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<mods:title id="7FE7183EB9B736FA722654F3D5E120F2">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="40527DADBAB7ECC30D5FB317956A45FA">Kim, Il-Hoi</mods:namePart>
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<mods:affiliation id="382EBE6271B52F0362B4689B14D1F7DB">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:namePart id="754CCB7077898879DCF375F45885CE8F">Boxshall, Geoff A.</mods:namePart>
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<mods:affiliation id="41537870081B892B9766175A859EE955">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<treatment id="03C487CBEEBC3BD8FCEFFEFAFBA2F95E" ID-DOI="http://doi.org/10.5281/zenodo.4591290" ID-GBIF-Taxon="194475203" ID-Zenodo-Dep="4591290" LSID="urn:lsid:plazi:treatment:03C487CBEEBC3BD8FCEFFEFAFBA2F95E" httpUri="http://treatment.plazi.org/id/03C487CBEEBC3BD8FCEFFEFAFBA2F95E" lastPageNumber="494" pageId="493" pageNumber="494">
<subSubSection id="C3776556EEBC3BD8FCEFFEFAFB87FE8E" box="[810,1090,308,335]" pageId="493" pageNumber="494" type="nomenclature">
<paragraph id="8BD236DDEEBC3BD8FCEFFEFAFB87FE8E" blockId="493.[810,1090,308,335]" box="[810,1090,308,335]" pageId="493" pageNumber="494">
<heading id="D09A81B1EEBC3BD8FCEFFEFAFB87FE8E" bold="true" box="[810,1090,308,335]" fontSize="11" level="1" pageId="493" pageNumber="494" reason="1">
<emphasis id="B919EACFEEBC3BD8FCEFFEFAFB87FE8E" bold="true" box="[810,1090,308,335]" pageId="493" pageNumber="494">
<taxonomicName id="4C6D4D5EEEBC3BD8FCEFFEFAFC08FE8E" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,973,308,335]" class="Maxillopoda" family="Notodelphyidae" genus="Pentachaetus" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEEBC3BD8FCEFFEFAFC08FE8E" bold="true" box="[810,973,308,335]" italics="true" pageId="493" pageNumber="494">Pentachaetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EEBC3BD8FC11FEFBFB87FE8E" box="[980,1090,309,335]" pageId="493" pageNumber="494" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3776556EEBC3BD8FCEFFEB7FA5BFB63" pageId="493" pageNumber="494" type="diagnosis">
<paragraph id="8BD236DDEEBC3BD8FCEFFEB7FA5BFB63" blockId="493.[810,1452,377,1696]" pageId="493" pageNumber="494">
<emphasis id="B919EACFEEBC3BD8FCEFFEB7FC58FE52" bold="true" box="[810,925,377,403]" pageId="493" pageNumber="494">Diagnosis</emphasis>
. Body form as in
<taxonomicName id="4C6D4D5EEEBC3BD8FBBAFEB4FB20FE52" authorityName="Kerschner" authorityYear="1879" box="[1151,1253,378,403]" class="Maxillopoda" family="Notodelphyidae" genus="Doroixys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEEBC3BD8FBBAFEB4FB20FE52" box="[1151,1253,378,403]" italics="true" pageId="493" pageNumber="494">Doroixys</emphasis>
</taxonomicName>
. Dorsal cephalic shield bearing paired horn-like processes posterolaterally. Fifth pedigerous somite fused with fourth. Free urosome 5-segmented. Caudal ramus bearing 5 well-developed setae (2 proximal and 3 distal). Rostrum well-developed. Antennule 6-segmented; first segment armed with 2 setae. Antenna 3-segmented, comprising coxa, basis, and unsegmented endopod; terminal claw small. Labrum weak, flexible. Mandible similar to that of
<taxonomicName id="4C6D4D5EEEBC3BD8FAFBFD44FA61FD62" authorityName="Kerschner" authorityYear="1879" box="[1342,1444,650,675]" class="Maxillopoda" family="Notodelphyidae" genus="Doroixys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEEBC3BD8FAFBFD44FA61FD62" box="[1342,1444,650,675]" italics="true" pageId="493" pageNumber="494">Doroixys</emphasis>
</taxonomicName>
; exopod armed with 5 setae; endopod 2-segmented with 1 seta on first segment and 4 setae on second. Maxillule as in
<taxonomicName id="4C6D4D5EEEBC3BD8FC8CFD3EFC6AFCC8" authorityName="Kerschner" authorityYear="1879" box="[841,943,752,777]" class="Maxillopoda" family="Notodelphyidae" genus="Doroixys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEEBC3BD8FC8CFD3EFC6AFCC8" box="[841,943,752,777]" italics="true" pageId="493" pageNumber="494">Doroixys</emphasis>
</taxonomicName>
, lacking coxal endite. Maxilla 4-segmented, bearing 4 setae on first endite of syncoxa, claw plus 2 setaeon basis, and 2-segmented endopodarmedwith 1 and 3 setae on first and second segments, respectively. Maxilliped unsegmented with 6 setae. Leg 1 with 2- or 3-segmented exopod and 3-segmented endopod; second exopodal segment (or original second exopodal segment) lacking outer element. Leg 2 with 3-segmented rami. Legs 3 and 4 with 3-segmented exopods and 2-segmented endopods. Inner coxal seta absent in legs 14. First and second exopodal segments of legs 24 lacking inner seta. Legs 24 with setal armature only on terminal endopodal segments. Leg 5 represented by 2 processes and 2 setae.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EEBC3BD8FC9AFB67FC66FAE8" pageId="493" pageNumber="494" type="etymology">
<paragraph id="8BD236DDEEBC3BD8FC9AFB67FC66FAE8" blockId="493.[810,1452,377,1696]" pageId="493" pageNumber="494">
<emphasis id="B919EACFEEBC3BD8FC9AFB67FC25FB02" bold="true" box="[863,992,1193,1219]" pageId="493" pageNumber="494">Etymology</emphasis>
. The generic name is derived from the Greek
<emphasis id="B919EACFEEBC3BD8FCBDFB02FC73FB24" box="[888,950,1228,1253]" italics="true" pageId="493" pageNumber="494">penta</emphasis>
(=five) and
<emphasis id="B919EACFEEBC3BD8FB8FFB05FB42FB24" box="[1098,1159,1227,1253]" italics="true" pageId="493" pageNumber="494">chaet</emphasis>
(=a bristle) and refers to the presence of five setae on the caudal ramus. Gender masculine.
</paragraph>
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<paragraph id="8BD236DDEEBC3BD8FC9AFAFFFBF1FAAC" blockId="493.[810,1452,377,1696]" pageId="493" pageNumber="494">
<emphasis id="B919EACFEEBC3BD8FC9AFAFFFC37FA8A" bold="true" box="[863,1010,1329,1355]" pageId="493" pageNumber="494">
<typeStatus id="54D6887FEEBC3BD8FC9AFAFFFC37FA8A" box="[863,1010,1329,1355]" pageId="493" pageNumber="494">Typespecies</typeStatus>
</emphasis>
.
<taxonomicName id="4C6D4D5EEEBC3BD8FC3AFAFFFB3CFA8A" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1023,1273,1329,1355]" class="Maxillopoda" family="Notodelphyidae" genus="Pentachaetus" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="species" species="spinatus" status="gen. et sp. nov.">
<emphasis id="B919EACFEEBC3BD8FC3AFAFFFB3CFA8A" box="[1023,1273,1329,1355]" italics="true" pageId="493" pageNumber="494">Pentachaetusspinatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEBC3BD8FAC5FAFFFA6EFA8A" bold="true" box="[1280,1451,1329,1355]" pageId="493" pageNumber="494">
<taxonomicNameLabel id="A22A57B4EEBC3BD8FAC5FAFFFA6EFA8A" box="[1280,1451,1329,1355]" pageId="493" pageNumber="494" rank="species">gen. etsp. nov.</taxonomicNameLabel>
</emphasis>
by original designation.
</paragraph>
<paragraph id="8BD236DDEEBC3BD8FC9AFABBFAA2FA70" blockId="493.[810,1452,377,1696]" pageId="493" pageNumber="494">
<emphasis id="B919EACFEEBC3BD8FC9AFABBFBBFFA4E" bold="true" box="[863,1146,1397,1423]" pageId="493" pageNumber="494">Other included species</emphasis>
.
<taxonomicName id="4C6D4D5EEEBC3BD8FB4AFABBFA69FA4E" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1167,1452,1397,1423]" class="Maxillopoda" family="Notodelphyidae" genus="Pentachaetus" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="species" species="palauensis" status="gen. et sp. nov.">
<emphasis id="B919EACFEEBC3BD8FB4AFABBFA69FA4E" box="[1167,1452,1397,1423]" italics="true" pageId="493" pageNumber="494">Pentachaetus palauensis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEBC3BD8FCEFFA59FC1DFA70" bold="true" box="[810,984,1431,1457]" pageId="493" pageNumber="494">
<taxonomicNameLabel id="A22A57B4EEBC3BD8FCEFFA59FC1DFA70" box="[810,984,1431,1457]" pageId="493" pageNumber="494" rank="species">gen. etsp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C6D4D5EEEBC3BD8FBCBFA56FB74FA70" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1038,1201,1431,1457]" class="Maxillopoda" family="Notodelphyidae" genus="Pentachaetus" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="species" species="longisetatus" status="gen. et sp. nov.">
<emphasis id="B919EACFEEBC3BD8FBCBFA56FB74FA70" box="[1038,1201,1431,1457]" italics="true" pageId="493" pageNumber="494">P. longisetatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEBC3BD8FB7DFA59FAA2FA70" bold="true" box="[1208,1383,1431,1457]" pageId="493" pageNumber="494">
<taxonomicNameLabel id="A22A57B4EEBC3BD8FB7DFA59FAA2FA70" box="[1208,1383,1431,1457]" pageId="493" pageNumber="494" rank="species">gen. etsp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3776556EEBC3BD8FC9AFA77FBA2F95E" pageId="493" pageNumber="494" type="discussion">
<paragraph id="8BD236DDEEBC3BD8FC9AFA77FBA2F95E" blockId="493.[810,1452,377,1696]" pageId="493" pageNumber="494">
<emphasis id="B919EACFEEBC3BD8FC9AFA77FC09FA12" bold="true" box="[863,972,1465,1491]" pageId="493" pageNumber="494">Remarks</emphasis>
. The three species placed in this new genus constituteahomogeneousgroup andcanbe separatedfrom the closely related genus,
<emphasis id="B919EACFEEBC3BD8FBA4FA30FB08F9D6" box="[1121,1229,1534,1559]" italics="true" pageId="493" pageNumber="494">
<taxonomicName id="4C6D4D5EEEBC3BD8FBA4FA30FB0CF9D6" authorityName="Kerschner" authorityYear="1879" box="[1121,1225,1534,1559]" class="Maxillopoda" family="Notodelphyidae" genus="Doroixys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="493" pageNumber="494" phylum="Arthropoda" rank="genus">Doroixys</taxonomicName>
,
</emphasis>
by two prominent features: firstly, the caudal ramus is armedwith 5 (not 6) well-developed setae, and secondly, the second exopodal segment (or original second exopodal segment) of leg 1 lacks an outer setal element.
</paragraph>
</subSubSection>
</treatment>
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