumber="439" reason="1">
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<emphasis id="B91AEAD0D642FF9CFF27FC03FE60FC7B" bold="true" box="[151,496,921,948]" pageId="9" pageNumber="439">
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<taxonomicName id="4C6E4D41D642FF9CFF27FC03FE1BFC7C" ID-CoL="9X6LC" authority="Medrano & Kury & Martinez, 2021" authorityName="Medrano & Kury & Martinez" authorityYear="2021" box="[151,395,921,948]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="439" phylum="Arthropoda" rank="species" species="villarreali" status="sp. nov.">
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||
<emphasis id="B91AEAD0D642FF9CFF27FC03FE1BFC7C" bold="true" box="[151,395,921,948]" italics="true" pageId="9" pageNumber="439">Cosmetus villarreali</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957ABD642FF9CFE22FC03FE60FC7B" box="[402,496,921,947]" pageId="9" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3746549D642FF9CFF27FC27FD8DFC1F" box="[151,541,957,983]" pageId="9" pageNumber="439" type="description">
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<paragraph id="8BD136C2D642FF9CFF27FC27FD8DFC1F" blockId="9.[151,541,921,983]" box="[151,541,957,983]" pageId="9" pageNumber="439">
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<figureCitation id="13552A47D642FF9CFF27FC27FF6EFC1F" box="[151,254,957,983]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Figures 8</figureCitation>
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A−D, 9A−H, 10A−C, 11.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3746549D642FF9CFF27FB9FFC41FBAF" pageId="9" pageNumber="439" type="materials_examined">
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<paragraph id="8BD136C2D642FF9CFF27FB9FFC41FBAF" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
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<emphasis id="B91AEAD0D642FF9CFF27FB9FFE85FBD7" bold="true" box="[151,277,1029,1055]" pageId="9" pageNumber="439">Type data.</emphasis>
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<materialsCitation id="3B063C9FD642FF9CFEADFB9CFB18FB8B" accessionNumber="ENT 47571" collectingDate="2007-04-26" collectionCode="MPUJ, ENT" collectorName="Jardin Botanico El Darien & J. Amaya" country="Colombia" county="Jardin Botanico El Darien" latitude="8.63975" location="Choco" longLatPrecision="1" longitude="-77.35042" municipality="Vereda Los Rios" pageId="9" pageNumber="439" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<specimenCount id="9D68FD4BD642FF9CFEADFB9CFEA2FBE8" box="[285,306,1030,1056]" pageId="9" pageNumber="439" type="male">♂</specimenCount>
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<typeStatus id="54D58860D642FF9CFE8AFB9FFE0CFBD7" box="[314,412,1029,1055]" pageId="9" pageNumber="439" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED7FAE07D642FF9CFE1BFB9FFE63FBD7" box="[427,499,1029,1055]" pageId="9" pageNumber="439">MPUJ</collectionCode>
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_
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<collectionCode id="ED7FAE07D642FF9CFDB1FB9FFDA8FBD7" box="[513,568,1029,1055]" country="Uganda" lsid="urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="9" pageNumber="439" type="Herbarium">ENT</collectionCode>
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47571) from
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<collectingCountry id="F3797652D642FF9CFD65FB9FFCD6FBD7" box="[725,838,1029,1055]" name="Colombia" pageId="9" pageNumber="439">Colombia</collectingCountry>
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,
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||
<location id="8EB16019D642FF9CFCE2FB9FFC0EFBD7" LSID="urn:lsid:plazi:treatment:03C787D4D642FF9EFF27FC03FC6EFED5:8EB16019D642FF9CFCE2FB9FFC0EFBD7" box="[850,926,1029,1055]" country="Colombia" county="Jardin Botanico El Darien" latitude="8.63975" longLatPrecision="1" longitude="-77.35042" municipality="Vereda Los Rios" name="Choco" pageId="9" pageNumber="439">Chocó</location>
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||
,
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||
<location id="8EB16019D642FF9CFC18FB9FFC6CFBD7" LSID="urn:lsid:plazi:treatment:03C787D4D642FF9EFF27FC03FC6EFED5:8EB16019D642FF9CFC18FB9FFC6CFBD7" box="[936,1020,1029,1055]" country="Colombia" county="Jardin Botanico El Darien" latitude="8.63975" longLatPrecision="1" longitude="-77.35042" municipality="Vereda Los Rios" name="Acandi" pageId="9" pageNumber="439">Acandí</location>
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||
,
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||
<location id="8EB16019D642FF9CFBB8FB9FFB14FBD7" LSID="urn:lsid:plazi:treatment:03C787D4D642FF9EFF27FC03FC6EFED5:8EB16019D642FF9CFBB8FB9FFB14FBD7" box="[1032,1156,1029,1055]" country="Colombia" county="Jardin Botanico El Darien" latitude="8.63975" longLatPrecision="1" longitude="-77.35042" municipality="Vereda Los Rios" name="Capurgana" pageId="9" pageNumber="439">Capurganá</location>
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,
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<collectingMunicipality id="6BB5ACB8D642FF9CFB3FFB9FFADCFBD7" box="[1167,1356,1029,1056]" pageId="9" pageNumber="439">Vereda Los Rios</collectingMunicipality>
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,
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<collectorName id="269B5314D642FF9CFAE8FB9FFEE1FB8B" pageId="9" pageNumber="439">
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||
<collectingCounty id="62B04E4ED642FF9CFAE8FB9FFEE1FB8B" pageId="9" pageNumber="439">Jardín Botánico El Darién</collectingCounty>
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</collectorName>
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(
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<geoCoordinate id="EE5A5005D642FF9CFE31FBB3FD82FB8B" box="[385,530,1065,1091]" degrees="8" direction="north" minutes="38" orientation="latitude" pageId="9" pageNumber="439" precision="1" seconds="23.1" value="8.63975">8°38’23.1”N</geoCoordinate>
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<geoCoordinate id="EE5A5005D642FF9CFDAAFBB3FD52FB8C" box="[538,706,1065,1092]" degrees="77" direction="west" minutes="21" orientation="longitude" pageId="9" pageNumber="439" precision="1" seconds="01.5" value="-77.35042">77°21’01.5”W</geoCoordinate>
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),
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<date id="FFD01002D642FF9CFD67FBB3FCC0FB8B" box="[727,848,1065,1091]" pageId="9" pageNumber="439" value="2007-04-26">
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||
<collectingDate id="EF94E9EAD642FF9CFD67FBB3FCC0FB8B" box="[727,848,1065,1091]" pageId="9" pageNumber="439" value="2007-04-26">
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||
26.iv.
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<quantity id="4C969B27D642FF9CFCA3FBB3FCC0FB8B" box="[787,848,1065,1091]" metricMagnitude="6" metricUnit="m" metricValue="2.00727" pageId="9" pageNumber="439" unit="m" value="2007270.0">2007</quantity>
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||
</collectingDate>
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||
</date>
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||
, 270 m,
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||
<collectorName id="269B5314D642FF9CFC02FBB0FB8BFB8B" box="[946,1051,1065,1092]" pageId="9" pageNumber="439">J. Amaya</collectorName>
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||
et. al leg
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B063C9FD642FF9CFB23FBB0FF2FFBAF" accessionNumber="ENT 47572" collectionCode="MPUJ, ENT" country="Colombia" pageId="9" pageNumber="439" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<specimenCount id="9D68FD4BD642FF9CFB23FBB0FB38FB8C" box="[1171,1192,1066,1092]" pageId="9" pageNumber="439" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7FAE07D642FF9CFB08FBB3FA90FB8B" box="[1208,1280,1065,1091]" pageId="9" pageNumber="439">MPUJ</collectionCode>
|
||
_
|
||
<collectionCode id="ED7FAE07D642FF9CFABEFBB3FAD5FB8B" box="[1294,1349,1065,1091]" country="Uganda" lsid="urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="9" pageNumber="439" type="Herbarium">ENT</collectionCode>
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||
47572) and
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||
</materialsCitation>
|
||
<materialsCitation id="3B063C9FD642FF9CFF77FBD4FC5EFBAF" accessionNumber="ENT 47570" box="[199,974,1101,1128]" collectingDate="2007-04-26" collectionCode="MPUJ, ENT" collectorName="C. Caltra" country="Colombia" county="Jardin Botanico El Darien" latitude="8.63975" location="Choco" longLatPrecision="1" longitude="-77.35042" municipality="Vereda Los Rios" pageId="9" pageNumber="439" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<specimenCount id="9D68FD4BD642FF9CFF77FBD4FF4CFBA0" box="[199,220,1102,1128]" pageId="9" pageNumber="439" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED7FAE07D642FF9CFF5AFBD7FEA2FBAF" box="[234,306,1101,1127]" pageId="9" pageNumber="439">MPUJ</collectionCode>
|
||
_
|
||
<collectionCode id="ED7FAE07D642FF9CFEF1FBD7FEE8FBAF" box="[321,376,1101,1127]" country="Uganda" lsid="urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="9" pageNumber="439" type="Herbarium">ENT</collectionCode>
|
||
47570)
|
||
<collectorName id="269B5314D642FF9CFE65FBD7FDA9FBAF" box="[469,569,1101,1127]" pageId="9" pageNumber="439">C. Caltra</collectorName>
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leg. with same data as the holotype
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</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C3746549D642FF9CFF77FBEBFDEBFB78" pageId="9" pageNumber="439" type="etymology">
|
||
<paragraph id="8BD136C2D642FF9CFF77FBEBFDEBFB78" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
|
||
<emphasis id="B91AEAD0D642FF9CFF77FBEBFEDEFB43" bold="true" box="[199,334,1137,1163]" pageId="9" pageNumber="439">Etymology.</emphasis>
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||
In honor of Osvaldo Villarreal, an arachnologist who made important contributions to the systematics of
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<taxonomicName id="4C6E4D41D642FF9CFF6CFB0FFEDDFB67" authorityName="Sundevall" authorityYear="1833" box="[220,333,1173,1199]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="439" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
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||
. Noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3746549D642FF9CFF77FB23FA01FA63" pageId="9" pageNumber="439" type="diagnosis">
|
||
<paragraph id="8BD136C2D642FF9CFF77FB23FA01FA63" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
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||
<emphasis id="B91AEAD0D642FF9CFF77FB23FED2FB1B" bold="true" box="[199,322,1209,1235]" pageId="9" pageNumber="439">Diagnosis.</emphasis>
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<taxonomicName id="4C6E4D41D642FF9CFEFAFB20FDA1FB1B" authority="Medrano & Kury & Martinez, 2021" authorityName="Medrano & Kury & Martinez" authorityYear="2021" box="[330,561,1209,1236]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="439" phylum="Arthropoda" rank="species" species="villarreali" status="sp. nov.">
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<emphasis id="B91AEAD0D642FF9CFEFAFB20FDA1FB1B" box="[330,561,1209,1236]" italics="true" pageId="9" pageNumber="439">Cosmetus villarreali</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="B91AEAD0D642FF9CFD89FB23FD05FB1B" bold="true" box="[569,661,1209,1235]" pageId="9" pageNumber="439">
|
||
<taxonomicNameLabel id="A22957ABD642FF9CFD89FB23FD05FB1B" box="[569,661,1209,1235]" pageId="9" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
|
||
differs from its congeners by having paired yellow spots in laterals of cephalothorax, posterior to ocularium, and by the shape of ventral plate of the penis: with a deep U-cleft and quadrangular latero-distal borders (
|
||
<figureCitation id="13552A47D642FF9CFD94FA9BFD1AFAD3" box="[548,650,1281,1307]" captionStart="FIGURE 10" captionStartId="12.[151,250,1148,1172]" captionTargetBox="[151,1437,382,1125]" captionTargetId="figure-119@12.[151,1437,381,1125]" captionTargetPageId="12" captionText="FIGURE 10A−C. Cosmetus villarreali sp. nov.: male (MPUJ_ENT 47570), penis. (A) dorsal view. (B) latero-dorsal view. (C) same, lateral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5120735" httpUri="https://zenodo.org/record/5120735/files/figure.png" pageId="9" pageNumber="439">Fig. 10C</figureCitation>
|
||
), and by the combination of the following features: (1) dorsal scutum beta-type (
|
||
<figureCitation id="13552A47D642FF9CFEA0FABFFE00FA88" box="[272,400,1317,1344]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Fig. 8A, D</figureCitation>
|
||
), (2) spine narrow, acuminate and divided, only fused at the base, length of the spine (SL) slightly shorter than dorsal scutum length (
|
||
<figureCitation id="13552A47D642FF9CFD30FAD3FD52FAAB" box="[640,706,1353,1379]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Fig. 8</figureCitation>
|
||
B−C), (3) femur IV long and unarmed, (4) male with hypertelic chelicerae, beta male has little incrassate chelicerae but larger than in female (
|
||
<figureCitation id="13552A47D642FF9CFBB3FAF7FBF0FA4F" box="[1027,1120,1389,1415]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Figs 8A</figureCitation>
|
||
,
|
||
<figureCitation id="13552A47D642FF9CFBDCFAF7FBE9FA4F" box="[1132,1145,1389,1415]" captionStart="FIGURE 9" captionStartId="11.[151,250,1579,1603]" captionTargetBox="[151,1436,309,1556]" captionTargetId="figure-83@11.[151,1436,309,1556]" captionTargetPageId="11" captionText="FIGURE 9A−I. Cosmetus villarreali sp. nov.: (A) alpha male, holotype (MPUJ_ENT 47571), left basichelicera, dorsal view. (B) same, left cheliceral hand, dorsal view. (C) beta male (MPUJ_ENT 47570), left basichelicera, dorsal view. (D) same, left cheliceral hand, dorsal view. (E) female (MPUJ_ENT 47572), left basichelicera, dorsal view. (F) same, left cheliceral hand, dorsal view. (G–I) alpha male (MPUJ_ENT 47571). (G) left pedipalp, tibia and tarsus, mesal view. (H) left pedipalp, femur and patella, ventral view. (I) left leg I, metatarsus and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120733" httpUri="https://zenodo.org/record/5120733/files/figure.png" pageId="9" pageNumber="439">9</figureCitation>
|
||
A−F), both forms of male with a triangular tooth in movable finger (6) coxa IV not visible in dorsal view and without groin warts (
|
||
<figureCitation id="13552A47D642FF9CFA9AFA0BFA14FA63" box="[1322,1412,1425,1451]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Fig. 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3746549D642FF9EFF77FA2FFC6EFED5" lastPageId="11" lastPageNumber="441" pageId="9" pageNumber="439" type="description">
|
||
<paragraph id="8BD136C2D642FF9CFF77FA2FFC73FA3B" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
|
||
<emphasis id="B91AEAD0D642FF9CFF77FA2FFD89FA07" bold="true" box="[199,537,1461,1487]" pageId="9" pageNumber="439">Description: Measurements.</emphasis>
|
||
CL: 1.18; CW: 3.07; AL: 4.24; CW: 3.05; IOD: 0.80; FeI: 5.65; TiI: 3.73; FeII: 12.94; TiII: 10.06; FeIII: 9.04; TiIII: 4.19; FeIV: 8.21; TiIV: 6.08; SL: 2.40.
|
||
</paragraph>
|
||
<paragraph id="8BD136C2D642FF9CFF77FA67FD83F927" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
|
||
<emphasis id="B91AEAD0D642FF9CFF77FA67FE95F9DF" bold="true" box="[199,261,1533,1559]" pageId="9" pageNumber="439">Body</emphasis>
|
||
(
|
||
<figureCitation id="13552A47D642FF9CFEA6FA67FEC7F9DF" box="[278,343,1533,1559]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Fig. 8</figureCitation>
|
||
A−D). Median cosmetid (
|
||
<quantity id="4C969B27D642FF9CFDCCFA67FD2FF9D0" box="[636,703,1533,1560]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="439" unit="mm" value="4.0">4 mm</quantity>
|
||
of body length) with slender legs. Outline of dorsal scutum
|
||
<typeStatus id="54D58860D642FF9CFADCFA64FA0CF9D0" box="[1388,1436,1534,1560]" pageId="9" pageNumber="439">type</typeStatus>
|
||
beta (β) with constrictions barely marked, with triangular ectal projections of cheliceral sockets. Scutal grooves not visible. All scutum unarmed, except for area III with two contiguous large spines fused in the base slanted backwards with a narrow base (a third of abdomen length). Abdomen uniformly granulate even over spine of area III and carapace poorly granulate, only some granules in the posterior region. Eyes slightly separated (one third of carapace width), ocularium with medial depression. Free tergites each with a transverse row of minute granules and anal operculum weakly granulate.
|
||
</paragraph>
|
||
<paragraph id="8BD136C2D642FF9CFF77F963FACEF8F0" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
|
||
<emphasis id="B91AEAD0D642FF9CFF77F963FF6FF8DB" bold="true" box="[199,255,1785,1811]" pageId="9" pageNumber="439">Legs</emphasis>
|
||
(
|
||
<figureCitation id="13552A47D642FF9CFEBEF963FEC2F8DB" box="[270,338,1785,1811]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Figs 8</figureCitation>
|
||
A−D, 9I). Coxa IV uniformly granulated excepted by a distal apophysis formed by some granules. All femurs straight and without ornamentation. Smooth claws III and IV. Tarsal counts: 7(3), 17(3)–?, 13, 15.
|
||
</paragraph>
|
||
<paragraph id="8BD136C2D642FF9CFF77F8DBFD3BF80F" blockId="9.[151,1437,1029,1992]" pageId="9" pageNumber="439">
|
||
<emphasis id="B91AEAD0D642FF9CFF77F8DBFEAAF893" bold="true" box="[199,314,1857,1883]" pageId="9" pageNumber="439">Chelicera</emphasis>
|
||
(
|
||
<figureCitation id="13552A47D642FF9CFEFCF8DBFE03F893" box="[332,403,1857,1883]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="9" pageNumber="439">Figs 8</figureCitation>
|
||
A−C, 9A−F). Basichelicerite tuberculated in basal and ectal sides, three tubercules bigger in ecto-basal corner. Two small tubercles in antero-mesal corner. Cheliceral hand incrassate, kidney-like, forming a dorsal protuberance. Fixed finger with a keel formed by seven six fused tubercles and movable finger with nine tubercles and basally with a triangular tubercle.
|
||
</paragraph>
|
||
<caption id="DF11664AD641FF9FFF27F9F5FBE4F963" ID-DOI="http://doi.org/10.5281/zenodo.5120731" ID-Zenodo-Dep="5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="10" pageNumber="440" startId="10.[151,250,1647,1671]" targetBox="[151,1436,182,1622]" targetPageId="10">
|
||
<paragraph id="8BD136C2D641FF9FFF27F9F5FBE4F963" blockId="10.[151,1437,1647,1708]" pageId="10" pageNumber="440">
|
||
<emphasis id="B91AEAD0D641FF9FFF27F9F5FED7F94F" bold="true" box="[151,327,1647,1671]" pageId="10" pageNumber="440">FIGURE 8A−D.</emphasis>
|
||
<taxonomicName id="4C6E4D41D641FF9FFEFFF9F5FD8FF94F" authority="Medrano & Kury & Martinez, 2021" authorityName="Medrano & Kury & Martinez" authorityYear="2021" box="[335,543,1647,1671]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="440" phylum="Arthropoda" rank="species" species="villarreali" status="sp. nov.">
|
||
<emphasis id="B91AEAD0D641FF9FFEFFF9F5FD8FF94F" box="[335,543,1647,1671]" italics="true" pageId="10" pageNumber="440">Cosmetus villarreali</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91AEAD0D641FF9FFD96F9F5FDE7F94F" bold="true" box="[550,631,1647,1671]" pageId="10" pageNumber="440">
|
||
<taxonomicNameLabel id="A22957ABD641FF9FFD96F9F5FDE7F94F" box="[550,631,1647,1671]" pageId="10" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD136C2D641FF9FFF77F943FF44F8F3" blockId="10.[151,1437,1753,2032]" pageId="10" pageNumber="440">
|
||
<emphasis id="B91AEAD0D641FF9FFF77F943FEACF93B" bold="true" box="[199,316,1753,1779]" pageId="10" pageNumber="440">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="13552A47D641FF9FFEFBF943FE56F93C" box="[331,454,1753,1780]" captionStart="FIGURE 9" captionStartId="11.[151,250,1579,1603]" captionTargetBox="[151,1436,309,1556]" captionTargetId="figure-83@11.[151,1436,309,1556]" captionTargetPageId="11" captionText="FIGURE 9A−I. Cosmetus villarreali sp. nov.: (A) alpha male, holotype (MPUJ_ENT 47571), left basichelicera, dorsal view. (B) same, left cheliceral hand, dorsal view. (C) beta male (MPUJ_ENT 47570), left basichelicera, dorsal view. (D) same, left cheliceral hand, dorsal view. (E) female (MPUJ_ENT 47572), left basichelicera, dorsal view. (F) same, left cheliceral hand, dorsal view. (G–I) alpha male (MPUJ_ENT 47571). (G) left pedipalp, tibia and tarsus, mesal view. (H) left pedipalp, femur and patella, ventral view. (I) left leg I, metatarsus and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120733" httpUri="https://zenodo.org/record/5120733/files/figure.png" pageId="10" pageNumber="440">Fig. 9G, H</figureCitation>
|
||
). Trochanter with strong ventral process. Femur with dorsally convex with some setiferous protuberances, ventrally with a row of 18 setiferous tubercles and mesodistal process. Tibia wider at the apical third.
|
||
</paragraph>
|
||
<paragraph id="8BD136C2D641FF9FFF77F8DFFE21F86F" blockId="10.[151,1437,1753,2032]" pageId="10" pageNumber="440">
|
||
<emphasis id="B91AEAD0D641FF9FFF77F8DFFE1CF897" bold="true" box="[199,396,1861,1887]" pageId="10" pageNumber="440">Color, in alcohol</emphasis>
|
||
(
|
||
<figureCitation id="13552A47D641FF9FFE2BF8DFFE4CF897" box="[411,476,1861,1887]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="10" pageNumber="440">Fig. 8</figureCitation>
|
||
A−D). Body uniformly vivid yellow (pale in
|
||
<typeStatus id="54D58860D641FF9FFC6AF8DFFBAEF8A8" box="[986,1086,1861,1888]" pageId="10" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
) with two unpigmented dots in posterior laterals of carapace. Legs with the same color of dorsal scutum except for Ta and tarsomeres of legs I and II with darker coloration.
|
||
</paragraph>
|
||
<paragraph id="8BD136C2D641FF9EFF77F82BFCB4FF1D" blockId="10.[151,1437,1753,2032]" lastBlockId="11.[151,1437,151,285]" lastPageId="11" lastPageNumber="441" pageId="10" pageNumber="440">
|
||
<emphasis id="B91AEAD0D641FF9FFF77F82BFEE1F803" bold="true" box="[199,369,1969,1995]" pageId="10" pageNumber="440">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="13552A47D641FF9FFECFF82BFE5EF803" box="[383,462,1969,1995]" captionStart="FIGURE 10" captionStartId="12.[151,250,1148,1172]" captionTargetBox="[151,1437,382,1125]" captionTargetId="figure-119@12.[151,1437,381,1125]" captionTargetPageId="12" captionText="FIGURE 10A−C. Cosmetus villarreali sp. nov.: male (MPUJ_ENT 47570), penis. (A) dorsal view. (B) latero-dorsal view. (C) same, lateral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5120735" httpUri="https://zenodo.org/record/5120735/files/figure.png" pageId="10" pageNumber="440">Fig. 10</figureCitation>
|
||
A−C). Ventral plate of penis, in dorsal view, wider at the base. Distal border remarkably concave. Stylus with a large distal caruncle, no stylar barbs visible. Tip of the stylus with a rounded dorsal process. Two pairs of MS C flattened and curved; two pairs of straight MS D, distal one stronger; two pairs of small MS A in basal portion of ventral plate grouped with basal MS D.
|
||
</paragraph>
|
||
<paragraph id="8BD136C2D640FF9EFF77FF44FC6EFED5" blockId="11.[151,1437,151,285]" pageId="11" pageNumber="441">
|
||
<emphasis id="B91AEAD0D640FF9EFF77FF44FECEFF30" bold="true" box="[199,350,222,248]" pageId="11" pageNumber="441">Dimorphism</emphasis>
|
||
. Two
|
||
<typeStatus id="54D58860D640FF9EFE17FF45FE72FF31" box="[423,482,223,249]" pageId="11" pageNumber="441">types</typeStatus>
|
||
of males: alpha male with hypertelic chelicerae and beta male with slightly swollen chelicerae (
|
||
<figureCitation id="13552A47D640FF9EFEA7FE99FECCFED5" box="[279,348,259,285]" captionStart="FIGURE 8" captionStartId="10.[151,250,1647,1671]" captionTargetBox="[151,1436,182,1622]" captionTargetId="figure-18@10.[151,1436,181,1623]" captionTargetPageId="10" captionText="FIGURE 8A−D. Cosmetus villarreali sp. nov.: (A) male holotype (MPUJ_ENT 47571), dorsal view. (B) same, frontal view. (C) same, lateral view. (D). female paratype (MPUJ_ENT 47571), lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120731" httpUri="https://zenodo.org/record/5120731/files/figure.png" pageId="11" pageNumber="441">Figs 8</figureCitation>
|
||
A−D, 9A−F). Male with basitarsomeres incrassate (
|
||
<figureCitation id="13552A47D640FF9EFC12FE99FC61FED5" box="[930,1009,259,285]" captionStart="FIGURE 9" captionStartId="11.[151,250,1579,1603]" captionTargetBox="[151,1436,309,1556]" captionTargetId="figure-83@11.[151,1436,309,1556]" captionTargetPageId="11" captionText="FIGURE 9A−I. Cosmetus villarreali sp. nov.: (A) alpha male, holotype (MPUJ_ENT 47571), left basichelicera, dorsal view. (B) same, left cheliceral hand, dorsal view. (C) beta male (MPUJ_ENT 47570), left basichelicera, dorsal view. (D) same, left cheliceral hand, dorsal view. (E) female (MPUJ_ENT 47572), left basichelicera, dorsal view. (F) same, left cheliceral hand, dorsal view. (G–I) alpha male (MPUJ_ENT 47571). (G) left pedipalp, tibia and tarsus, mesal view. (H) left pedipalp, femur and patella, ventral view. (I) left leg I, metatarsus and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5120733" httpUri="https://zenodo.org/record/5120733/files/figure.png" pageId="11" pageNumber="441">Fig. 9I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |