360 lines
61 KiB
XML
360 lines
61 KiB
XML
<document id="96FB997E7891713B74BE13FFEB523E2C" ID-DOI="10.1080/00222933.2014.961987" ID-ISSN="1464-5262" ID-Zenodo-Dep="4607733" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615896067452" checkinUser="carolina" docAuthor="Kocot, Kevin M. & Todt, Christiane" docDate="2014" docId="03C7536DFFFBFFC8FE16FD1BFB9A2D06" docLanguage="en" docName="JNATHIST.48.45-48.3007-3031.pdf" docOrigin="Journal of Natural History 48 (45 - 48)" docSource="http://dx.doi.org/10.1080/00222933.2014.961987" docStyle="DocumentStyle:CD936EBDDF5274E0936345450CC2DAE4.5:JNatHist.2013-2015.journal_article.1cover" docStyleId="CD936EBDDF5274E0936345450CC2DAE4" docStyleName="JNatHist.2013-2015.journal_article.1cover" docStyleVersion="5" docTitle="Macellomenia schanderi Kocot & Todt 2014, sp. nov." docType="treatment" docVersion="3" lastPageNumber="3016" masterDocId="FFFE2B15FFFEFFC2FFEAFFC7FFBA2F44" masterDocTitle="Three new meiofaunal solenogaster species (Mollusca: Aplacophora) from the north-east Pacific" masterLastPageNumber="3031" masterPageNumber="3007" pageNumber="3011" updateTime="1698886697010" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="48963DBAAEC34D52F73C640AE1D8EC1F">Three new meiofaunal solenogaster species (Mollusca: Aplacophora) from the north-east Pacific</mods:title>
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<mods:name id="7E8D4F81025F62C563BA70CCD0110766" type="personal">
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<mods:roleTerm id="B556470728B8FAD3C1834309D7439F16">Author</mods:roleTerm>
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<mods:namePart id="B8EC6CF51D15AA254668755734DE4A59">Kocot, Kevin M.</mods:namePart>
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<mods:affiliation id="BF0D33C377D47A2CA69EBCAC130DDEB8">Department of Biological Sciences, Auburn University, Auburn, AL, USA;</mods:affiliation>
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<mods:roleTerm id="ADAC7BCBFD95A1A42F602AE6B07A4C3F">Author</mods:roleTerm>
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<mods:namePart id="3197DD5474716DEF1C3D8642BE96E839">Todt, Christiane</mods:namePart>
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<mods:affiliation id="A5FECCF2C6229A83842313DEB8F31CF5">University Museum of Bergen - The Natural History Collections, University of Bergen, Bergen, Norway</mods:affiliation>
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<mods:typeOfResource id="A249962F20A1E9A269F28836FC7DB275">text</mods:typeOfResource>
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<mods:title id="13697B56A5E10A98244DEE5178F5E236">Journal of Natural History</mods:title>
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<mods:part id="E1A766927994639147937FA13FAC6B22">
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<mods:date id="561DD4022DCDA2D241A4617C747C93DC">2014</mods:date>
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<mods:number id="6FBFD4B8F96532A853EFA06796707A2E">2014-10-13</mods:number>
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<mods:detail id="4FB80C66266DE602D23BA11DF4F54D1C" type="volume">
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<mods:number id="42A8686629B877F2E36F1ED2BE87F3EF">48</mods:number>
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<mods:number id="5FE1EECF1878F2656E4C86FAED2C8C4D">45 - 48</mods:number>
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<mods:start id="1AECFA7071E570474AD77D4C9D691449">3007</mods:start>
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<mods:url id="8871B8B9744B4182715202309875C932">http://dx.doi.org/10.1080/00222933.2014.961987</mods:url>
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<mods:classification id="E2AAB6953CDB795AC24025607F00138D">journal article</mods:classification>
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<mods:identifier id="6E529C1ED7FE38AEC57BDF82CEB0A213" type="DOI">10.1080/00222933.2014.961987</mods:identifier>
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<mods:identifier id="76D5EF6C1E90A9FBDE31348547DD201F" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03C7536DFFFBFFC8FE16FD1BFB9A2D06" LSID="urn:lsid:plazi:treatment:03C7536DFFFBFFC8FE16FD1BFB9A2D06" httpUri="http://treatment.plazi.org/id/03C7536DFFFBFFC8FE16FD1BFB9A2D06" lastPageId="10" lastPageNumber="3016" pageId="5" pageNumber="3011">
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<subSubSection id="C374B1F0FFFBFFC7FE16FD1BFCD62DB1" box="[508,876,732,757]" pageId="5" pageNumber="3011" type="nomenclature">
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<paragraph id="8BD1E27BFFFBFFC7FE16FD1BFCD62DB1" blockId="5.[508,876,732,788]" box="[508,876,732,757]" pageId="5" pageNumber="3011">
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<heading id="D0995517FFFBFFC7FE16FD1BFCD62DB1" box="[508,876,732,757]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="3011" reason="2">
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<taxonomicName id="4C6E99F8FFFBFFC7FE16FD1BFCB32DB0" ID-CoL="3WXV6" authority="Kocot & Todt, 2014" authorityName="Kocot & Todt" authorityYear="2014" box="[508,777,732,756]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="5" pageNumber="3011" phylum="Mollusca" rank="species" species="schanderi" status="sp. nov.">
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<emphasis id="B91A3E69FFFBFFC7FE16FD1BFCB32DB0" bold="true" box="[508,777,732,756]" italics="true" pageId="5" pageNumber="3011">Macellomenia schanderi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2298312FFFBFFC7FCF8FD1AFCD62DB1" box="[786,876,733,757]" pageId="5" pageNumber="3011" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C374B1F0FFFBFFC7FDD7FD3BFC902C50" box="[573,810,764,788]" pageId="5" pageNumber="3011" type="description">
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<paragraph id="8BD1E27BFFFBFFC7FDD7FD3BFC902C50" blockId="5.[508,876,732,788]" box="[573,810,764,788]" pageId="5" pageNumber="3011">
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(
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<figureCitation id="1355FEFEFFFBFFC7FDACFD3BFD5D2C50" box="[582,743,764,788]" captionStart="Figure 2" captionStartId="6.[144,211,1157,1179]" captionTargetBox="[144,1104,168,1125]" captionTargetId="figure-75@6.[144,1104,168,1125]" captionTargetPageId="6" captionText="Figure 2. (A) Three new meiofaunal solenogaster species from Reid Rock, Friday Harbor: Ms, Macellomenia schanderi; Mm, Macellomenia morseae; Hs, Hypomenia sanjuanensis; note the similar size, colour, and body shape. Scale bar 1 mm; (B) posterior body end of M. schanderi showing: asp, abdominal spicules. Scale bar 150 µm; (C, D) epidermal sclerites of H. sanjuanensis sp. nov., light micrographs, polarized filter; (C) note the criss-cross arrangement of sclerites (cuticle removed with household bleach); scale bar 50 µm; (D) sclerite types: a, dorsolateral sclerites are acicular with small or almost reduced cavity, often serrated; b, most anterior sclerites are recurved and serrated; c, ventral scales; d, ventral scales found close to the pallial cavity opening; scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607737" httpUri="https://zenodo.org/record/4607737/files/figure.png" pageId="5" pageNumber="3011">Figures 2A, B</figureCitation>
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,
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<figureCitation id="1355FEFEFFFBFFC7FD1DFD3BFCBE2C50" box="[759,772,764,788]" captionStart="Figure 3" captionStartId="7.[212,279,1306,1328]" captionTargetBox="[212,1172,168,1274]" captionTargetId="figure-1@7.[212,1172,168,1275]" captionTargetPageId="7" captionText="Figure 3. Macellomenia schanderi sp. nov. (A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="5" pageNumber="3011">3</figureCitation>
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,
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<figureCitation id="1355FEFEFFFBFFC7FCFEFD3BFC9B2C50" box="[788,801,764,788]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="5" pageNumber="3011">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C374B1F0FFFBFFC7FF3EFC9FFEF72A5D" pageId="5" pageNumber="3011" type="materials_examined">
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<paragraph id="8BD1E27BFFFBFFC7FF3EFC9FFD9E2C34" blockId="5.[212,1172,856,983]" box="[212,548,856,880]" pageId="5" pageNumber="3011">
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<heading id="D0995517FFFBFFC7FF3EFC9FFD9E2C34" box="[212,548,856,880]" fontSize="10" level="3" pageId="5" pageNumber="3011" reason="5">
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<emphasis id="B91A3E69FFFBFFC7FF3EFC9FFD9E2C34" box="[212,548,856,880]" italics="true" pageId="5" pageNumber="3011">
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Type material (
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<specimenCount id="9D6829F2FFFBFFC7FE68FC9FFDA62C34" box="[386,540,856,880]" pageId="5" pageNumber="3011" type="generic">10 specimens</specimenCount>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD1E27BFFFBFFC7FF3EFC47FEFB2C93" blockId="5.[212,1172,856,983]" pageId="5" pageNumber="3011">
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<materialsCitation id="3B06E826FFFBFFC7FF3EFC47FEFB2C93" collectionCode="ZMBN" country="Norway" county="Reid Rock" latitude="48.323887" location="University Museum of Bergen" longLatPrecision="18" longitude="122.585556" municipality="San Juan Channel" pageId="5" pageNumber="3011" specimenCount="1" typeStatus="holotype">
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<emphasis id="B91A3E69FFFBFFC7FF3EFC47FEF82CDC" box="[212,322,896,920]" italics="true" pageId="5" pageNumber="3011">
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<typeStatus id="54D55CD9FFFBFFC7FF3EFC47FE842CDC" box="[212,318,896,920]" pageId="5" pageNumber="3011" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED7F7ABEFFFBFFC7FEA1FCB8FE242CD3" box="[331,414,895,919]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">ZMBN</collectionCode>
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(
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<location id="8EB1B4A0FFFBFFC7FE5AFCB8FCAA2CD3" LSID="urn:lsid:plazi:treatment:03C7536DFFFBFFC8FE16FD1BFB9A2D06:8EB1B4A0FFFBFFC7FE5AFCB8FCAA2CD3" box="[432,784,895,919]" country="Norway" county="Reid Rock" latitude="48.323887" longLatPrecision="18" longitude="122.585556" municipality="San Juan Channel" name="University Museum of Bergen" pageId="5" pageNumber="3011">University Museum of Bergen</location>
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,
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<collectingCountry id="F379A2EBFFFBFFC7FCF7FCB8FCC02CD3" box="[797,890,895,919]" name="Norway" pageId="5" pageNumber="3011">Norway</collectingCountry>
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) 94118, histological section series (6 slides),
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||
<collectingMunicipality id="6BB57801FFFBFFC7FE50FC58FD3C2CF3" box="[442,646,927,951]" pageId="5" pageNumber="3011">San Juan Channel</collectingMunicipality>
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near
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<collectingCounty id="62B09AF7FFFBFFC7FD2EFC58FC842CF3" box="[708,830,927,951]" pageId="5" pageNumber="3011">Reid Rock</collectingCounty>
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(
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<figureCitation id="1355FEFEFFFBFFC7FCA7FC58FC112CF3" box="[845,939,927,951]" captionStart="Figure 1" captionStartId="3.[212,279,1135,1157]" captionTargetBox="[212,1171,169,1102]" captionTargetId="figure-182@3.[212,1172,168,1103]" captionTargetPageId="3" captionText="Figure 1. Schematic map of the San Juan Islands showing the type locality of the species described herein with a star." figureDoi="http://doi.org/10.5281/zenodo.4607735" httpUri="https://zenodo.org/record/4607735/files/figure.png" pageId="5" pageNumber="3011">Figure 1</figureCitation>
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);
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,
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<geoCoordinate id="EE5A84BCFFFBFFC7FB8BFC58FE872C93" degrees="122" direction="east" minutes="35" orientation="longitude" pageId="5" pageNumber="3011" precision="15" seconds="08" value="122.585556">122° 35' 08'' E</geoCoordinate>
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.
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</paragraph>
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<paragraph id="8BD1E27BFFFBFFC7FF3EFBC5FB2E2B9D" blockId="5.[212,1172,1026,1305]" pageId="5" pageNumber="3011">
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<materialsCitation id="3B06E826FFFBFFC7FF3EFBC5FC3F2B5E" box="[212,901,1026,1050]" collectionCode="ZMBN" pageId="5" pageNumber="3011" specimenCode="ZMBN 94119" specimenCount="1" typeStatus="paratype">
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<emphasis id="B91A3E69FFFBFFC7FF3EFBC5FEF02B5E" box="[212,330,1026,1050]" italics="true" pageId="5" pageNumber="3011">
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<typeStatus id="54D55CD9FFFBFFC7FF3EFBC5FEFD2B5E" box="[212,327,1026,1050]" pageId="5" pageNumber="3011" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBC84A00FFFBFFC7FEBBFBC5FE482B5E" box="[337,498,1026,1050]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">ZMBN 94119</specimenCode>
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, histological section series (6 slides)
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||
</materialsCitation>
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||
;
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<materialsCitation id="3B06E826FFFBFFC7FC7AFBC5FDF02B7E" collectionCode="ZMBN" pageId="5" pageNumber="3011" specimenCode="ZMBN 94120, 1" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DBC84A00FFFBFFC7FC7AFBC5FB8B2B5E" box="[912,1073,1026,1050]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">ZMBN 94120</specimenCode>
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,
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<specimenCode id="DBC84A00FFFBFFC7FBD6FBC5FBF32B5E" box="[1084,1097,1026,1050]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">1</specimenCode>
|
||
whole animal mounted on a glass slide
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B06E826FFFBFFC7FDB3FBE5FEA62B1E" collectionCode="ZMBN" pageId="5" pageNumber="3011" specimenCode="ZMBN 94121" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DBC84A00FFFBFFC7FDB3FBE5FD422B7E" box="[601,760,1058,1082]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">ZMBN 94121</specimenCode>
|
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and 94122, sclerite mounts (2 glass slides)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B06E826FFFBFFC7FEC3FB85FCB12B3E" collectionCode="ZMBN" pageId="5" pageNumber="3011" specimenCode="ZMBN 94123, 2" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DBC84A00FFFBFFC7FEC3FB85FE712B1E" box="[297,459,1090,1114]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">ZMBN 94123</specimenCode>
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,
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<specimenCode id="DBC84A00FFFBFFC7FE32FB85FE5F2B1E" box="[472,485,1090,1114]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="5" pageNumber="3011">2</specimenCode>
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||
whole animals fixed in 4% glutaraldehyde in 0.1 M sodium cacodylate buffer and preserved in 70% ethanol
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B06E826FFFBFFC7FCF6FBA5FE912BFE" collectionCode="USNM" location="Smithsonian Institution National Museum of Natural History" pageId="5" pageNumber="3011" specimenCount="1" typeStatus="paratype">
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<collectionCode id="ED7F7ABEFFFBFFC7FCF6FBA5FCCA2B3E" box="[796,880,1122,1146]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="3011" type="Museum">USNM</collectionCode>
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(
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<location id="8EB1B4A0FFFBFFC7FC6EFBA5FD3E2BDE" LSID="urn:lsid:plazi:treatment:03C7536DFFFBFFC8FE16FD1BFB9A2D06:8EB1B4A0FFFBFFC7FC6EFBA5FD3E2BDE" name="Smithsonian Institution National Museum of Natural History" pageId="5" pageNumber="3011">Smithsonian Institution National Museum of Natural History</location>
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) 1231346, 3 whole animals preserved in 100% ethanol
|
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</materialsCitation>
|
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;
|
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<materialsCitation id="3B06E826FFFBFFC7FED5FB65FB912BFE" box="[319,1067,1186,1210]" collectionCode="USNM" pageId="5" pageNumber="3011" specimenCode="USNM 1231345, 1" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DBC84A00FFFBFFC7FED5FB65FDB92BFE" box="[319,515,1186,1210]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="3011" type="Museum">USNM 1231345</specimenCode>
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,
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<specimenCode id="DBC84A00FFFBFFC7FDFDFB65FD9E2BFE" box="[535,548,1186,1210]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="3011" type="Museum">1</specimenCode>
|
||
whole animal mounted on an SEM stub
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B06E826FFFBFFC7FBD5FB65FB2A2B9D" collectionCode="USNM" collectorName="All" pageId="5" pageNumber="3011" specimenCode="USNM 1231347, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBC84A00FFFBFFC7FBD5FB65FE8E2B9D" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="3011" type="Museum">USNM 1231347</specimenCode>
|
||
,
|
||
<specimenCode id="DBC84A00FFFBFFC7FED5FB06FEF62B9D" box="[319,332,1217,1241]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="3011" type="Museum">1</specimenCode>
|
||
whole animal mounted on an SEM stub.
|
||
<collectorName id="269B87ADFFFBFFC7FCCFFB06FCF22B9D" box="[805,840,1217,1241]" pageId="5" pageNumber="3011">All</collectorName>
|
||
<typeStatus id="54D55CD9FFFBFFC7FCBAFB06FC072B9D" box="[848,957,1217,1241]" pageId="5" pageNumber="3011" type="paratype">paratypes</typeStatus>
|
||
from type locality
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFFBFFC7FF14FB25FEF72A5D" blockId="5.[212,1172,1026,1305]" pageId="5" pageNumber="3011">
|
||
<emphasis id="B91A3E69FFFBFFC7FF14FB25FDB62BBE" box="[254,524,1250,1274]" italics="true" pageId="5" pageNumber="3011">
|
||
Macellomena
|
||
<taxonomicName id="4C6E99F8FFFBFFC7FE4FFB25FDB62BBE" authorityName="Kocot & Todt" authorityYear="2014" box="[421,524,1250,1274]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="5" pageNumber="3011" phylum="Mollusca" rank="species" species="schanderi">schanderi</taxonomicName>
|
||
</emphasis>
|
||
full length COI sequence:
|
||
<collectionCode id="ED7F7ABEFFFBFFC7FC87FB26FC082BBD" box="[877,946,1249,1273]" pageId="5" pageNumber="3011">NCBI</collectionCode>
|
||
accession number
|
||
<accessionNumber id="943D7F98FFFBFFC7FF3EFAC6FEF32A5D" box="[212,329,1281,1305]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ568516" pageId="5" pageNumber="3011">KJ568516</accessionNumber>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374B1F0FFFBFFC7FF3EFA9BFE9629DF" pageId="5" pageNumber="3011" type="diagnosis">
|
||
<paragraph id="8BD1E27BFFFBFFC7FF3EFA9BFEFA2A30" blockId="5.[212,1172,1372,1691]" box="[212,320,1372,1396]" pageId="5" pageNumber="3011">
|
||
<heading id="D0995517FFFBFFC7FF3EFA9BFEFA2A30" box="[212,320,1372,1396]" fontSize="10" level="3" pageId="5" pageNumber="3011" reason="5">
|
||
<emphasis id="B91A3E69FFFBFFC7FF3EFA9BFEFA2A30" box="[212,320,1372,1396]" italics="true" pageId="5" pageNumber="3011">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFFBFFC7FF3EFA43FE9629DF" blockId="5.[212,1172,1372,1691]" pageId="5" pageNumber="3011">
|
||
Body up to around
|
||
<quantity id="4C964F9EFFFBFFC7FE54FA43FDB82AD8" box="[446,514,1412,1436]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="3011" unit="mm" value="2.0">2 mm</quantity>
|
||
long. Mantle sclerites up to
|
||
<quantity id="4C964F9EFFFBFFC7FCBDFA43FC1A2AD8" box="[855,928,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="3011" unit="mm" value="50.0">50 µm</quantity>
|
||
in length with ovate bases. Acicular spine of mantle sclerites flattened and broadest medially. Peripedal scales of one
|
||
<typeStatus id="54D55CD9FFFBFFC7FE86FA03FE252A98" box="[364,415,1476,1500]" pageId="5" pageNumber="3011">type</typeStatus>
|
||
, leaf-shaped and pointed. Atrium (= vestibulum) with unbranched sensory papillae. Mouth opening separate from atrium. Rostral midgut caecum present, frontally paired. Radula with seven equally sized denticles per tooth. Oesophagus longer than radula. Midgut without regular constrictions. Abdominal spicules present, attached to a retractable genital cone. Posterior mantle glands present. Mantle cavity without respiratory papillae. Dorsoterminal sense organ present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF11B2F3FFF8FFC4FF7AFB42FD932AC1" ID-DOI="http://doi.org/10.5281/zenodo.4607737" ID-Zenodo-Dep="4607737" httpUri="https://zenodo.org/record/4607737/files/figure.png" pageId="6" pageNumber="3012" startId="6.[144,211,1157,1179]" targetBox="[144,1104,168,1125]" targetPageId="6">
|
||
<paragraph id="8BD1E27BFFF8FFC4FF7AFB42FD932AC1" blockId="6.[144,1104,1157,1413]" pageId="6" pageNumber="3012">
|
||
Figure 2. (A) Three new meiofaunal solenogaster species from Reid Rock, Friday Harbor: Ms,
|
||
<taxonomicName id="4C6E99F8FFF8FFC4FF7AFB64FE392BFC" authorityName="Kocot & Todt" authorityYear="2014" box="[144,387,1187,1208]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="6" pageNumber="3012" phylum="Mollusca" rank="species" species="schanderi">
|
||
<emphasis id="B91A3E69FFF8FFC4FF7AFB64FE392BFC" box="[144,387,1187,1208]" italics="true" pageId="6" pageNumber="3012">Macellomenia schanderi</emphasis>
|
||
</taxonomicName>
|
||
; Mm,
|
||
<taxonomicName id="4C6E99F8FFF8FFC4FE25FB64FD0C2BFC" authorityName="Kocot & Todt" authorityYear="2014" box="[463,694,1187,1208]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="6" pageNumber="3012" phylum="Mollusca" rank="species" species="morseae">
|
||
<emphasis id="B91A3E69FFF8FFC4FE25FB64FD0C2BFC" box="[463,694,1187,1208]" italics="true" pageId="6" pageNumber="3012">Macellomenia morseae</emphasis>
|
||
</taxonomicName>
|
||
; Hs,
|
||
<taxonomicName id="4C6E99F8FFF8FFC4FD1EFB64FC502BFC" authorityName="Kocot & Todt" authorityYear="2014" box="[756,1002,1187,1208]" class="Solenogastres" family="Pruvotinidae" genus="Hypomenia" kingdom="Animalia" order="Cavibelonia" pageId="6" pageNumber="3012" phylum="Mollusca" rank="species" species="sanjuanensis">
|
||
<emphasis id="B91A3E69FFF8FFC4FD1EFB64FC502BFC" box="[756,1002,1187,1208]" italics="true" pageId="6" pageNumber="3012">Hypomenia sanjuanensis</emphasis>
|
||
</taxonomicName>
|
||
; note the similar size, colour, and body shape. Scale bar 1 mm; (B) posterior body end of
|
||
<taxonomicName id="4C6E99F8FFF8FFC4FC26FB07FBEA2B91" authorityName="Kocot & Todt" authorityYear="2014" box="[972,1104,1216,1237]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="6" pageNumber="3012" phylum="Mollusca" rank="species" species="schanderi">
|
||
<emphasis id="B91A3E69FFF8FFC4FC26FB07FBEA2B91" box="[972,1104,1216,1237]" italics="true" pageId="6" pageNumber="3012">M. schanderi</emphasis>
|
||
</taxonomicName>
|
||
showing: asp, abdominal spicules. Scale bar 150 µm; (C, D) epidermal sclerites of
|
||
<taxonomicName id="4C6E99F8FFF8FFC4FC0CFB1AFF712A4B" authority="Kocot & Todt, 2014" authorityName="Kocot & Todt" authorityYear="2014" class="Solenogastres" family="Pruvotinidae" genus="Hypomenia" kingdom="Animalia" order="Cavibelonia" pageId="6" pageNumber="3012" phylum="Mollusca" rank="species" species="sanjuanensis" status="sp. nov.">
|
||
<emphasis id="B91A3E69FFF8FFC4FC0CFB1AFF712A4B" italics="true" pageId="6" pageNumber="3012">H. sanjuanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2298312FFF8FFC4FF3CFB3DFE972A54" box="[214,301,1274,1296]" pageId="6" pageNumber="3012" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, light micrographs, polarized filter; (C) note the criss-cross arrangement of sclerites (cuticle removed with household bleach); scale bar 50 µm; (D) sclerite types: a, dorsolateral sclerites are acicular with small or almost reduced cavity, often serrated; b, most anterior sclerites are recurved and serrated; c, ventral scales; d, ventral scales found close to the pallial cavity opening; scale bar: 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C374B1F0FFF8FFC8FF7AFA1FFDB62E64" lastPageId="10" lastPageNumber="3016" pageId="6" pageNumber="3012" type="description">
|
||
<paragraph id="8BD1E27BFFF8FFC4FF7AFA1FFEB52AB4" blockId="6.[144,1104,1496,1687]" box="[144,271,1496,1520]" pageId="6" pageNumber="3012">
|
||
<heading id="D0995517FFF8FFC4FF7AFA1FFEB52AB4" box="[144,271,1496,1520]" fontSize="10" level="3" pageId="6" pageNumber="3012" reason="5">
|
||
<emphasis id="B91A3E69FFF8FFC4FF7AFA1FFEB52AB4" box="[144,271,1496,1520]" italics="true" pageId="6" pageNumber="3012">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFF8FFCAFF7AFA38FBA52A97" blockId="6.[144,1104,1496,1687]" lastBlockId="8.[144,1103,1435,1491]" lastPageId="8" lastPageNumber="3014" pageId="6" pageNumber="3012">
|
||
<emphasis id="B91A3E69FFF8FFC4FF7AFA38FF552953" box="[144,239,1535,1559]" italics="true" pageId="6" pageNumber="3012">Habitus.</emphasis>
|
||
Relaxed specimens are up to about
|
||
<quantity id="4C964F9EFFF8FFC4FD49FA38FD522953" box="[675,744,1535,1559]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="3012" unit="mm" value="2.0">2 mm</quantity>
|
||
in length by
|
||
<quantity id="4C964F9EFFF8FFC4FC66FA38FC5E2953" box="[908,996,1535,1559]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="3012" unit="mm" value="200.0">200 µm</quantity>
|
||
in width (
|
||
<figureCitation id="1355FEFEFFF8FFC4FF72F9D8FE912973" box="[152,299,1567,1591]" captionStart="Figure 3" captionStartId="7.[212,279,1306,1328]" captionTargetBox="[212,1172,168,1274]" captionTargetId="figure-1@7.[212,1172,168,1275]" captionTargetPageId="7" captionText="Figure 3. Macellomenia schanderi sp. nov. (A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="6" pageNumber="3012">Figure 3A, B</figureCitation>
|
||
). The body is without any keels or bumps. Specimens are creamy whitecoloured and slightly translucent in life with similar colour when fixed in formalin and preserved in 70% ethanol (although less translucent). Specimens have a shiny appearance (from the sclerites) when illuminated from above. Orange material was visible in the midgut of some specimens. Living animals maintained in glass dishes were observed to actively crawl around the dish, frequently raising their anterior end.
|
||
</paragraph>
|
||
<caption id="DF11B2F3FFF9FFC5FF3EFADDFEA12911" ID-DOI="http://doi.org/10.5281/zenodo.4607739" ID-Zenodo-Dep="4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="7" pageNumber="3013" startId="7.[212,279,1306,1328]" targetBox="[212,1172,168,1274]" targetPageId="7">
|
||
<paragraph id="8BD1E27BFFF9FFC5FF3EFADDFEA12911" blockId="7.[212,1173,1306,1621]" pageId="7" pageNumber="3013">
|
||
Figure 3.
|
||
<taxonomicName id="4C6E99F8FFF9FFC5FED4FADCFD8B2A74" authority="Kocot & Todt, 2014" authorityName="Kocot & Todt" authorityYear="2014" box="[318,561,1307,1328]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="7" pageNumber="3013" phylum="Mollusca" rank="species" species="schanderi" status="sp. nov.">
|
||
<emphasis id="B91A3E69FFF9FFC5FED4FADCFD8B2A74" box="[318,561,1307,1328]" italics="true" pageId="7" pageNumber="3013">Macellomenia schanderi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2298312FFF9FFC5FDD0FADDFD372A74" box="[570,653,1306,1328]" pageId="7" pageNumber="3013" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11B2F3FFF6FFCAFF7AFB44FEED2A22" ID-DOI="http://doi.org/10.5281/zenodo.4607741" ID-Zenodo-Dep="4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="8" pageNumber="3014" startId="8.[144,211,1155,1177]" targetBox="[147,1101,171,1121]" targetPageId="8">
|
||
<paragraph id="8BD1E27BFFF6FFCAFF7AFB44FEED2A22" blockId="8.[144,1104,1155,1382]" pageId="8" pageNumber="3014">
|
||
Figure 4. Light micrographs of histological cross sections through various body regions of
|
||
<taxonomicName id="4C6E99F8FFF6FFCAFF7AFB66FE392BF2" authorityName="Kocot & Todt" authorityYear="2014" box="[144,387,1185,1206]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="8" pageNumber="3014" phylum="Mollusca" rank="species" species="schanderi">
|
||
<emphasis id="B91A3E69FFF6FFCAFF7AFB66FE392BF2" box="[144,387,1185,1206]" italics="true" pageId="8" pageNumber="3014">Macellomenia schanderi</emphasis>
|
||
</taxonomicName>
|
||
. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD1E27BFFF6FFCBFF7AF9C3FC072E64" blockId="8.[144,1104,1539,1691]" lastBlockId="9.[212,1172,168,288]" lastPageId="9" lastPageNumber="3015" pageId="8" pageNumber="3014">
|
||
<emphasis id="B91A3E69FFF6FFCAFF7AF9C3FF522958" box="[144,232,1540,1564]" italics="true" pageId="8" pageNumber="3014">Mantle.</emphasis>
|
||
Sclerites are typical of the genus with an ovate, cup-shaped basal plate sharply tapering to a solid, slightly recurved and flattened acicular spine (
|
||
<figureCitation id="1355FEFEFFF6FFCAFF72F984FE8A291F" box="[152,304,1603,1627]" captionStart="Figure 3" captionStartId="7.[212,279,1306,1328]" captionTargetBox="[212,1172,168,1274]" captionTargetId="figure-1@7.[212,1172,168,1275]" captionTargetPageId="7" captionText="Figure 3. Macellomenia schanderi sp. nov. (A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="8" pageNumber="3014">Figure 3C, E</figureCitation>
|
||
). The acicular spines are widest medially rather than proximally and are covered in minute spinelets. The mantle sclerites are up to
|
||
<quantity id="4C964F9EFFF6FFCAFCB4F9A4FC1F293F" box="[862,933,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="3014" unit="mm" value="50.0">50 µm</quantity>
|
||
in total length with the basal plate being up to
|
||
<quantity id="4C964F9EFFF6FFCAFDEEF944FDF129DF" box="[516,587,1667,1691]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="3014" unit="mm" value="18.0">18 µm</quantity>
|
||
long and
|
||
<quantity id="4C964F9EFFF6FFCAFD2BF944FCB329DF" box="[705,777,1667,1691]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="3014" unit="mm" value="10.0">10 µm</quantity>
|
||
wide, and the acicular spine being up to about
|
||
<quantity id="4C964F9EFFF7FFCBFE4BFF6FFE602F84" box="[417,474,168,192]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="3015" unit="mm" value="6.0">6 µm</quantity>
|
||
in width. Only one
|
||
<typeStatus id="54D55CD9FFF7FFCBFD52FF6FFD522F84" box="[696,744,168,192]" pageId="9" pageNumber="3015">type</typeStatus>
|
||
of scale was observed to surround the foot and pedal pit (
|
||
<figureCitation id="1355FEFEFFF7FFCBFE54FF0FFDE72FA4" box="[446,605,200,224]" captionStart="Figure 3" captionStartId="7.[212,279,1306,1328]" captionTargetBox="[212,1172,168,1274]" captionTargetId="figure-1@7.[212,1172,168,1275]" captionTargetPageId="7" captionText="Figure 3. Macellomenia schanderi sp. nov. (A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="9" pageNumber="3015">Figure 3C, D</figureCitation>
|
||
). These peripedal sclerites are blade-shaped and distally pointed, measuring up to
|
||
<quantity id="4C964F9EFFF7FFCBFDBEFF2FFD212E44" box="[596,667,232,256]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="3015" unit="mm" value="30.0">30 µm</quantity>
|
||
in length. The cuticle is uniformly thin over the body with a maximum thickness of around
|
||
<quantity id="4C964F9EFFF7FFCBFD1EFECFFC812E64" box="[756,827,264,288]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="3015" unit="mm" value="27.0">27 µm</quantity>
|
||
(
|
||
<figureCitation id="1355FEFEFFF7FFCBFCA7FECFFC162E64" box="[845,940,264,288]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFF7FFCBFF3EFE8FFD672D3B" blockId="9.[212,1172,328,639]" pageId="9" pageNumber="3015">
|
||
<emphasis id="B91A3E69FFF7FFCBFF3EFE8FFDFF2E24" box="[212,581,328,352]" italics="true" pageId="9" pageNumber="3015">Pedal groove and mantle cavity.</emphasis>
|
||
The pedal pit is large and eversible (
|
||
<figureCitation id="1355FEFEFFF7FFCBFBE4FE8FFB3E2E24" box="[1038,1156,328,352]" captionStart="Figure 3" captionStartId="7.[212,279,1306,1328]" captionTargetBox="[212,1172,168,1274]" captionTargetId="figure-1@7.[212,1172,168,1275]" captionTargetPageId="7" captionText="Figure 3. Macellomenia schanderi sp. nov. (A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="9" pageNumber="3015">Figure 3B</figureCitation>
|
||
). Large, darkly staining pedal glands are associated with the pedal pit (
|
||
<figureCitation id="1355FEFEFFF7FFCBFC05FEAFFB3E2EC4" box="[1007,1156,360,384]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4B, C</figureCitation>
|
||
). Small sole glands similar in appearance to the pedal glands (but more darkly stained) are associated with the foot. The foot in fixed specimens consists of a single fold (
|
||
<figureCitation id="1355FEFEFFF7FFCBFEB0FE00FE552E9B" box="[346,495,455,479]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4B, C</figureCitation>
|
||
). The mantle cavity lacks respiratory papillae (
|
||
<figureCitation id="1355FEFEFFF7FFCBFBE4FE00FB3E2E9B" box="[1038,1156,455,479]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4D</figureCitation>
|
||
). No adhesive glands (such as that of
|
||
<taxonomicName id="4C6E99F8FFF7FFCBFD4CFE20FC982EBB" authorityName="M.P.Morse" authorityYear="1979" box="[678,802,487,511]" class="Solenogastres" family="Meiomeniidae" genus="Meiomenia" kingdom="Animalia" order="Pholidoskepia" pageId="9" pageNumber="3015" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91A3E69FFF7FFCBFD4CFE20FC982EBB" box="[678,802,487,511]" italics="true" pageId="9" pageNumber="3015">Meiomenia</emphasis>
|
||
</taxonomicName>
|
||
) were observed but there are glandular cells in the posterior mantle rim (not shown). The mantle cavity has a tubular ventral pouch, which at its muscular end holds a paired group of about 30– 40 abdominal spicules. This pouch is projectable and the whole organ could be called a genital cone (
|
||
<figureCitation id="1355FEFEFFF7FFCBFE3EFDA0FDF62D3B" box="[468,588,615,639]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4D</figureCitation>
|
||
; see below).
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFF7FFCBFF3EFD60FBFB2C99" blockId="9.[212,1172,678,989]" pageId="9" pageNumber="3015">
|
||
<emphasis id="B91A3E69FFF7FFCBFF3EFD60FE2C2DFB" box="[212,406,679,703]" italics="true" pageId="9" pageNumber="3015">Digestive system.</emphasis>
|
||
The mouth opening of
|
||
<taxonomicName id="4C6E99F8FFF7FFCBFD5EFD60FCF22DFB" authorityName="Kocot & Todt" authorityYear="2014" box="[692,840,679,703]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="9" pageNumber="3015" phylum="Mollusca" rank="species" species="schanderi">
|
||
<emphasis id="B91A3E69FFF7FFCBFD5EFD60FCF22DFB" box="[692,840,679,703]" italics="true" pageId="9" pageNumber="3015">M. schanderi</emphasis>
|
||
</taxonomicName>
|
||
is separate from the atrium (
|
||
<figureCitation id="1355FEFEFFF7FFCBFF36FD01FEEA2D9A" box="[220,336,710,734]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4A</figureCitation>
|
||
) but immediately posterior to it. There are no sclerites separating the two openings although there may be cuticle between them. The pharynx is narrow and muscular. It is about the same length as the distinct oesophagus (
|
||
<figureCitation id="1355FEFEFFF7FFCBFC2EFCC1FB8D2C5A" box="[964,1079,774,798]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4C</figureCitation>
|
||
), which has dorsally located oesophageal glands. There is a tiny monostichous radula (
|
||
<quantity id="4C964F9EFFF7FFCBFF31FC81FE832C1A" box="[219,313,838,862]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="3015" unit="mm" value="16.5">16.5 µm</quantity>
|
||
wide) that bears seven equally sized denticles per tooth (
|
||
<figureCitation id="1355FEFEFFF7FFCBFC24FC81FBFB2C1A" box="[974,1089,838,862]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4B</figureCitation>
|
||
). Each ventrolateral foregut gland consists of a long duct surrounded by densely packed subepithelial (extraepithelial) gland cells (
|
||
<typeStatus id="54D55CD9FFF7FFCBFD52FC41FD532CDA" box="[696,745,902,926]" pageId="9" pageNumber="3015">type</typeStatus>
|
||
A). The midgut has a short rostral caecum that is paired at the frontal end. No nematocysts were observed in the digestive system of either of the histologically sectioned specimens examined.
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFF7FFCBFF3EFBC1FE222BF9" blockId="9.[212,1172,1029,1213]" pageId="9" pageNumber="3015">
|
||
<emphasis id="B91A3E69FFF7FFCBFF3EFBC1FDDF2B5A" box="[212,613,1030,1054]" italics="true" pageId="9" pageNumber="3015">Nervous system and sensory organs.</emphasis>
|
||
The nervous system includes a cerebral ganglion that is large relative to the size of the animal. The cerebral ganglion spans 19 of the histological sections, making it about
|
||
<quantity id="4C964F9EFFF7FFCBFD7AFB82FD602B19" box="[656,730,1093,1117]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="3015" unit="mm" value="38.0">38 µm</quantity>
|
||
in length. The vestibular sense organ (atrium or vestibulum) holds about 20 simple sensory papillae. A large, distinct dorsoterminal sense organ is present (
|
||
<figureCitation id="1355FEFEFFF7FFCBFD60FB42FCB82BD9" box="[650,770,1157,1181]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4D</figureCitation>
|
||
). This organ was readily observed in living animals.
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFF7FFC8FF3EFB22FDB62E64" blockId="9.[212,1172,1253,1691]" lastBlockId="10.[144,1104,168,288]" lastPageId="10" lastPageNumber="3016" pageId="9" pageNumber="3015">
|
||
<emphasis id="B91A3E69FFF7FFCBFF3EFB22FE7B2BB9" box="[212,449,1253,1277]" italics="true" pageId="9" pageNumber="3015">Reproductive system.</emphasis>
|
||
In the region of the mantle cavity of
|
||
<taxonomicName id="4C6E99F8FFF7FFCBFC6DFB22FBA12BB9" authorityName="Kocot & Todt" authorityYear="2014" box="[903,1051,1253,1277]" class="Solenogastres" family="Macellomeniidae" genus="Macellomenia" kingdom="Animalia" order="Pholidoskepia" pageId="9" pageNumber="3015" phylum="Mollusca" rank="species" species="schanderi">
|
||
<emphasis id="B91A3E69FFF7FFCBFC6DFB22FBA12BB9" box="[903,1051,1253,1277]" italics="true" pageId="9" pageNumber="3015">M. schanderi</emphasis>
|
||
</taxonomicName>
|
||
there is a ventral muscular organ (
|
||
<figureCitation id="1355FEFEFFF7FFCBFDEBFAC3FDC42A58" box="[513,638,1284,1308]" captionStart="Figure 4" captionStartId="8.[144,211,1155,1177]" captionTargetBox="[147,1101,171,1121]" captionTargetId="figure-99@8.[144,1104,168,1124]" captionTargetPageId="8" captionText="Figure 4. Light micrographs of histological cross sections through various body regions of Macellomenia schanderi. Scale bars 100 µm. (A) Anterior body-end with atrium (a) showing atrial papillae; cu, cuticle; ep, epidermal papilla; (B) radular region with cerebral ganglion (cer), radula (r), and pedal pit (pp) with pedal glands (pgl); the insert shows the radula in higher magnification; (C) section slightly posterior to B with oesophagus (es), lateroventral foregut glands (fgl), and radular sheath (rs); cer, cerebral ganglion; ep, epidermal papilla; (D) posterior body-end with pallial cavity (pc), part of the dorsoterminal sense organ (dts), hind gut (hg) and genital cone (genc)." figureDoi="http://doi.org/10.5281/zenodo.4607741" httpUri="https://zenodo.org/record/4607741/files/figure.png" pageId="9" pageNumber="3015">Figure 4D</figureCitation>
|
||
) similar to the genital cone of
|
||
<taxonomicName id="4C6E99F8FFF7FFCBFBFAFAC2FE772A78" authority="(Salvini-Plawen 1968)" baseAuthorityName="Salvini-Plawen" baseAuthorityYear="1968" class="Solenogastres" family="Gymnomeniidae" genus="Genitoconia" kingdom="Animalia" order="Pholidoskepia" pageId="9" pageNumber="3015" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91A3E69FFF7FFCBFBFAFAC2FB2E2A59" box="[1040,1172,1285,1309]" italics="true" pageId="9" pageNumber="3015">Genitoconia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFF9F8AFFF7FFCBFF31FAE3FE782A78" author="Salvini-Plawen L" box="[219,450,1316,1340]" pageId="9" pageNumber="3015" pagination="231 - 272" refId="ref10302" refString="Salvini-Plawen L. 1968. Neue Formen im marinen Mesopsammon: Kamptozoa und Aculifera. Annalen des Naturhistorischen Museums in Wien. 72: 231 - 272." type="journal article" year="1968">Salvini-Plawen 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Within this structure are up to 40 posterior-pointing abdominal spicules. Examination of histological sections indicates that this structure is not covered by cuticle. When relaxed, most specimens seemed to partially evert the cone and abdominal spicules (
|
||
<figureCitation id="1355FEFEFFF7FFCBFE10FA43FDD52AD8" box="[506,623,1412,1436]" captionStart="Figure 2" captionStartId="6.[144,211,1157,1179]" captionTargetBox="[144,1104,168,1125]" captionTargetId="figure-75@6.[144,1104,168,1125]" captionTargetPageId="6" captionText="Figure 2. (A) Three new meiofaunal solenogaster species from Reid Rock, Friday Harbor: Ms, Macellomenia schanderi; Mm, Macellomenia morseae; Hs, Hypomenia sanjuanensis; note the similar size, colour, and body shape. Scale bar 1 mm; (B) posterior body end of M. schanderi showing: asp, abdominal spicules. Scale bar 150 µm; (C, D) epidermal sclerites of H. sanjuanensis sp. nov., light micrographs, polarized filter; (C) note the criss-cross arrangement of sclerites (cuticle removed with household bleach); scale bar 50 µm; (D) sclerite types: a, dorsolateral sclerites are acicular with small or almost reduced cavity, often serrated; b, most anterior sclerites are recurved and serrated; c, ventral scales; d, ventral scales found close to the pallial cavity opening; scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607737" httpUri="https://zenodo.org/record/4607737/files/figure.png" pageId="9" pageNumber="3015">Figure 2B</figureCitation>
|
||
). One specimen imaged by SEM had mucous covering up part of its posterior end including the base of the ‘genital cone’ but it appeared that this specimen had either completely ‘ejected’ the abdominal spicules or they had broken off near their point of attachment to the body and were stuck in this mucous (
|
||
<figureCitation id="1355FEFEFFF7FFCBFED4F9C3FE672958" box="[318,477,1540,1564]" captionStart="Figure 3" captionStartId="7.[212,279,1306,1328]" captionTargetBox="[212,1172,168,1274]" captionTargetId="figure-1@7.[212,1172,168,1275]" captionTargetPageId="7" captionText="Figure 3. Macellomenia schanderi sp. nov. (A) SEM micrograph of entire specimen, oblique ventral view. Position of anterior end (ant) and pedal groove (pg) indicated. Scale bar 200 µm; (B) ventral view of anterior body tip showing the atrium (a), mouth (mo) and the partially everted, ciliated pedal pit (pp). Scale bar 50 µm; (C) SEM micrograph of epidermal sclerites. The small insert shows a pedal scale (ps), the larger image illustrates the morphology of the main sclerite type with broadened base (bs) and flattened, recurved spine (s) covered in minute spinelets. Scale bar 10 µm; (D) SEM micrograph of the partially everted genital cone and pallial cavity opening (pc) with abdominal spines (asp) covered in mucus (muc). Scale bar 50 µm; (E) light micrograph of sclerites. Most of the body sclerites are intact although the acicular portion has been separated from the basal plate from several. There is one pedal scale (ps). Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4607739" httpUri="https://zenodo.org/record/4607739/files/figure.png" pageId="9" pageNumber="3015">Figure 3A, D</figureCitation>
|
||
). The unpaired portion of the spawning duct is very large, filling the ventral half of the posterior portion of the body. Only close to where the spawning duct fuses with the pericardioducts, the lumen of the spawning duct becomes paired. There are darkly stained vesicles in the cells of the spawning duct. The frontal end of the spawning duct is bilobed. One pair of tube-shaped seminal receptacles is attached to the spawning duct, just laterofrontal to where the pericardioducts fuse with its frontal end. The pericardioducts originate at the posterior end of the pericardium. The gonad is distinctly paired, with each half up against the body wall on either side of the midgut.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374B1F0FFF4FFC8FF7AFEA5FE432EE5" box="[144,505,354,417]" pageId="10" pageNumber="3016" type="distribution">
|
||
<paragraph id="8BD1E27BFFF4FFC8FF7AFEA5FEAE2E3E" blockId="10.[144,505,354,417]" box="[144,276,354,378]" pageId="10" pageNumber="3016">
|
||
<heading id="D0995517FFF4FFC8FF7AFEA5FEAE2E3E" box="[144,276,354,378]" fontSize="10" level="3" pageId="10" pageNumber="3016" reason="5">
|
||
<emphasis id="B91A3E69FFF4FFC8FF7AFEA5FEAE2E3E" box="[144,276,354,378]" italics="true" pageId="10" pageNumber="3016">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD1E27BFFF4FFC8FF7AFE4EFE432EE5" blockId="10.[144,505,354,417]" box="[144,505,393,417]" pageId="10" pageNumber="3016">
|
||
Only known from
|
||
<typeStatus id="54D55CD9FFF4FFC8FE8FFE4EFE2F2EE5" box="[357,405,393,417]" pageId="10" pageNumber="3016">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374B1F0FFF4FFC8FF7AFE24FB9A2D06" pageId="10" pageNumber="3016" type="etymology">
|
||
<paragraph id="8BD1E27BFFF4FFC8FF7AFE24FEB32EBF" blockId="10.[144,1104,483,578]" box="[144,265,483,507]" pageId="10" pageNumber="3016">
|
||
<heading id="D0995517FFF4FFC8FF7AFE24FEB32EBF" box="[144,265,483,507]" fontSize="10" level="3" pageId="10" pageNumber="3016" reason="5">
|
||
<emphasis id="B91A3E69FFF4FFC8FF7AFE24FEB32EBF" box="[144,265,483,507]" italics="true" pageId="10" pageNumber="3016">Etymology</emphasis>
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||
</heading>
|
||
</paragraph>
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||
<paragraph id="8BD1E27BFFF4FFC8FF7AFDCCFB9A2D06" blockId="10.[144,1104,483,578]" pageId="10" pageNumber="3016">This species is named to honour the late Dr Christoffer Schander, an expert on aplacophorans and other molluscs who was a mentor and friend to the authors.</paragraph>
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</subSubSection>
|
||
</treatment>
|
||
</document> |