136 lines
16 KiB
XML
136 lines
16 KiB
XML
<document id="43F24796E54CB573ADFCCF791DD5AE1C" ID-DOI="10.5281/zenodo.195310" ID-GBIF-Dataset="3a7afe4d-9527-4653-81a9-b3305e84bd0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="195310" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460225599725" checkinUser="plazi" docAuthor="Purchart, Luboš" docDate="2010" docId="03C487C7FFC9035F53B5FEC3A4746DEA" docLanguage="en" docName="zt02476p013.pdf" docOrigin="Zootaxa 2476" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hexarhopalus (Hexarhopalus) becvari Purchart, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="5" masterDocId="FFFDFFBFFFCA035B5322FFB1A6086D16" masterDocTitle="Review of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Borneo with description of new species" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1698238246819" updateUser="plazi">
|
||
<mods:mods id="06A3C7B301837598C1CA3FEA3ACD9BA1" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="84A94F8559F1DB02261DBFD6653E5978">
|
||
<mods:title id="81A09361529F4D1FF3ED8A3390553373">Review of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Borneo with description of new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="541B2E0D8B4BAB6F941E241E10A68170" type="personal">
|
||
<mods:role id="D96FB4A1F0C72211972B27DFD3D9EFC1">
|
||
<mods:roleTerm id="D7A8C39355D07780A162A01551313A35">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FBE501ED7FFF53DA21856F3300423E2D">Purchart, Luboš</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8C5AE80FC0C4EC67C4B7D3C7C25E969F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B69BAF83859170264001F635F276CAE9" type="host">
|
||
<mods:titleInfo id="1C0E31BEC5A18CA114D690DABE9377CE">
|
||
<mods:title id="DF98A3D8D708644E6B5C2FD70517C00E">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="00E40D97FFEF5A35FA7739D72F5DC069">
|
||
<mods:date id="42D612520C2B28AB3AC0EFAB6A84B908">2010</mods:date>
|
||
<mods:detail id="340F22894A98D357CEF8D4822E651C86" type="volume">
|
||
<mods:number id="3194754398E226CE219B7D193A06BB33">2476</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="64F138788FA6A279B49D98300C16CF34" unit="page">
|
||
<mods:start id="D07F402C7415DB9C85831F156530C499">1</mods:start>
|
||
<mods:end id="758E1BFE4F80A2AC3557012190FD36A7">13</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="81B0E341707E7648592E49610A76C1CB">journal article</mods:classification>
|
||
<mods:identifier id="395537F4A48F197E7532B1E38ED5F80C" type="DOI">10.5281/zenodo.195310</mods:identifier>
|
||
<mods:identifier id="077DF92CAC6C753EA983D44BD9FAFBEC" type="GBIF-Dataset">3a7afe4d-9527-4653-81a9-b3305e84bd0d</mods:identifier>
|
||
<mods:identifier id="B337DDBAC86447240546ABB316500402" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="3D1CB825BD7F81E4C9BBE7F581864FD4" type="Zenodo-Dep">195310</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C487C7FFC9035F53B5FEC3A4746DEA" ID-DOI="http://doi.org/10.5281/zenodo.6196167" ID-GBIF-Taxon="119386198" ID-Zenodo-Dep="6196167" LSID="urn:lsid:plazi:treatment:03C487C7FFC9035F53B5FEC3A4746DEA" httpUri="http://treatment.plazi.org/id/03C487C7FFC9035F53B5FEC3A4746DEA" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
<subSubSection id="C377655AFFC9035853B5FEC3A6E76CB9" pageId="3" pageNumber="4" type="nomenclature">
|
||
<paragraph id="8BD236D1FFC9035853B5FEC3A4C76C9A" blockId="3.[151,719,370,431]" box="[151,719,370,396]" pageId="3" pageNumber="4">
|
||
<heading id="D09A81BDFFC9035853B5FEC3A4C76C9A" bold="true" box="[151,719,370,396]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
|
||
<emphasis id="B919EAC3FFC9035853B5FEC3A4C76C9A" bold="true" box="[151,719,370,396]" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C6D4D52FFC9035853B5FEC3A4626C9A" box="[151,618,370,396]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="becvari" status="sp. nov." subGenus="Hexarhopalus">
|
||
<emphasis id="B919EAC3FFC9035853B5FEC3A74B6C9A" bold="true" box="[151,323,370,396]" italics="true" pageId="3" pageNumber="4">Hexarhopalus</emphasis>
|
||
(
|
||
<emphasis id="B919EAC3FFC903585276FEC3A4086C9A" bold="true" box="[340,512,370,396]" italics="true" pageId="3" pageNumber="4">Hexarhopalus</emphasis>
|
||
)
|
||
<emphasis id="B919EAC3FFC903585130FEC3A4626C9A" bold="true" box="[530,618,370,396]" italics="true" pageId="3" pageNumber="4">becvari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A57B8FFC903585153FEC3A4C76C9A" box="[625,719,370,396]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853B5FE24A6E76CB9" blockId="3.[151,719,370,431]" box="[151,239,405,431]" pageId="3" pageNumber="4">
|
||
(
|
||
<figureCitation id="13562A54FFC9035853BDFE24A6ED6CB9" box="[159,229,405,431]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1948,1972]" captionTargetBox="[157,1430,197,1925]" captionTargetId="figure@2.[151,1436,197,1925]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Habitus, dorsal view. 1. Hexarhopalus barclayi sp. nov. (holotype); 2. H. becvari sp. nov. (holotype); 3. H. ferreri sp. nov. (holotype); 4. H. iwani sp. nov. (holotype); 5. H. murudensis sp. nov. (holotype); 6. H. rolandi sp. nov. (holotype)." httpUri="https://zenodo.org/record/195311/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377655AFFC9035853B5FE6EA5896E82" pageId="3" pageNumber="4" type="materials_examined">
|
||
<paragraph id="8BD236D1FFC9035853B5FE6EA4DE6CEC" blockId="3.[151,1437,479,2036]" box="[151,726,479,506]" pageId="3" pageNumber="4">
|
||
<emphasis id="B919EAC3FFC9035853B5FE6EA7356CEF" bold="true" box="[151,317,479,505]" pageId="3" pageNumber="4">
|
||
<typeStatus id="54D68873FFC9035853B5FE6EA6DC6CEF" box="[151,212,479,505]" pageId="3" pageNumber="4">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Borneo, Sabah, Mount Trus Madi.
|
||
</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6FDB4A79D6F52" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">
|
||
<emphasis id="B919EAC3FFC9035853E6FDB4A7726F09" bold="true" box="[196,378,517,543]" pageId="3" pageNumber="4">
|
||
<typeStatus id="54D68873FFC9035853E6FDB4A7086F09" box="[196,256,517,543]" pageId="3" pageNumber="4">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54D68873FFC9035852A7FDB4A7FB6F09" box="[389,499,517,543]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
|
||
(♀
|
||
<collectionCode id="ED7CAE14FFC903585106FDB4A48B6F09" box="[548,643,517,543]" pageId="3" pageNumber="4">BMNH</collectionCode>
|
||
): BORNEO, Sabah, Mount Trus Madi,
|
||
<date id="FFD31011FFC90358577FFDB4A2E56F09" box="[1117,1261,517,543]" pageId="3" pageNumber="4" value="2005-04-12">12. iv. 2005</date>
|
||
, S. Chew leg.,
|
||
<collectionCode id="ED7CAE14FFC9035853B5FD9BA6E76F52" box="[151,239,554,580]" pageId="3" pageNumber="4">BMNH</collectionCode>
|
||
{E}
|
||
<date id="FFD31011FFC903585209FD9BA7996F52" box="[299,401,554,580]" pageId="3" pageNumber="4" value="2006" valueMax="2036">2006-36</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6FDFEA5896E82" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">
|
||
<emphasis id="B919EAC3FFC9035853E6FDFEA7ED6F7F" bold="true" box="[196,485,591,617]" pageId="3" pageNumber="4">Differential diagnosis.</emphasis>
|
||
Within the entire genus
|
||
<taxonomicName id="4C6D4D52FFC90358503AFDE1A5CD6F7F" box="[792,965,592,617]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B919EAC3FFC90358503AFDE1A5CD6F7F" box="[792,965,592,617]" italics="true" pageId="3" pageNumber="4">Hexarhopalus</emphasis>
|
||
</taxonomicName>
|
||
this species comes, due to corrugated surface of elytra and transversely connected interstriae, closest to
|
||
<taxonomicName id="4C6D4D52FFC9035850CEFDC4A3376F99" authority="Kaszab, 1960" authorityName="Kaszab" authorityYear="1960" box="[1004,1343,629,655]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sculptilis">
|
||
<emphasis id="B919EAC3FFC9035850CEFDC4A2816F98" box="[1004,1161,629,654]" italics="true" pageId="3" pageNumber="4">H. sculptilis</emphasis>
|
||
Kaszab, 1960
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6D4D52FFC90358565CFDC4A7C06FA2" authority="Kaszab, 1960" authorityName="Kaszab" authorityYear="1960" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis id="B919EAC3FFC90358565CFDC4A7136FA5" italics="true" pageId="3" pageNumber="4">H. birmanicus</emphasis>
|
||
Kaszab, 1960
|
||
</taxonomicName>
|
||
. It is, however very easily distinguishable as follows: pronotum longer than broad, oblique indentation in posterior angles large and deep, continuing with shallow groove anteriad, anterior half of pronotum dorso-laterally with two large swellings separated by a large, deep midlongitudinal depression situated in the first two thirds of pronotum, some elytral interstriae costate (in both above mentioned species pronotum is approximately as long as broad or slightly broader, oblique indentation in posterior angles extremely slightly indicated, pronotum without or with only slightly developed swellings and without large and deep depression, elytral interstriae convex, but not costate).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377655AFFC9035853E6FC2EA38E6AD9" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="8BD236D1FFC9035853E6FC2EA33A6EC9" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">
|
||
<emphasis id="B919EAC3FFC9035853E6FC2EA7556EAF" bold="true" box="[196,349,927,953]" pageId="3" pageNumber="4">Description.</emphasis>
|
||
Size of
|
||
<typeStatus id="54D68873FFC9035852E3FC11A4206EAC" box="[449,552,928,954]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
10.8 x
|
||
<quantity id="4C959B34FFC9035851A1FC11A4EA6EAC" box="[643,738,928,954]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="4" unit="mm" value="4.6">4.6 mm</quantity>
|
||
. Dark brown, dull, with labrum, maxillary palpi, antennae, tibiae, tarsi, basal portion of femora and elytral interstriae pale brown. Body elongate oval, glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6FC5BA2AF6962" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">Head smooth, glabrous, shagreened, entire surface simply and finely punctate. Narrower than pronotum in ratio 1:1.5. Frontoclypeal sulcus at the base U-shaped, very deep, almost reaching the clypeal margin. Distance between eyes 1.4 times as wide as their diameter (DV). Ocular sulcus normally developed, deep. Vertex very slightly depressed centrally. Antenna slender, 1.2 times as long as pronotum.</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6FB31A6F86889" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">Pronotum glabrous, shagreened, slightly dilated towards the anterior margin, broadest in anterior third, PW:PL ratio is 1:0.9. Base of pronotum rimmed, roughly and densely punctate (size of punctures approximately as large as an eye facet). Anterior half of pronotum with two large swellings situated dorsolaterally. The latter extremely finely and simply punctate. The space between the swellings depressed midlongitudinally and very finely sculptured. Oblique indentation in posterior angles deep, continuing with lateral shallow groove, reaching anterior third of pronotum. Lateral carina normally developed, only slightly obliterated anteriorly, bent upwards and creating rim of anterior margin of pronotum. The latter obliterated at middle.</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6FA1BA7856A2C" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">Elytra elongate oval, dilated posteriad, broadest behind middle and then directly narrowing towards the apex of elytra. 1.4 times as long as its width and 2.1 times longer and 1.4 as wide as pronotum. The basal margin of the elytra strongly carinated. Elytral suture flat. Each elytron with three corrugated costae, all of which end in two thirds of the elytral length. The first costae (inner – those nearest to elytral suture) not coalescent with the costa at elytral base but separated from them by approximately the distance of the combined length of the third and fourth antennomeres. The space between basal margin and the beginning of each first inner costa transversely with two large tubercles approximately as large as scutellum. Second and third costae coalescent with basal costa, apically (on elytral declivity) continuing with large longitudinal tubercles. The costae are connected by transverse wrinkles. The space between, and also after third (outer), costae with two rows of deep punctures. Elytral disc slightly convex (LV). Elytral declivity moderately steep. Scutellum triangular.</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6F8F4A3836A92" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">Prosternum, prosternal process, mesosventrite and metaventrite as smooth as prothoracic hypomeron and micropubescent. Metaventrite, 0.7 of length of mesocoxa. Abdominal ventrites smooth and micropubescent.</paragraph>
|
||
<paragraph id="8BD236D1FFC9035853E6F821A38E6AD9" blockId="3.[151,1437,479,2036]" pageId="3" pageNumber="4">Legs smooth, simply and finely punctate. Femora glabrous. Apical half of tibiae with relatively long yellowish setae underneath. Tarsi sparsely pubescent above and with brush-like yellowish setae underneath.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377655AFFC9035853E6F86BA4D16AE2" box="[196,729,2010,2036]" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="8BD236D1FFC9035853E6F86BA4D16AE2" blockId="3.[151,1437,479,2036]" box="[196,729,2010,2036]" pageId="3" pageNumber="4">
|
||
<emphasis id="B919EAC3FFC9035853E6F86BA76C6AE2" bold="true" box="[196,356,2010,2036]" pageId="3" pageNumber="4">Distribution.</emphasis>
|
||
Borneo, Sabah, Mt. Trus Madi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377655AFFCE035F53E6FF26A4746DEA" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="8BD236D1FFCE035F53E6FF26A4746DEA" blockId="4.[151,1436,151,252]" pageId="4" pageNumber="5">
|
||
<emphasis id="B919EAC3FFCE035F53E6FF26A7506DA7" bold="true" box="[196,344,151,177]" pageId="4" pageNumber="5">Etymology.</emphasis>
|
||
I am very pleased to name this new species in honour of my dear colleague and friend Stanislav Beĕvář jr. (
|
||
<collectingCountry id="F37A7641FFCE035F52AAFF0CA4416DC1" box="[392,585,189,215]" name="Czech Republic" pageId="4" pageNumber="5">Czech Republic</collectingCountry>
|
||
), who kindly allowed me to participate in his recent revision of the genus
|
||
<taxonomicName id="4C6D4D52FFCE035F53B5FF53A4716DEA" authority="Beevar & Purchart 2008" authorityName="Beevar & Purchart" authorityYear="2008" box="[151,633,225,252]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B919EAC3FFCE035F53B5FF53A7346DED" box="[151,316,226,251]" italics="true" pageId="4" pageNumber="5">Hexarhopalus</emphasis>
|
||
(Beĕvář &
|
||
<bibRefCitation id="EFFC4B20FFCE035F52E4FF53A4786DEA" author="Purchart" box="[454,624,226,252]" pageId="4" pageNumber="13" refString="Becvar, S. & Purchart, L. (2008) Revision of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera: Tenebrionidae: Cnodaloninae), with description of Malaysphena gen. nov. Annales Zoologici, 58, 35 - 70." type="journal article" year="2008">Purchart 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |