treatments-xml/data/03/C4/87/03C487D0C45BFFB0174FF957260FA0E4.xml
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<document id="DB72401B4050BB389EC19BDE562111DA" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105362570" checkinUser="felipe" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel-Angel" docDate="2005" docId="03C487D0C45BFFB0174FF957260FA0E4" docLanguage="en" docName="zt01012p037.pdf" docOrigin="Zootaxa 1012 (1012)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrodactylus tibialis Arce-Perez and Moron 2005, new species" docType="treatment" docVersion="3" lastPageNumber="31" masterDocId="FFFDFFA8C45DFFB81647FFEA2418A64A" masterDocTitle="New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua" masterLastPageNumber="37" masterPageNumber="23" pageNumber="29" updateTime="1698980141271" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="664270B3C6B55E053C394F8A97EDBE7F">New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua</mods:title>
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<mods:namePart id="8791924FFB0DBD52BDF45C294B0292ED">Arce-Pérez, Roberto</mods:namePart>
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<mods:date id="F52C4D7EF9DAF8CA4DA6ACF22BC37E7D">2005</mods:date>
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<mods:number id="2FF0C8486BDD05CB428DC74254FF4F53">2005-06-28</mods:number>
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<treatment id="03C487D0C45BFFB0174FF957260FA0E4" ID-DOI="http://doi.org/10.5281/zenodo.5049194" ID-Zenodo-Dep="5049194" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4" httpUri="http://treatment.plazi.org/id/03C487D0C45BFFB0174FF957260FA0E4" lastPageId="8" lastPageNumber="31" pageId="6" pageNumber="29">
<subSubSection id="C377654DC45BFFBE174FF95727C1A09D" box="[264,985,1725,1751]" pageId="6" pageNumber="29" type="nomenclature">
<paragraph id="8BD236C6C45BFFBE174FF95727C1A09D" blockId="6.[264,985,1725,1786]" box="[264,985,1725,1751]" pageId="6" pageNumber="29">
<heading id="D09A81AAC45BFFBE174FF95727C1A09D" bold="true" box="[264,985,1725,1751]" fontSize="11" level="1" pageId="6" pageNumber="29" reason="1">
<emphasis id="B919EAD4C45BFFBE174FF95727C1A09D" bold="true" box="[264,985,1725,1751]" pageId="6" pageNumber="29">
<taxonomicName id="4C6D4D45C45BFFBE174FF9572724A09D" authority="Arce-Perez and Moron" authorityName="Arce-Perez and Moron" authorityYear="2005" box="[264,828,1725,1751]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B919EAD4C45BFFBE174FF957260BA09D" bold="true" box="[264,531,1725,1751]" italics="true" pageId="6" pageNumber="29">Macrodactylus tibialis</emphasis>
Arce­Pérez and Morón
</taxonomicName>
,
<taxonomicNameLabel id="A22A57AFC45BFFBE150DF95727C1A09D" box="[842,985,1725,1751]" pageId="6" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45BFFBE174FF90A25F0A0B0" box="[264,488,1760,1786]" pageId="6" pageNumber="29" type="description">
<paragraph id="8BD236C6C45BFFBE174FF90A25F0A0B0" blockId="6.[264,985,1725,1786]" box="[264,488,1760,1786]" pageId="6" pageNumber="29">
(
<figureCitation id="13562A43C45BFFBE1756F90A2590A0B0" box="[273,392,1760,1786]" captionStart="FIGURES 712" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 712. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="6" pageNumber="29">Figs. 7,10</figureCitation>
,
<figureCitation id="13562A43C45BFFBE17D2F90A25C6A0B0" box="[405,478,1760,1786]" captionStart="FIGURES 1321" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 1321. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="6" pageNumber="29">1315</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45BFFBF174FF8C5204AA714" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29" type="diagnosis">
<paragraph id="8BD236C6C45BFFBF174FF8C5204AA714" blockId="6.[264,1323,1839,1906]" lastBlockId="7.[264,1324,284,1870]" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">
<emphasis id="B919EAD4C45BFFBE174FF8C52590A103" bold="true" box="[264,392,1839,1865]" pageId="6" pageNumber="29">Diagnosis.</emphasis>
This species is distinguished by the following combination of characters: antennal club unicolored; pronotum, elytra, and pygidium glabrous, orange; tarsi without rings of white setae; male with a strong, acute spine near the middle of the inner border of each protibia; parameres are stout without setae on the external borders.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45AFFBF177FFE81272CA104" pageId="7" pageNumber="30" type="description">
<paragraph id="8BD236C6C45AFFBF177FFE8127DEA3F4" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="30">
<emphasis id="B919EAD4C45AFFBF177FFE812683A7CF" bold="true" box="[312,667,363,389]" pageId="7" pageNumber="30">
Description.
<typeStatus id="54D68864C45AFFBF179DFE812656A7CF" box="[474,590,363,389]" pageId="7" pageNumber="30" type="holotype">Holotype</typeStatus>
. Male
</emphasis>
: Body length
<quantity id="4C959B23C45AFFBF150FFE8627AEA7CC" box="[840,950,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="30" unit="mm" value="12.5">12.5 mm</quantity>
; humeral width
<quantity id="4C959B23C45AFFBF123DFE8620C1A7CC" box="[1146,1241,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="30" unit="mm" value="3.5">3.5 mm</quantity>
.
<emphasis id="B919EAD4C45AFFBF12A1FE862132A7CF" box="[1254,1322,364,389]" italics="true" pageId="7" pageNumber="30">Color</emphasis>
(
<figureCitation id="13562A43C45AFFBF1756FE7E2541A7E4" box="[273,345,404,430]" captionStart="FIGURES 712" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 712. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="7" pageNumber="30">Fig. 7</figureCitation>
): clypeus, antennae, mouthparts, pronotum, elytra, pygidium, last sternite, femora, and tibiae orange; frons, scutellum, pterothorax, abdomen, and tarsi black.
<emphasis id="B919EAD4C45AFFBF12C9FE5620FFA79F" box="[1166,1255,444,469]" italics="true" pageId="7" pageNumber="30">Surface</emphasis>
: dorsally glabrous with iridescent silky shine when viewed without magnification, coarsely punctate or finely rugose with magnification; ventrally with short, fine, yellow setae.
<emphasis id="B919EAD4C45AFFBF174FFDDE2549A407" box="[264,337,564,589]" italics="true" pageId="7" pageNumber="30">Head:</emphasis>
clypeus trapezoidal, anterior border nearly straight; antennal club yellow; mentum elongate, longitudinally furrowed, anterior border straight, anterolateral emargination exposing basal segment of labial palpus.
<emphasis id="B919EAD4C45AFFBF14B8FD6E274BA4D7" box="[767,851,644,669]" italics="true" pageId="7" pageNumber="30">Thorax</emphasis>
: pronotum hexagonal, convex,
<quantity id="4C959B23C45AFFBF1297FD6E2134A4D4" box="[1232,1324,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="30" unit="mm" value="3.1">3.1 mm</quantity>
length,
<quantity id="4C959B23C45AFFBF1725FD4625A6A48C" box="[354,446,684,710]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="30" unit="mm" value="2.8">2.8 mm</quantity>
width; prosternal process small, subtriangular, slightly depressed, hidden in frontal view; scutellum elongate with rounded apex; elytra with punctate striae. Metasternum with two longitudinal rows of short, red setae between meso­ and metacoxae. Procoxa conical, prominent. Protibia with two rounded teeth at the apex of external border and one strong, acute spine near the middle of inner border (
<figureCitation id="13562A43C45AFFBF1598FCA6203EA52C" box="[991,1062,844,870]" captionStart="FIGURES 16" captionStartId="6.[264,383,1501,1525]" captionTargetBox="[275,1312,313,1474]" captionTargetId="figure-83@6.[275,1312,309,1474]" captionTargetPageId="6" captionText="FIGURES 16. 1­­­ Macrodactylus costulatus Bates, male; 2­­­ M. sericeicollis Bates, male; 3­­­ M. sericinus Bates, male; 4­­­ M. suavis Bates, male; 5­­­ M. dimidiatus Guérin­Méneville, male; 6­ ­­ M. sylphis Bates, male." pageId="7" pageNumber="30">Fig. 1</figureCitation>
). Protibial articulated spur absent. Protarsus shortened, with spine­like setae at the apex of each segment, without rings of white setae. Meso­ and metafemur with short, spine­like setae scattered on ventral surface. Mesotibia widened toward the apex with two long, narrow, straight, acute spurs and long spine­like setae around the apex. Metatibia widened toward the apex with some preapical long, black setae on the inner surface, and long, spine­like setae around the apex. Meso­ and metatarsi enlarged with short, spine­like setae on the surface and long spine­like setae around the apex of each segment, without rings of white setae. All tarsal claws narrowly cleft.
<emphasis id="B919EAD4C45AFFBF1458FB662695A2EF" box="[543,653,1164,1189]" italics="true" pageId="7" pageNumber="30">Abdomen</emphasis>
: sternites 24 moderately concave at middle with two spine­like, erect, reddish, short setae at the sides of midline; length of sternite 5 twice the length of sternite 4 with numerous spine­like, erect, long setae near the posterior border; sternite 6 ovate, with numerous spine­like, red setae on the surface and long, black setae on the apical border. Pygidium elongate, strongly convex, with scattered spine­like setae.
<emphasis id="B919EAD4C45AFFBF174FFABE256FA327" box="[264,375,1364,1389]" italics="true" pageId="7" pageNumber="30">Genitalia</emphasis>
: total length
<quantity id="4C959B23C45AFFBF1456FABE2664A324" box="[529,636,1364,1390]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" pageId="7" pageNumber="30" unit="mm" value="3.75">3.75 mm</quantity>
; parameres stout, glabrous, completely fused dorsobasally, widely curved toward the apex, forming an oval in distal view with
<quantity id="4C959B23C45AFFBF127DFA9620BDA3DC" box="[1082,1189,1404,1430]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="30" unit="mm" value="1.5">1.50 mm</quantity>
length and
<quantity id="4C959B23C45AFFBF174FFA4E2568A3F4" box="[264,368,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="30" unit="mm" value="1.24">1.24 mm</quantity>
width; each apex weakly lanceolate (
<figureCitation id="13562A43C45AFFBF156EFA4E27A0A3F4" box="[809,952,1444,1470]" captionStart="FIGURES 1321" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 1321. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="7" pageNumber="30">Figs. 1314</figureCitation>
).
</paragraph>
<paragraph id="8BD236C6C45AFFBF177FFA26202EA0CC" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="30">
<emphasis id="B919EAD4C45AFFBF177FFA262613A3AC" bold="true" box="[312,523,1484,1510]" pageId="7" pageNumber="30">
<typeStatus id="54D68864C45AFFBF177FFA2625BCA3AC" box="[312,420,1484,1510]" pageId="7" pageNumber="30" type="allotype">Allotype</typeStatus>
. Female
</emphasis>
: Body length 12.0 mm; humeral width
<quantity id="4C959B23C45AFFBF1599FA262020A3AC" box="[990,1080,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="30" unit="mm" value="3.9">3.9 mm</quantity>
(
<figureCitation id="13562A43C45AFFBF120EFA2620B8A3AC" box="[1097,1184,1484,1510]" captionStart="FIGURES 712" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 712. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="7" pageNumber="30">Fig. 10</figureCitation>
). Similar to male except as follows: pronotum
<quantity id="4C959B23C45AFFBF14D9FA1E271DA044" box="[670,773,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="7" pageNumber="30" unit="mm" value="3.05">3.05 mm</quantity>
length,
<quantity id="4C959B23C45AFFBF1521FA1E27D7A044" box="[870,975,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="30" unit="mm" value="3.1">3.10 mm</quantity>
width; protibia without acute spine near the middle of inner border with preapical articulated spur; abdominal sternites 24 convex at middle without erect, long setae; pygidium short, weakly convex, nearly triangular. Genital plates large, external border widely curved (
<figureCitation id="13562A43C45AFFBF1596F986203FA0CC" box="[977,1063,1644,1670]" captionStart="FIGURES 1321" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 1321. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="7" pageNumber="30">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BD236C6C45AFFBF177FF97E272CA104" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="30">
<emphasis id="B919EAD4C45AFFBF177FF97E25B6A0E4" bold="true" box="[312,430,1684,1710]" pageId="7" pageNumber="30">Variation</emphasis>
(
<specimenCount id="9D6BFD4FC45AFFBF1787F97E2674A0E4" box="[448,620,1684,1710]" pageId="7" pageNumber="30" type="generic" typeStatus="paratypes">131 paratypes</specimenCount>
). Body length 12.0
<quantity id="4C959B23C45AFFBF1526F97E27D6A0E4" box="[865,974,1684,1710]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="30" unit="mm" value="13.5">13.5 mm</quantity>
; humeral width 3.0
<quantity id="4C959B23C45AFFBF1280F97E213EA0E4" box="[1223,1318,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="30" unit="mm" value="3.6">3.6 mm</quantity>
; prosternal process slightly elongate in some specimens; pronotum completely dark green or orange with a dark green stripe along the midline or a dark brown spot at the middle of pronotal disc in some specimens from
<collectingCountry id="F37A7656C45AFFBF14A5F8E62746A16C" box="[738,862,1804,1830]" name="Nicaragua" pageId="7" pageNumber="30">Nicaragua</collectingCountry>
. Posterior half of the elytra iridescent dark brown in some specimens from
<collectingCountry id="F37A7656C45AFFBF14FDF8DE2728A104" box="[698,816,1844,1870]" name="Honduras" pageId="7" pageNumber="30">Honduras</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC455FFB0177FFEF120D6A21C" pageId="8" pageNumber="31" type="materials_examined">
<paragraph id="8BD236C6C455FFB0177FFEF120D6A21C" blockId="8.[264,1323,283,1710]" pageId="8" pageNumber="31">
<emphasis id="B919EAD4C455FFB0177FFEF1263DA77F" bold="true" box="[312,549,283,309]" pageId="8" pageNumber="31">Material examined</emphasis>
(
<specimenCount id="9D6BFD4FC455FFB01471FEF626BCA77C" box="[566,676,284,310]" pageId="8" pageNumber="31" type="male">79 males</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB014F6FEF6272FA77C" box="[689,823,284,310]" pageId="8" pageNumber="31" type="female">57 females</specimenCount>
).
<materialsCitation id="3B053C9BC455FFB0150BFEF625B3A7CC" collectingDate="1984-07-01" collectionCode="IEXA" collectorName="Col. J. M. Maes" country="Nicaragua" elevation="1000" location="Matagalpa-Jinotega" municipality="Km" pageId="8" pageNumber="31" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D68864C455FFB0150BFEF627A1A77C" box="[844,953,284,310]" pageId="8" pageNumber="31" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F37A7656C455FFB0124DFEF620A6A77C" box="[1034,1214,284,310]" name="Nicaragua" pageId="8" pageNumber="31">NICARAGUA</collectingCountry>
.
<collectingMunicipality id="6BB6ACBCC455FFB0128DFEF620EEA77C" box="[1226,1270,284,310]" pageId="8" pageNumber="31">Km</collectingMunicipality>
137 carretera
<location id="8EB2601DC455FFB0173FFEAE267AA714" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB0173FFEAE267AA714" box="[376,610,324,350]" country="Nicaragua" municipality="Km" name="Matagalpa-Jinotega" pageId="8" pageNumber="31">Matagalpa­Jinotega</location>
,
<quantity id="4C959B23C455FFB01437FEAE26D5A714" box="[624,717,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="31" unit="m" value="1000.0">
<elevation id="0040D1F5C455FFB01437FEAE26D5A714" box="[624,717,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="31" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB0149BFEAE2774A714" box="[732,876,324,350]" pageId="8" pageNumber="31" value="1984-07-01">
<collectingDate id="EF97E9EEC455FFB0149BFEAE2774A714" box="[732,876,324,350]" pageId="8" pageNumber="31" value="1984-07-01">1 July 1984</collectingDate>
</date>
, on weeds along the road,
<collectorName id="26985310C455FFB012F6FEAE255FA7CC" pageId="8" pageNumber="31">Col. J. M. Maes</collectorName>
(
<collectionCode id="ED7CAE03C455FFB01710FE8625BBA7CC" box="[343,419,364,390]" pageId="8" pageNumber="31">IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B053C9BC455FFB017F1FE862030A7E4" collectingDate="1995-07-09" country="Nicaragua" elevation="1500" location="Fuente Pura" pageId="8" pageNumber="31" specimenCount="1" stateProvince="Matagalpa" typeStatus="allotype">
<typeStatus id="54D68864C455FFB017F1FE862604A7CC" box="[438,540,364,390]" pageId="8" pageNumber="31" type="allotype">Allotype</typeStatus>
female:
<collectingCountry id="F37A7656C455FFB014C5FE86272EA7CC" box="[642,822,364,390]" name="Nicaragua" pageId="8" pageNumber="31">NICARAGUA</collectingCountry>
.
<collectingRegion id="49A9F824C455FFB01507FE862052A7CC" box="[832,1098,364,390]" country="Nicaragua" name="Matagalpa" pageId="8" pageNumber="31">Matagalpa department</collectingRegion>
,
<location id="8EB2601DC455FFB01212FE8620FCA7CC" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB01212FE8620FCA7CC" box="[1109,1252,364,390]" country="Nicaragua" name="Fuente Pura" pageId="8" pageNumber="31" stateProvince="Matagalpa">Fuente Pura</location>
,
<quantity id="4C959B23C455FFB012A8FE862539A7E4" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="31" unit="m" value="1500.0">
<elevation id="0040D1F5C455FFB012A8FE862539A7E4" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="31" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB01769FE7E25A2A7E4" box="[302,442,404,430]" pageId="8" pageNumber="31" value="1995-07-09">
<collectingDate id="EF97E9EEC455FFB01769FE7E25A2A7E4" box="[302,442,404,430]" pageId="8" pageNumber="31" value="1995-07-09">9 July 1995</collectingDate>
</date>
,
<collectedFrom id="3871753BC455FFB01781FE7E2030A7E4" box="[454,1064,404,430]" pageId="8" pageNumber="31">
on
<taxonomicName id="4C6D4D45C455FFB017ABFE7E2030A7E4" authority=", Cols. J. M. Maes &amp; P. Jolivet (IEXA)" box="[492,1064,404,430]" class="Liliopsida" family="Poaceae" genus="Zea" kingdom="Plantae" order="Poales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="species" species="mays">
<emphasis id="B919EAD4C455FFB017ABFE7E2644A7E7" box="[492,604,404,429]" italics="true" pageId="8" pageNumber="31">Zea mays</emphasis>
, Cols. J. M. Maes &amp; P. Jolivet (IEXA)
</taxonomicName>
</collectedFrom>
</materialsCitation>
.
<materialsCitation id="3B053C9BC455FFB01273FE7E269EA79C" collectingDate="1984-07-01" collectionCode="MXAL" collectorName="Col. J. M. Maes" country="Nicaragua" elevation="1000" location="Matagalpa-Jinotega" municipality="Km" pageId="8" pageNumber="31" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54D68864C455FFB01273FE7E20B2A7E4" box="[1076,1194,404,430]" pageId="8" pageNumber="31" type="paratype">Paratypes</typeStatus>
: same data as holotype (
<specimenCount id="9D6BFD4FC455FFB017E9FE56261BA79C" box="[430,515,444,470]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01466FE562666A79C" box="[545,638,444,470]" pageId="8" pageNumber="31">MXAL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB014DFFE562085A79C" box="[664,1181,444,470]" collectingDate="1995-07-09" collectionCode="MELN" country="Nicaragua" elevation="1500" location="Fuente Pura" pageId="8" pageNumber="31" specimenCount="2" specimenCount-male="2" stateProvince="Matagalpa" typeStatus="paratype">
same data as allotype (
<specimenCount id="9D6BFD4FC455FFB015FCFE562006A79C" box="[955,1054,444,470]" pageId="8" pageNumber="31" type="male">2 males</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB0127BFE56208EA79C" box="[1084,1174,444,470]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB012F7FE5625CDA46C" collectingDate="1994-06-12" collectionCode="MXAL, MELN" collectorName="Cols. &amp; J, M. Maes, J &amp; Tellez &amp; Hernandez" country="Nicaragua" elevation="1500" location="Fuente Pura" pageId="8" pageNumber="31" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Matagalpa" typeStatus="paratype">
same data except:
<date id="FFD31006C455FFB01721FE0E2612A7B4" box="[358,522,484,510]" pageId="8" pageNumber="31" value="1994-06-12">
<collectingDate id="EF97E9EEC455FFB01721FE0E2612A7B4" box="[358,522,484,510]" pageId="8" pageNumber="31" value="1994-06-12">12 June 1994</collectingDate>
</date>
,
<collectorName id="26985310C455FFB0145EFE0E2710A7B4" box="[537,776,484,510]" pageId="8" pageNumber="31">Cols. J, M. Maes, J</collectorName>
.
<collectorName id="26985310C455FFB01551FE0E2747A7B4" box="[790,863,484,510]" pageId="8" pageNumber="31">Téllez</collectorName>
&amp; J.
<collectorName id="26985310C455FFB015E0FE0E203EA7B4" box="[935,1062,484,510]" pageId="8" pageNumber="31">Hernández</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB0127CFE0E2095A7B4" box="[1083,1165,484,510]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB012E0FE0E2119A7B4" box="[1191,1281,484,510]" pageId="8" pageNumber="31">MXAL</collectionCode>
) (
<specimenCount id="9D6BFD4FC455FFB01359FE0E2542A46C" pageId="8" pageNumber="31" type="female">1 female</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01734FDE625D5A46C" box="[371,461,524,550]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<date id="FFD31006C455FFB017A3FDE6269EA46C" box="[484,646,524,550]" pageId="8" pageNumber="31" value="1994-06-26">26 June 1994</date>
(
<specimenCount id="9D6BFD4FC455FFB014DCFDE626F6A46C" box="[667,750,524,550]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB014BBFDE62771A46C" box="[764,873,524,550]" pageId="8" pageNumber="31" type="female">1 female</specimenCount>
);
<materialsCitation id="3B053C9BC455FFB015C7FDE6267FA404" collectingDate="1994-07-06" collectionCode="MELN" collectorName="Cols. &amp; Maes &amp; Hernandez" country="Nicaragua" pageId="8" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Matagalpa" typeStatus="paratype">
<date id="FFD31006C455FFB015C7FDE62008A46C" box="[896,1040,524,550]" pageId="8" pageNumber="31" value="1994-07-06">
<collectingDate id="EF97E9EEC455FFB015C7FDE62008A46C" box="[896,1040,524,550]" pageId="8" pageNumber="31" value="1994-07-06">6 July 1994</collectingDate>
</date>
,
<collectorName id="26985310C455FFB01258FDE62044A46C" box="[1055,1116,524,550]" pageId="8" pageNumber="31">Cols.</collectorName>
J. M.
<collectorName id="26985310C455FFB012E9FDE620F5A46C" box="[1198,1261,524,550]" pageId="8" pageNumber="31">Maes</collectorName>
&amp; J.
<collectorName id="26985310C455FFB0174FFDDE259FA404" box="[264,391,564,590]" pageId="8" pageNumber="31">Hernández</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB017DCFDDE25F3A404" box="[411,491,564,590]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01442FDDE2647A404" box="[517,607,564,590]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB01432FDDE213EA43C" collectingDate="1995-05-08" collectionCode="MELN" collectorName="Cols. &amp; Maes &amp; J. Tellez &amp; J. Hernandez" country="Nicaragua" elevation="1300" location="Selva Negra" pageId="8" pageNumber="31" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Matagalpa" typeStatus="paratype">
<collectingCountry id="F37A7656C455FFB01432FDDE2731A404" box="[629,809,564,590]" name="Nicaragua" pageId="8" pageNumber="31">NICARAGUA</collectingCountry>
.
<collectingRegion id="49A9F824C455FFB01572FDDE2059A404" box="[821,1089,564,590]" country="Nicaragua" name="Matagalpa" pageId="8" pageNumber="31">Matagalpa department</collectingRegion>
,
<location id="8EB2601DC455FFB01209FDDE20FAA404" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB01209FDDE20FAA404" box="[1102,1250,564,590]" country="Nicaragua" name="Selva Negra" pageId="8" pageNumber="31" stateProvince="Matagalpa">Selva Negra</location>
,
<quantity id="4C959B23C455FFB012B7FDDE2539A43C" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="31" unit="m" value="1300.0">
<elevation id="0040D1F5C455FFB012B7FDDE2539A43C" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="31" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB01769FDB625A7A43C" box="[302,447,604,630]" pageId="8" pageNumber="31" value="1995-05-08">
<collectingDate id="EF97E9EEC455FFB01769FDB625A7A43C" box="[302,447,604,630]" pageId="8" pageNumber="31" value="1995-05-08">8 May 1995</collectingDate>
</date>
,
<collectorName id="26985310C455FFB0178BFDB62611A43C" box="[460,521,604,630]" pageId="8" pageNumber="31">Cols.</collectorName>
J. M.
<collectorName id="26985310C455FFB01412FDB6268FA43C" box="[597,663,604,630]" pageId="8" pageNumber="31">Maes</collectorName>
,
<collectorName id="26985310C455FFB014E3FDB62710A43C" box="[676,776,604,630]" pageId="8" pageNumber="31">J. Téllez</collectorName>
&amp;
<collectorName id="26985310C455FFB01569FDB627D0A43C" box="[814,968,604,630]" pageId="8" pageNumber="31">J. Hernández</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB0159CFDB62034A43C" box="[987,1068,604,630]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB0127FFDB620B4A43C" box="[1080,1196,604,630]" pageId="8" pageNumber="31" type="female">7 females</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01283FDB62106A43C" box="[1220,1310,604,630]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB0174FFD6E25DBA48C" collectingDate="2004-06-25" collectingDateMax="2004-06-29" collectingDateMin="2004-06-25" collectionCode="PSPC" collectorName="Col. P. Schoolmeesters" country="Nicaragua" location="Selva Negra" pageId="8" pageNumber="31" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Matagalpa" typeStatus="paratype">
<collectingRegion id="49A9F824C455FFB0174FFD6E259EA4D4" box="[264,390,644,670]" country="Nicaragua" name="Matagalpa" pageId="8" pageNumber="31">Matagalpa</collectingRegion>
,
<location id="8EB2601DC455FFB017D0FD6E2634A4D4" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB017D0FD6E2634A4D4" box="[407,556,644,670]" country="Nicaragua" name="Selva Negra" pageId="8" pageNumber="31" stateProvince="Matagalpa">Selva Negra</location>
(hotel)
<date id="FFD31006C455FFB014D4FD6E2770A4D4" box="[659,872,644,670]" pageId="8" pageNumber="31" value="2004-06-25" valueMax="2004-06-29" valueMin="2004-06-25">
<collectingDate id="EF97E9EEC455FFB014D4FD6E2770A4D4" box="[659,872,644,670]" pageId="8" pageNumber="31" value="2004-06-25" valueMax="2004-06-29" valueMin="2004-06-25">2529 June 2004</collectingDate>
</date>
,
<collectorName id="26985310C455FFB0153DFD6E2088A4D4" box="[890,1168,644,670]" pageId="8" pageNumber="31">Col. P. Schoolmeesters</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB012E1FD6E2111A4D4" box="[1190,1289,644,670]" pageId="8" pageNumber="31" type="male">2 males</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB0135DFD6E2542A48C" pageId="8" pageNumber="31" type="female">1 female</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01734FD4625A3A48C" box="[371,443,684,710]" pageId="8" pageNumber="31">PSPC</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B053C9BC455FFB0178AFD46277CA4A4" collectingDate="2000-07-29" collectionCode="MELN" collectorName="Cols. &amp; Maes &amp; Tellez" country="Nicaragua" elevation="1370" location="San Lucas" municipality="Somoto" pageId="8" pageNumber="31" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Madriz" typeStatus="paratype">
<collectingRegion id="49A9F824C455FFB0178AFD4626AAA48C" box="[461,690,684,710]" country="Nicaragua" name="Madriz" pageId="8" pageNumber="31">Madriz department</collectingRegion>
,
<collectingMunicipality id="6BB6ACBCC455FFB01487FD462706A48C" box="[704,798,684,710]" pageId="8" pageNumber="31">Somoto</collectingMunicipality>
,
<location id="8EB2601DC455FFB0156BFD4627B2A48C" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB0156BFD4627B2A48C" box="[812,938,684,710]" country="Nicaragua" municipality="Somoto" name="San Lucas" pageId="8" pageNumber="31" stateProvince="Madriz">San Lucas</location>
,
<quantity id="4C959B23C455FFB015FFFD46200EA48C" box="[952,1046,684,710]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="8" pageNumber="31" unit="m" value="1370.0">
<elevation id="0040D1F5C455FFB015FFFD46200EA48C" box="[952,1046,684,710]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="8" pageNumber="31" unit="m" value="1370.0">1370 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB01263FD4620DBA48C" box="[1060,1219,684,710]" pageId="8" pageNumber="31" value="2000-07-29">
<collectingDate id="EF97E9EEC455FFB01263FD4620DBA48C" box="[1060,1219,684,710]" pageId="8" pageNumber="31" value="2000-07-29">29 July 2000</collectingDate>
</date>
,
<collectorName id="26985310C455FFB01296FD462116A48C" box="[1233,1294,684,710]" pageId="8" pageNumber="31">Cols.</collectorName>
J. M.
<collectorName id="26985310C455FFB01774FD3E256AA4A4" box="[307,370,724,750]" pageId="8" pageNumber="31">Maes</collectorName>
&amp; B.
<collectorName id="26985310C455FFB01787FD3E2611A4A4" box="[448,521,724,750]" pageId="8" pageNumber="31">Téllez</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01458FD3E2669A4A4" box="[543,625,724,750]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB01438FD3E26F1A4A4" box="[639,745,724,750]" pageId="8" pageNumber="31" type="female">1 female</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01545FD3E2744A4A4" box="[770,860,724,750]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB01535FD3E256AA574" collectingDate="1998-05-17" collectionCode="MELN" collectorName="Cols. J. M. Maes &amp; B. Hernandez" country="Nicaragua" location="Camp." municipality="Cerro Kilambe" pageId="8" pageNumber="31" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Jinotega" typeStatus="paratype">
<collectingRegion id="49A9F824C455FFB01535FD3E207CA4A4" box="[882,1124,724,750]" country="Nicaragua" name="Jinotega" pageId="8" pageNumber="31">Jinotega department</collectingRegion>
,
<collectingMunicipality id="6BB6ACBCC455FFB01235FD3E213EA4A4" box="[1138,1318,724,750]" pageId="8" pageNumber="31">Cerro Kilambe</collectingMunicipality>
,
<location id="8EB2601DC455FFB0174FFD16254EA55C" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB0174FFD16254EA55C" box="[264,342,764,790]" country="Nicaragua" municipality="Cerro Kilambe" name="Camp." pageId="8" pageNumber="31" stateProvince="Jinotega">Camp.</location>
2, 1550 m, 10/
<date id="FFD31006C455FFB0144EFD1626BEA55C" box="[521,678,764,790]" pageId="8" pageNumber="31" value="1998-05-17">
<collectingDate id="EF97E9EEC455FFB0144EFD1626BEA55C" box="[521,678,764,790]" pageId="8" pageNumber="31" value="1998-05-17">17 May 1998</collectingDate>
</date>
,
<collectorName id="26985310C455FFB014F4FD162761A55C" box="[691,889,764,790]" pageId="8" pageNumber="31">Cols. J. M. Maes</collectorName>
&amp;
<collectorName id="26985310C455FFB015E7FD162059A55C" box="[928,1089,764,790]" pageId="8" pageNumber="31">B. Hernández</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01215FD1620B7A55C" box="[1106,1199,764,790]" pageId="8" pageNumber="31" type="male">3 males</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB012FCFD16213BA55C" box="[1211,1315,764,790]" pageId="8" pageNumber="31" type="female">1 female</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01757FCCE2571A574" box="[272,361,804,830]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB01739FCCE2670A52C" collectingDate="2003-05-30" collectionCode="MELN" collectorName="Col. D. Roiz" country="Nicaragua" elevation="1250" location="Finca Santa Enriqueta" pageId="8" pageNumber="31" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Jinotega" typeStatus="paratype">
<locationDeviation id="9A602FE4C455FFB01739FCCE2627A574" box="[382,575,804,830]" location="Jinotega" pageId="8" pageNumber="31">
<quantity id="4C959B23C455FFB01739FCCE25A1A574" box="[382,441,804,830]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="31" unit="km" value="9.0">9 km</quantity>
E
<collectingRegion id="49A9F824C455FFB0179CFCCE2627A574" box="[475,575,804,830]" country="Nicaragua" name="Jinotega" pageId="8" pageNumber="31">Jinotega</collectingRegion>
</locationDeviation>
,
<location id="8EB2601DC455FFB0140CFCCE2749A574" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB0140CFCCE2749A574" box="[587,849,804,830]" country="Nicaragua" name="Finca Santa Enriqueta" pageId="8" pageNumber="31" stateProvince="Jinotega">Finca Santa Enriqueta</location>
,
<quantity id="4C959B23C455FFB0151AFCCE27A0A574" box="[861,952,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="31" unit="m" value="1250.0">
<elevation id="0040D1F5C455FFB0151AFCCE27A0A574" box="[861,952,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="31" unit="m" value="1250.0">1250 m</elevation>
</quantity>
, 24/
<date id="FFD31006C455FFB015ADFCCE2091A574" box="[1002,1161,804,830]" pageId="8" pageNumber="31" value="2003-05-30">
<collectingDate id="EF97E9EEC455FFB015ADFCCE2091A574" box="[1002,1161,804,830]" pageId="8" pageNumber="31" value="2003-05-30">30 May 2003</collectingDate>
</date>
,
<collectorName id="26985310C455FFB012D1FCCE2132A574" box="[1174,1322,804,830]" pageId="8" pageNumber="31">Col. D. Roiz</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01756FCA62577A52C" box="[273,367,844,870]" pageId="8" pageNumber="31" type="male">6 males</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB0173CFCA625F7A52C" box="[379,495,844,870]" pageId="8" pageNumber="31" type="female">3 females</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01441FCA62678A52C" box="[518,608,844,870]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB01432FCA62709A5C4" collectingDate="1992-05-29" collectionCode="MELN" collectorName="Cols. &amp; Maes &amp; Pineda" country="Nicaragua" elevation="1500" location="Los Pinares" pageId="8" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Jinotega" typeStatus="paratype">
<locationDeviation id="9A602FE4C455FFB01432FCA6272CA52C" box="[629,820,844,870]" location="Jinotega" pageId="8" pageNumber="31">
<quantity id="4C959B23C455FFB01432FCA626A8A52C" box="[629,688,844,870]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="31" unit="km" value="5.0">5 km</quantity>
S
<collectingRegion id="49A9F824C455FFB01497FCA6272CA52C" box="[720,820,844,870]" country="Nicaragua" name="Jinotega" pageId="8" pageNumber="31">Jinotega</collectingRegion>
</locationDeviation>
,
<location id="8EB2601DC455FFB01506FCA627D5A52C" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB01506FCA627D5A52C" box="[833,973,844,870]" country="Nicaragua" name="Los Pinares" pageId="8" pageNumber="31" stateProvince="Jinotega">Los Pinares</location>
,
<quantity id="4C959B23C455FFB0159EFCA6202DA52C" box="[985,1077,844,870]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="31" unit="m" value="1500.0">
<elevation id="0040D1F5C455FFB0159EFCA6202DA52C" box="[985,1077,844,870]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="31" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB01205FCA620F9A52C" box="[1090,1249,844,870]" pageId="8" pageNumber="31" value="1992-05-29">
<collectingDate id="EF97E9EEC455FFB01205FCA620F9A52C" box="[1090,1249,844,870]" pageId="8" pageNumber="31" value="1992-05-29">29 May 1992</collectingDate>
</date>
,
<collectorName id="26985310C455FFB012AAFCA62133A52C" box="[1261,1323,844,870]" pageId="8" pageNumber="31">Cols.</collectorName>
J. M.
<collectorName id="26985310C455FFB01717FC9E2597A5C4" box="[336,399,884,910]" pageId="8" pageNumber="31">Maes</collectorName>
&amp; C.
<collectorName id="26985310C455FFB01799FC9E2635A5C4" box="[478,557,884,910]" pageId="8" pageNumber="31">Pineda</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01404FC9E268DA5C4" box="[579,661,884,910]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB014E8FC9E2711A5C4" box="[687,777,884,910]" pageId="8" pageNumber="31">MELN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B053C9BC455FFB0156FFC9E27CDA594" collectingDate="1994-06-08" collectionCode="HAHC, BDGC, EAPZ, IEXA, MXAL" country="Honduras" elevation="1030" location="Yuscaran" pageId="8" pageNumber="31" specimenCount="84" specimenCount-female="31" specimenCount-male="53" stateProvince="El Paraiso" typeStatus="paratype">
<collectingCountry id="F37A7656C455FFB0156FFC9E27D7A5C4" box="[808,975,884,910]" name="Honduras" pageId="8" pageNumber="31">HONDURAS</collectingCountry>
.
<collectingRegion id="49A9F824C455FFB0159BFC9E20FEA5C4" box="[988,1254,884,910]" country="Honduras" name="El Paraiso" pageId="8" pageNumber="31">El Paraiso department</collectingRegion>
,
<location id="8EB2601DC455FFB012B3FC9E2552A5FC" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB012B3FC9E2552A5FC" country="Honduras" name="Yuscaran" pageId="8" pageNumber="31" stateProvince="El Paraiso">Yuscarán</location>
,
<quantity id="4C959B23C455FFB0171EFC7625AFA5FC" box="[345,439,924,950]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="31" unit="m" value="1030.0">
<elevation id="0040D1F5C455FFB0171EFC7625AFA5FC" box="[345,439,924,950]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="31" unit="m" value="1030.0">1030 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB01781FC762643A5FC" box="[454,603,924,950]" pageId="8" pageNumber="31" value="1994-06-08">
<collectingDate id="EF97E9EEC455FFB01781FC762643A5FC" box="[454,603,924,950]" pageId="8" pageNumber="31" value="1994-06-08">8 June 1994</collectingDate>
</date>
,
<collectedFrom id="3871753BC455FFB0142CFC7620E1A5FC" box="[619,1273,924,950]" pageId="8" pageNumber="31">
on
<taxonomicName id="4C6D4D45C455FFB014D5FC7620E1A5FC" authority=", Cols. H. &amp; A. Howden &amp; B. Gill" authorityName="Cols. H. &amp; A. Howden &amp; B. Gill" box="[658,1273,924,950]" class="Magnoliopsida" family="Sapotaceae" genus="Pouteria" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="species" species="sapota">
<emphasis id="B919EAD4C455FFB014D5FC762749A5FF" box="[658,849,924,949]" italics="true" pageId="8" pageNumber="31">Pouteria sapota</emphasis>
, Cols. H. &amp; A. Howden &amp; B. Gill
</taxonomicName>
</collectedFrom>
(
<specimenCount id="9D6BFD4FC455FFB0134BFC762557A594" pageId="8" pageNumber="31" type="male">53 males</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB0171AFC2E25F8A594" box="[349,480,964,990]" pageId="8" pageNumber="31" type="female">31 females</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB017BEFC2E2648A594" box="[505,592,964,990]" pageId="8" pageNumber="31">HAHC</collectionCode>
,
<collectionCode id="ED7CAE03C455FFB01419FC2E26ABA594" box="[606,691,964,990]" pageId="8" pageNumber="31">BDGC</collectionCode>
,
<collectionCode id="ED7CAE03C455FFB01486FC2E2715A594" box="[705,781,964,990]" country="Honduras" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33268" lsid="urn:lsid:biocol.org:col:33268" name="Escuela Agricola Panamericana" pageId="8" pageNumber="31">EAPZ</collectionCode>
,
<collectionCode id="ED7CAE03C455FFB0155DFC2E277BA594" box="[794,867,964,990]" pageId="8" pageNumber="31">IEXA</collectionCode>
,
<collectionCode id="ED7CAE03C455FFB01536FC2E27D4A594" box="[881,972,964,990]" pageId="8" pageNumber="31">MXAL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB015A5FC2E25CAA264" collectingDate="1994-06-08" collectionCode="HAHC, BDGC" collectorName="Cols. &amp; H. &amp; A. Howden &amp; Gill" country="Honduras" elevation="1800" location="Cerro Monserrat" pageId="8" pageNumber="31" specimenCount="13" specimenCount-female="10" specimenCount-male="3" stateProvince="El Paraiso" typeStatus="paratype">
<location id="8EB2601DC455FFB015A5FC2E20B2A594" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB015A5FC2E20B2A594" box="[994,1194,964,990]" country="Honduras" name="Cerro Monserrat" pageId="8" pageNumber="31" stateProvince="El Paraiso">Cerro Monserrat</location>
,
<locationDeviation id="9A602FE4C455FFB012F0FC2E256EA24C" location="Yuscaran" pageId="8" pageNumber="31">
<quantity id="4C959B23C455FFB012F0FC2E20ECA594" box="[1207,1268,964,990]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="31" unit="km" value="7.0">7 km</quantity>
SW Yuscarán
</locationDeviation>
,
<quantity id="4C959B23C455FFB017C4FC0625F8A24C" box="[387,480,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="31" unit="m" value="1800.0">
<elevation id="0040D1F5C455FFB017C4FC0625F8A24C" box="[387,480,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="31" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FFD31006C455FFB017A8FC06269AA24C" box="[495,642,1004,1030]" pageId="8" pageNumber="31" value="1994-06-08">
<collectingDate id="EF97E9EEC455FFB017A8FC06269AA24C" box="[495,642,1004,1030]" pageId="8" pageNumber="31" value="1994-06-08">8 June 1994</collectingDate>
</date>
,
<collectorName id="26985310C455FFB014D7FC0627BDA24C" box="[656,933,1004,1030]" pageId="8" pageNumber="31">Cols. H. &amp; A. Howden</collectorName>
&amp; B.
<collectorName id="26985310C455FFB015B4FC062038A24C" box="[1011,1056,1004,1030]" pageId="8" pageNumber="31">Gill</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01274FC06208AA24C" box="[1075,1170,1004,1030]" pageId="8" pageNumber="31" type="male">3 males</specimenCount>
,
<specimenCount id="9D6BFD4FC455FFB012D8FC06213BA24C" box="[1183,1315,1004,1030]" pageId="8" pageNumber="31" type="female">10 females</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01757FBFE2570A264" box="[272,360,1044,1070]" pageId="8" pageNumber="31">HAHC</collectionCode>
,
<collectionCode id="ED7CAE03C455FFB01734FBFE25D2A264" box="[371,458,1044,1070]" pageId="8" pageNumber="31">BDGC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B053C9BC455FFB0179AFBFE256AA21C" collectingDate="1995-05-24" collectionCode="R, HAHC" collectorName="La Muralla &amp; Col. &amp; Morris" country="Honduras" location="Olancho department" pageId="8" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Olancho" typeStatus="paratype">
<collectingRegion id="49A9F824C455FFB0179AFBFE26D7A264" box="[477,719,1044,1070]" country="Honduras" name="Olancho" pageId="8" pageNumber="31">Olancho department</collectingRegion>
,
<collectorName id="26985310C455FFB0149DFBFE2747A264" box="[730,863,1044,1070]" pageId="8" pageNumber="31">La Muralla</collectorName>
,
<date id="FFD31006C455FFB0152BFBFE2012A264" box="[876,1034,1044,1070]" pageId="8" pageNumber="31" value="1995-05-24">
<collectingDate id="EF97E9EEC455FFB0152BFBFE2012A264" box="[876,1034,1044,1070]" pageId="8" pageNumber="31" value="1995-05-24">24 May 1995</collectingDate>
</date>
,
<collectorName id="26985310C455FFB01251FBFE2050A264" box="[1046,1096,1044,1070]" pageId="8" pageNumber="31">Col.</collectorName>
<collectionCode id="ED7CAE03C455FFB01208FBFE207EA264" box="[1103,1126,1044,1070]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="31">R</collectionCode>
.
<collectorName id="26985310C455FFB01235FBFE20DAA264" box="[1138,1218,1044,1070]" pageId="8" pageNumber="31">Morris</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01293FBFE213BA264" box="[1236,1315,1044,1070]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB01757FBD62571A21C" box="[272,361,1084,1110]" pageId="8" pageNumber="31">HAHC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053C9BC455FFB01739FBD620D2A21C" box="[382,1226,1084,1110]" collectingDate="1995-05-27" collectionCode="HAHC" collectorName="Col. J. E. Wappes" country="Honduras" location="La Muralla" pageId="8" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Olancho" typeStatus="paratype">
P. N.
<location id="8EB2601DC455FFB017F9FBD6265CA21C" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB017F9FBD6265CA21C" box="[446,580,1084,1110]" country="Honduras" name="La Muralla" pageId="8" pageNumber="31" stateProvince="Olancho">La Muralla</location>
, 24/
<date id="FFD31006C455FFB01432FBD6270BA21C" box="[629,787,1084,1110]" pageId="8" pageNumber="31" value="1995-05-27">
<collectingDate id="EF97E9EEC455FFB01432FBD6270BA21C" box="[629,787,1084,1110]" pageId="8" pageNumber="31" value="1995-05-27">27 May 1995</collectingDate>
</date>
,
<collectorName id="26985310C455FFB01567FBD627F7A21C" box="[800,1007,1084,1110]" pageId="8" pageNumber="31">Col. J. E. Wappes</collectorName>
(
<specimenCount id="9D6BFD4FC455FFB01245FBD62049A21C" box="[1026,1105,1084,1110]" pageId="8" pageNumber="31" type="male">1 male</specimenCount>
) (
<collectionCode id="ED7CAE03C455FFB0122EFBD620DBA21C" box="[1129,1219,1084,1110]" pageId="8" pageNumber="31">HAHC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC455FFB0177FFB8E262FA014" pageId="8" pageNumber="31" type="biology_ecology">
<paragraph id="8BD236C6C455FFB0177FFB8E2073A2BC" blockId="8.[264,1323,283,1710]" pageId="8" pageNumber="31">
<emphasis id="B919EAD4C455FFB0177FFB8E258CA234" bold="true" box="[312,404,1124,1150]" pageId="8" pageNumber="31">Biology</emphasis>
. This species inhabits montane moist forests located at altitudes from
<quantity id="4C959B23C455FFB01297FB8E257DA2EC" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="8" pageNumber="31" unit="m" value="1400.0" valueMax="1800.0" valueMin="1000.0">1000 to 1800 m</quantity>
. Months of collection: May (24), June (101), July (8). Specimens were collected on
<taxonomicName id="4C6D4D45C455FFB01769FB5E25F3A287" box="[302,491,1204,1229]" class="Magnoliopsida" family="Sapotaceae" genus="Pouteria" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="species" species="sapota">
<emphasis id="B919EAD4C455FFB01769FB5E25F3A287" box="[302,491,1204,1229]" italics="true" pageId="8" pageNumber="31">Pouteria sapota</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C6D4D45C455FFB0145BFB5E26BCA284" box="[540,676,1204,1230]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
) in
<collectingRegion id="49A9F824C455FFB01494FB5E27C3A284" box="[723,987,1204,1230]" country="Honduras" name="El Paraiso" pageId="8" pageNumber="31">El Paraiso department</collectingRegion>
,
<collectingCountry id="F37A7656C455FFB015AFFB5E2042A284" box="[1000,1114,1204,1230]" name="Honduras" pageId="8" pageNumber="31">Honduras</collectingCountry>
and on
<taxonomicName id="4C6D4D45C455FFB012FDFB5E2132A287" box="[1210,1322,1204,1229]" class="Liliopsida" family="Poaceae" genus="Zea" kingdom="Plantae" order="Poales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="species" species="mays">
<emphasis id="B919EAD4C455FFB012FDFB5E2132A287" box="[1210,1322,1204,1229]" italics="true" pageId="8" pageNumber="31">Zea mays</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C6D4D45C455FFB01777FB36258BA2BC" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[304,403,1244,1270]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="31" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) and unidentified weeds at
<collectingRegion id="49A9F824C455FFB01491FB3627C7A2BC" box="[726,991,1244,1270]" country="Nicaragua" name="Matagalpa" pageId="8" pageNumber="31">Matagalpa department</collectingRegion>
,
<collectingCountry id="F37A7656C455FFB015ACFB36207FA2BC" box="[1003,1127,1244,1270]" name="Nicaragua" pageId="8" pageNumber="31">Nicaragua</collectingCountry>
.
</paragraph>
<paragraph id="8BD236C6C455FFB0177FFAEE279BA30C" blockId="8.[264,1323,283,1710]" pageId="8" pageNumber="31">
<emphasis id="B919EAD4C455FFB0177FFAEE2664A354" bold="true" box="[312,636,1284,1310]" pageId="8" pageNumber="31">Geographical distribution</emphasis>
.
<materialsCitation id="3B053C9BC455FFB014CCFAEE2767A30C" country="Honduras" latitude="15.0" location="Mountains" longLatPrecision="40878" longitude="-87.0" municipality="Central America" pageId="8" pageNumber="31" specimenCount="1">
<location id="8EB2601DC455FFB014CCFAEE2710A354" LSID="urn:lsid:plazi:treatment:03C487D0C45BFFB0174FF957260FA0E4:8EB2601DC455FFB014CCFAEE2710A354" box="[651,776,1284,1310]" country="Honduras" latitude="15.0" longLatPrecision="40878" longitude="-87.0" municipality="Central America" name="Mountains" pageId="8" pageNumber="31">Mountains</location>
in the southeast of
<collectingCountry id="F37A7656C455FFB015ABFAEE2046A354" box="[1004,1118,1284,1310]" name="Honduras" pageId="8" pageNumber="31">Honduras</collectingCountry>
and northwest of
<collectingCountry id="F37A7656C455FFB0174FFAC6259BA30C" box="[264,387,1324,1350]" name="Nicaragua" pageId="8" pageNumber="31">Nicaragua</collectingCountry>
,
<collectingMunicipality id="6BB6ACBCC455FFB017C8FAC62649A30C" box="[399,593,1324,1350]" pageId="8" pageNumber="31">Central America</collectingMunicipality>
(13°
<geoCoordinate id="EE595001C455FFB014DCFAC626FAA30C" box="[667,738,1324,1350]" degrees="15" direction="north" orientation="latitude" pageId="8" pageNumber="31" precision="55555" value="15.0">15° N</geoCoordinate>
, 86°
<geoCoordinate id="EE595001C455FFB01560FAC6276EA30C" box="[807,886,1324,1350]" degrees="87" direction="west" orientation="longitude" pageId="8" pageNumber="31" precision="55555" value="-87.0">87° W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD236C6C455FFB0177FFABE262FA014" blockId="8.[264,1323,283,1710]" pageId="8" pageNumber="31">
<emphasis id="B919EAD4C455FFB0177FFABE2655A324" bold="true" box="[312,589,1364,1390]" pageId="8" pageNumber="31">Taxonomic comments</emphasis>
.
<taxonomicName id="4C6D4D45C455FFB01428FABE2764A327" box="[623,892,1364,1389]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B919EAD4C455FFB01428FABE2764A327" box="[623,892,1364,1389]" italics="true" pageId="8" pageNumber="31">Macrodactylus tibialis</emphasis>
</taxonomicName>
is the only known species in this genus with an acute spine on the inner border of the male protibia. It is included in the “
<taxonomicName id="4C6D4D45C455FFB0134CFA96259CA3F7" authorityName="Guerin-Meneville" authorityYear="1844" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B919EAD4C455FFB0134CFA96213BA3DF" box="[1291,1315,1404,1429]" italics="true" pageId="8" pageNumber="31">M</emphasis>
.
<emphasis id="B919EAD4C455FFB0174FFA4E259CA3F7" box="[264,388,1444,1469]" italics="true" pageId="8" pageNumber="31">dimidiatus</emphasis>
</taxonomicName>
” group (
<emphasis id="B919EAD4C455FFB017B2FA4E262EA3F7" box="[501,566,1444,1469]" italics="true" pageId="8" pageNumber="31">sensu</emphasis>
Arce­Pérez and
<bibRefCitation id="EFFC4B37C455FFB01542FA4E27BCA3F4" author="Moron, M. A." box="[773,932,1444,1470]" pageId="8" pageNumber="31" pagination="123 - 239" refId="ref6864" refString="Arce-Perez R. &amp; Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) de Mexico y America del Norte. Acta Zoologica Mexicana (n. s), 79, 123 - 239." type="journal article" year="2000">Morón, 2000</bibRefCitation>
) because the dorsal surface and pygidium are glabrous and lack the scale­like setae typical of other groups of species, the tarsal segments do not have rings of white setae, the parameres are stout and are without setae on the external borders, and the genital plates of the females are large with the lateral borders widely expanded.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC455FFB0177FF986260FA0E4" pageId="8" pageNumber="31" type="etymology">
<paragraph id="8BD236C6C455FFB0177FF986260FA0E4" blockId="8.[264,1323,283,1710]" pageId="8" pageNumber="31">
<emphasis id="B919EAD4C455FFB0177FF98625A5A0CC" bold="true" box="[312,445,1644,1670]" pageId="8" pageNumber="31">Etymology</emphasis>
. The Latinized name refers to the sexually dimorphic structure of the protibia in this species.
</paragraph>
</subSubSection>
</treatment>
</document>