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<document id="D240A01B6F87F91432151399AD836E20" ID-CLB-Dataset="22479" ID-DOI="10.5281/zenodo.3776222" ID-GBIF-Dataset="c1129e00-0019-4411-9efb-a9d6c83fcf9d" ID-ISSN="0253-116X" ID-Zenodo-Dep="3776222" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588180845528" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2018" docId="03C487B82905B83ACE8FF8AEFD23767F" docLanguage="en" docName="Linzer.biol.Beitr.50.2.1015-1032.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Myllaena ambulans Assing 2018, nov.sp." docType="treatment" docVersion="7" lastPageNumber="1030" masterDocId="FFFDFFC0290AB835CE3DF876FFAB7050" masterDocTitle="On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1032" masterPageNumber="1015" pageNumber="1030" updateTime="1698838018870" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="52300448FB356F1039B86956C6BD0FAF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7A189997F916D5289C53AE6ACF68B00">
<mods:title id="75323B84E3FB1A49EE29F7F1E8BF6330">On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="F3CCB9054817408235A5AEDCA2FF2A63" type="personal">
<mods:role id="F179B354F7DA49CF0378D8AB1D71AAE0">
<mods:roleTerm id="C304CD1AC4679B8DB05FF7FC0474BF43">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4A2A87A4696279BE2BF2C9343BA059C">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="375FAA6E835C17FCCB53D92EEE9B8F71">text</mods:typeOfResource>
<mods:relatedItem id="E2027A5E856A952FA0C3BD7D5C172EEA" type="host">
<mods:titleInfo id="8DA38CF4C2CBC9AD09E87284CDDB07A9">
<mods:title id="7F41198B90496F26A3B30BDAD8BE9808">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="A0E7B1BC77E0646DB4F4BC664C9A85D1">
<mods:date id="C65252EF0296051F6F77C19988781798">2018</mods:date>
<mods:detail id="DA1631F2696C3E7CD3358459FF596A58" type="pubDate">
<mods:number id="59B7B0E8AC4EA70B6592D303313720F3">2018-12-17</mods:number>
</mods:detail>
<mods:detail id="383CE82FC12E30EA73DF543F82F781BA" type="volume">
<mods:number id="D81DEB06E8AC68E9859CD69B2F59B41F">50</mods:number>
</mods:detail>
<mods:detail id="4EAC78FFB90A96E1F83AED160237172A" type="issue">
<mods:number id="8C471DA01B377DD77B4B5D0EAE9B7768">2</mods:number>
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<mods:extent id="55F9D7B34938ECD349613E4C576C4159" unit="page">
<mods:start id="36574225D84ACB958717521751CCAB9F">1015</mods:start>
<mods:end id="ACC4E07AFC65D081270102843F643A6C">1032</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="EB61A1337E1497640E6AAF725E2BEBFC">journal article</mods:classification>
<mods:identifier id="655AE0B14EC562F8FC3D6AEB527ED40C" type="CLB-Dataset">22479</mods:identifier>
<mods:identifier id="4855376D43006D4AE06A0ACFBF79EAA4" type="DOI">10.5281/zenodo.3776222</mods:identifier>
<mods:identifier id="BD2B26893D234D5FE7A2820EB8FC58A1" type="GBIF-Dataset">c1129e00-0019-4411-9efb-a9d6c83fcf9d</mods:identifier>
<mods:identifier id="75404622FEA0A2BD7C54C7CB79C49AD4" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="9B12F0E1E8D7E9BAE3252F33CF836AAB" type="Zenodo-Dep">3776222</mods:identifier>
</mods:mods>
<treatment id="03C487B82905B83ACE8FF8AEFD23767F" ID-DOI="http://doi.org/10.5281/zenodo.3796589" ID-GBIF-Taxon="163876663" ID-Zenodo-Dep="3796589" LSID="urn:lsid:plazi:treatment:03C487B82905B83ACE8FF8AEFD23767F" httpUri="http://treatment.plazi.org/id/03C487B82905B83ACE8FF8AEFD23767F" lastPageNumber="1030" pageId="15" pageNumber="1030">
<subSubSection id="C37765252905B83ACE8FF8AEFE677199" pageId="15" pageNumber="1030" type="nomenclature">
<paragraph id="8BD236AE2905B83ACE8FF8AEFD0D70BF" blockId="15.[178,678,216,239]" box="[178,678,216,239]" pageId="15" pageNumber="1030">
<heading id="D09A81C22905B83ACE8FF8AEFD0D70BF" box="[178,678,216,239]" fontSize="8" level="2" pageId="15" pageNumber="1030" reason="2">
<emphasis id="B919EABC2905B83ACE8FF8AEFE7F70BE" bold="true" box="[178,468,216,238]" pageId="15" pageNumber="1030">
<taxonomicName id="4C6D4D2D2905B83ACE8FF8AEFE2A70BE" ID-CoL="8Y6J2" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[178,385,216,238]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="ambulans" status="sp. nov.">
<emphasis id="B919EABC2905B83ACE8FF8AEFE2A70BE" bold="true" box="[178,385,216,238]" italics="true" pageId="15" pageNumber="1030">Myllaena ambulans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57C72905B83ACFB5F8AFFE7F70BE" box="[392,468,217,238]" pageId="15" pageNumber="1030" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13562A2B2905B83ACFDCF8AFFDFA70BE" box="[481,593,217,239]" captionStart-0="Figs 33-38" captionStart-1="Figs 39-47" captionStartId-0="10.[178,218,1506,1528]" captionStartId-1="11.[178,218,1506,1528]" captionTargetBox-0="[179,1080,211,1483]" captionTargetBox-1="[185,1079,211,1485]" captionTargetId-0="figure@10.[178,1085,208,1493]" captionTargetId-1="figure@11.[177,1084,208,1493]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs 33-38: Myllaena ambulans: (33) forebody; (34-37) median lobe of aedeagus in lateral and in ventral view; (8) paramere. Scale bars: 33: 0.5 mm; 34-38: 0.1 mm." captionText-1="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3776234" figureDoi-1="http://doi.org/10.5281/zenodo.3776236" httpUri-0="https://zenodo.org/record/3776234/files/figure.png" httpUri-1="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Figs 33-43</figureCitation>
,
<figureCitation id="13562A2B2905B83ACC61F8AFFD3570BF" box="[604,670,217,239]" captionStart="Map 3" captionStartId="16.[178,224,1007,1028]" captionTargetBox="[177,1085,206,993]" captionTargetId="figure@16.[177,1085,206,993]" captionTargetPageId="16" captionText="Map 3: Distribution of Myllaena ambulans in Armenia." figureDoi="http://doi.org/10.5281/zenodo.3776242" httpUri="https://zenodo.org/record/3776242/files/figure.png" pageId="15" pageNumber="1030">Map 3</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FF888FE677199" blockId="15.[178,1085,253,457]" pageId="15" pageNumber="1030">
<materialsCitation id="3B053CF32905B83ACE8FF888FDE07111" ID-GBIF-Occurrence="2609436656" collectingDate="2016-07-03" country="Armenia" determinerName="V. Assing" elevation="2070" latitude="39.683056" location="35 km NW Sisian" longLatPrecision="19" longitude="45.780556" pageId="15" pageNumber="1030" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
T y p e m a t e r i a l:
<typeStatus id="54D6880C2905B83ACF8AF88BFDA07143" box="[439,523,253,275]" pageId="15" pageNumber="1030" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6BFD272905B83ACC25F888FD877143" box="[536,556,254,275]" pageId="15" pageNumber="1030" type="male">♂</specimenCount>
: &quot;
<collectingCountry id="F37A763E2905B83ACC79F88BFD1B7142" box="[580,688,253,274]" name="Armenia" pageId="15" pageNumber="1030">ARMENIA</collectingCountry>
[19] -
<location id="8EB260752905B83ACD3EF88BFC167143" LSID="urn:lsid:plazi:treatment:03C487B82905B83ACE8FF8AEFD23767F:8EB260752905B83ACD3EF88BFC167143" box="[771,957,253,275]" country="Armenia" latitude="39.683056" longLatPrecision="19" longitude="45.780556" name="35 km NW Sisian" pageId="15" pageNumber="1030">
<quantity id="4C959B4B2905B83ACD3EF88BFCEF7143" box="[771,836,253,275]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="1030" unit="km" value="35.0">35 km</quantity>
NW Sisian
</location>
,
<geoCoordinate id="EE5950692905B83ACDF0F88BFB927143" box="[973,1081,253,275]" degrees="39" direction="north" minutes="40" orientation="latitude" pageId="15" pageNumber="1030" precision="15" seconds="59" value="39.683056">39°40'59''N</geoCoordinate>
,
<geoCoordinate id="EE5950692905B83ACE8FF962FEB0717A" box="[178,283,276,298]" degrees="45" direction="east" minutes="46" orientation="longitude" pageId="15" pageNumber="1030" precision="15" seconds="50" value="45.780556">45°46'50''E</geoCoordinate>
,
<quantity id="4C959B4B2905B83ACF11F962FED17179" box="[300,378,276,297]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="15" pageNumber="1030" unit="m" value="2070.0">
<elevation id="0040D19D2905B83ACF11F962FED17179" box="[300,378,276,297]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="15" pageNumber="1030" unit="m" value="2070.0">2070 m</elevation>
</quantity>
, stream valley, sifted,
<date id="FFD3106E2905B83ACC58F962FD61717A" box="[613,714,276,298]" pageId="15" pageNumber="1030" value="2016-07-03">
<collectingDate id="EF97E9862905B83ACC58F962FD61717A" box="[613,714,276,298]" pageId="15" pageNumber="1030" value="2016-07-03">3.VII.2016</collectingDate>
</date>
, V. Assing /
<typeStatus id="54D6880C2905B83ACD60F962FC10717A" box="[861,955,276,298]" pageId="15" pageNumber="1030" type="holotype">Holotypus</typeStatus>
<specimenCount id="9D6BFD272905B83ACDF4F962FC717179" box="[969,986,276,297]" pageId="15" pageNumber="1030" type="male">♂</specimenCount>
<taxonomicName id="4C6D4D2D2905B83ACDDAF962FEA17110" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="ambulans" status="sp. nov.">
<emphasis id="B919EABC2905B83ACDDAF962FEA17110" italics="true" pageId="15" pageNumber="1030">Myllaena ambulans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57C72905B83ACF2DF95DFEE97110" box="[272,322,299,320]" pageId="15" pageNumber="1030" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="7E0EDA6A2905B83ACF52F95DFE617111" box="[367,458,299,321]" pageId="15" pageNumber="1030">V. Assing</determinerName>
2018&quot; (cAss)
</materialsCitation>
.
<materialsCitation id="3B053CF32905B83ACC69F95DFB927111" ID-GBIF-Occurrence="2609436624" box="[596,1081,299,321]" collectingDate="2016-07-03" country="Armenia" determinerName="V. Assing" elevation="2070" latitude="39.683056" location="35 km NW Sisian" longLatPrecision="19" longitude="45.780556" pageId="15" pageNumber="1030" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54D6880C2905B83ACC69F95DFD1B7110" box="[596,688,299,320]" pageId="15" pageNumber="1030" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D6BFD272905B83ACC84F95DFD437110" box="[697,744,299,320]" pageId="15" pageNumber="1030" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D6BFD272905B83ACCCFF95DFC897110" box="[754,802,299,320]" pageId="15" pageNumber="1030" type="female">3♀♀</specimenCount>
: same data as holotype (cAss)
</materialsCitation>
;
<materialsCitation id="3B053CF32905B83ACE8FF937FDEA7107" ID-GBIF-Occurrence="2609436633" box="[178,577,321,343]" collectingDate="2016-07-03" collectionCode="MNB" collectorName="Schulke" country="Armenia" determinerName="V. Assing" elevation="2070" latitude="39.683056" location="35 km NW Sisian" longLatPrecision="19" longitude="45.780556" pageId="15" pageNumber="1030" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D6BFD272905B83ACE8FF937FF7B7106" box="[178,208,321,342]" pageId="15" pageNumber="1030" type="female">1♀</specimenCount>
: same data, but leg.
<collectorName id="269853782905B83ACF9BF937FE447107" box="[422,495,321,343]" pageId="15" pageNumber="1030">Schülke</collectorName>
(
<collectionCode id="ED7CAE6B2905B83ACC3FF934FD917107" box="[514,570,322,343]" pageId="15" pageNumber="1030">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32905B83ACC72F937FC15713E" ID-GBIF-Occurrence="2609436635" collectingDate="2016-07-03" country="Armenia" elevation="2080" latitude="39.683613" location="35 km NW Sisian" longLatPrecision="19" longitude="45.779446" pageId="15" pageNumber="1030" specimenCount="10" specimenCount-female="7" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D6BFD272905B83ACC72F937FDD57106" box="[591,638,321,342]" pageId="15" pageNumber="1030" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D6BFD272905B83ACCB0F934FD157107" box="[653,702,322,343]" pageId="15" pageNumber="1030" type="female">7♀♀</specimenCount>
: &quot;
<collectingCountry id="F37A763E2905B83ACCE9F937FCEB7106" box="[724,832,321,342]" name="Armenia" pageId="15" pageNumber="1030">ARMENIA</collectingCountry>
[20]-
<location id="8EB260752905B83ACDB9F937FB917107" LSID="urn:lsid:plazi:treatment:03C487B82905B83ACE8FF8AEFD23767F:8EB260752905B83ACDB9F937FB917107" box="[900,1082,321,343]" country="Armenia" latitude="39.683613" longLatPrecision="19" longitude="45.779446" name="35 km NW Sisian" pageId="15" pageNumber="1030">
<quantity id="4C959B4B2905B83ACDB9F937FC687107" box="[900,963,321,343]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="1030" unit="km" value="35.0">35 km</quantity>
NW Sisian
</location>
,
<geoCoordinate id="EE5950692905B83ACE8FF92EFEB5713D" box="[178,286,344,365]" degrees="39" direction="north" minutes="41" orientation="latitude" pageId="15" pageNumber="1030" precision="15" seconds="01" value="39.683613">39°41'01''N</geoCoordinate>
,
<geoCoordinate id="EE5950692905B83ACF14F92EFE39713E" box="[297,402,344,366]" degrees="45" direction="east" minutes="46" orientation="longitude" pageId="15" pageNumber="1030" precision="15" seconds="46" value="45.779446">45°46'46''E</geoCoordinate>
,
<quantity id="4C959B4B2905B83ACFA3F92EFE43713D" box="[414,488,344,365]" metricMagnitude="3" metricUnit="m" metricValue="2.08" pageId="15" pageNumber="1030" unit="m" value="2080.0">
<elevation id="0040D19D2905B83ACFA3F92EFE43713D" box="[414,488,344,365]" metricMagnitude="3" metricUnit="m" metricValue="2.08" pageId="15" pageNumber="1030" unit="m" value="2080.0">2080 m</elevation>
</quantity>
, swampy meadow,
<date id="FFD3106E2905B83ACC9CF92EFCAD713E" box="[673,774,344,366]" pageId="15" pageNumber="1030" value="2016-07-03">
<collectingDate id="EF97E9862905B83ACC9CF92EFCAD713E" box="[673,774,344,366]" pageId="15" pageNumber="1030" value="2016-07-03">3.VII.2016</collectingDate>
</date>
, V. Assing&quot; (cAss)
</materialsCitation>
;
<materialsCitation id="3B053CF32905B83ACDF4F92EFE6E71D5" ID-GBIF-Occurrence="2609436662" collectingDate="2016-07-03" collectionCode="MNB" collectorName="Schulke" country="Armenia" elevation="2080" latitude="39.683613" location="35 km NW Sisian" longLatPrecision="19" longitude="45.779446" pageId="15" pageNumber="1030" specimenCount="7" typeStatus="paratype">
<specimenCount id="9D6BFD272905B83ACDF4F92EFBA8713D" box="[969,1027,344,365]" pageId="15" pageNumber="1030" type="generic">7 exs.</specimenCount>
, same data, but leg.
<collectorName id="269853782905B83ACF12F919FED371D5" box="[303,376,367,389]" pageId="15" pageNumber="1030">Schülke</collectorName>
(
<collectionCode id="ED7CAE6B2905B83ACFB8F919FE1671D5" box="[389,445,367,389]" pageId="15" pageNumber="1030">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32905B83ACFF3F919FD3D71CB" ID-GBIF-Occurrence="2609436626" collectingDate="2017-07-07" collectorName="V. Assing" country="Armenia" elevation="2000" latitude="40.7625" location="Spitak" longLatPrecision="19" longitude="44.273056" pageId="15" pageNumber="1030" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D6BFD272905B83ACFF3F919FE4671D4" box="[462,493,367,388]" pageId="15" pageNumber="1030" type="male">1♂</specimenCount>
: &quot;
<collectingCountry id="F37A763E2905B83ACFC2F919FDC071D4" box="[511,619,367,388]" name="Armenia" pageId="15" pageNumber="1030">ARMENIA</collectingCountry>
[33] – S
<location id="8EB260752905B83ACCFFF919FD5571D5" LSID="urn:lsid:plazi:treatment:03C487B82905B83ACE8FF8AEFD23767F:8EB260752905B83ACCFFF919FD5571D5" box="[706,766,367,389]" country="Armenia" latitude="40.7625" longLatPrecision="19" longitude="44.273056" name="Spitak" pageId="15" pageNumber="1030">Spitak</location>
,
<quantity id="4C959B4B2905B83ACD35F919FCFB71D4" box="[776,848,367,388]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="1030" unit="m" value="2000.0">
<elevation id="0040D19D2905B83ACD35F919FCFB71D4" box="[776,848,367,388]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="1030" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<geoCoordinate id="EE5950692905B83ACD67F918FC6D71D4" box="[858,966,366,388]" degrees="40" direction="north" minutes="45" orientation="latitude" pageId="15" pageNumber="1030" precision="15" seconds="45" value="40.7625">40°45'45''N</geoCoordinate>
,
<geoCoordinate id="EE5950692905B83ACDEDF919FB9271D5" box="[976,1081,367,389]" degrees="44" direction="east" minutes="16" orientation="longitude" pageId="15" pageNumber="1030" precision="15" seconds="23" value="44.273056">44°16'23''E</geoCoordinate>
, bank of small stream,
<date id="FFD3106E2905B83ACF43F9F3FE4871CA" box="[382,483,389,410]" pageId="15" pageNumber="1030" value="2017-07-07">
<collectingDate id="EF97E9862905B83ACF43F9F3FE4871CA" box="[382,483,389,410]" pageId="15" pageNumber="1030" value="2017-07-07">7.VII.2017</collectingDate>
</date>
,
<collectorName id="269853782905B83ACFD0F9F0FDE071CB" box="[493,587,389,411]" pageId="15" pageNumber="1030">V. Assing</collectorName>
&quot; (cAss)
</materialsCitation>
;
<materialsCitation id="3B053CF32905B83ACC9DF9F3FBBB71CB" ID-GBIF-Occurrence="2609436619" box="[672,1040,389,411]" collectingDate="2017-07-07" collectionCode="MNB" collectorName="Schulke" country="Armenia" elevation="2000" latitude="40.7625" location="Spitak" longLatPrecision="19" longitude="44.273056" pageId="15" pageNumber="1030" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D6BFD272905B83ACC9DF9F3FD1571CA" box="[672,702,389,410]" pageId="15" pageNumber="1030" type="female">1♀</specimenCount>
: same data, but leg.
<collectorName id="269853782905B83ACD46F9F3FC6F71CB" box="[891,964,389,411]" pageId="15" pageNumber="1030">Schülke</collectorName>
(
<collectionCode id="ED7CAE6B2905B83ACDECF9F3FBA271CB" box="[977,1033,389,411]" pageId="15" pageNumber="1030">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32905B83ACA27F9F3FE637199" ID-GBIF-Occurrence="2609436616" collectingDate="2017-07-08" collectorName="V. Assing" country="Armenia" elevation="2340" latitude="39.96611" location="Martuni" longLatPrecision="19" longitude="45.236946" municipality="Sulema Pass" pageId="15" pageNumber="1030" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D6BFD272905B83ACA27F9F3FB9271CA" box="[1050,1081,389,410]" pageId="15" pageNumber="1030" type="male">1♂</specimenCount>
: &quot;
<collectingCountry id="F37A763E2905B83ACE84F9EAFE8E71E1" box="[185,293,412,433]" name="Armenia" pageId="15" pageNumber="1030">ARMENIA</collectingCountry>
[36] – S
<location id="8EB260752905B83ACFAFF9EAFE7571E2" LSID="urn:lsid:plazi:treatment:03C487B82905B83ACE8FF8AEFD23767F:8EB260752905B83ACFAFF9EAFE7571E2" box="[402,478,412,434]" country="Armenia" latitude="39.96611" longLatPrecision="19" longitude="45.236946" municipality="Sulema Pass" name="Martuni" pageId="15" pageNumber="1030">Martuni</location>
,
<collectingMunicipality id="6BB6ACD42905B83ACFD0F9EAFDC371E1" box="[493,616,412,434]" pageId="15" pageNumber="1030">Sulema Pass</collectingMunicipality>
,
<geoCoordinate id="EE5950692905B83ACC4AF9EAFD4871E2" box="[631,739,412,434]" degrees="39" direction="north" minutes="57" orientation="latitude" pageId="15" pageNumber="1030" precision="15" seconds="58" value="39.96611">39°57'58''N</geoCoordinate>
,
<geoCoordinate id="EE5950692905B83ACCCEF9EAFCF771E1" box="[755,860,412,434]" degrees="45" direction="east" minutes="14" orientation="longitude" pageId="15" pageNumber="1030" precision="15" seconds="13" value="45.236946">45°14'13''E</geoCoordinate>
,
<quantity id="4C959B4B2905B83ACD56F9EAFC1271E1" box="[875,953,412,433]" metricMagnitude="3" metricUnit="m" metricValue="2.34" pageId="15" pageNumber="1030" unit="m" value="2340.0">
<elevation id="0040D19D2905B83ACD56F9EAFC1271E1" box="[875,953,412,433]" metricMagnitude="3" metricUnit="m" metricValue="2.34" pageId="15" pageNumber="1030" unit="m" value="2340.0">2340 m</elevation>
</quantity>
, near stream,
<date id="FFD3106E2905B83ACE8FF9C5FEBC7198" box="[178,279,435,456]" pageId="15" pageNumber="1030" value="2017-07-08">
<collectingDate id="EF97E9862905B83ACE8FF9C5FEBC7198" box="[178,279,435,456]" pageId="15" pageNumber="1030" value="2017-07-08">8.VII.2017</collectingDate>
</date>
,
<collectorName id="269853782905B83ACF1DF9C5FED67199" box="[288,381,435,457]" pageId="15" pageNumber="1030">V. Assing</collectorName>
&quot; (cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37765252905B83ACE8FF9A1FD23767F" pageId="15" pageNumber="1030" type="description">
<paragraph id="8BD236AE2905B83ACE8FF9A1FD6D725A" blockId="15.[178,1085,471,522]" pageId="15" pageNumber="1030">E t y m o l o g y: The specific epithet is the present participle of the Latin verb ambulare (to walk) and alludes to the reduced hind wings.</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFA6CFE5B723A" blockId="15.[178,1086,537,618]" pageId="15" pageNumber="1030">
D e s c r i p t i o n: Body length
<quantity id="4C959B4B2905B83ACC0CFA6CFD1A727F" box="[561,689,538,559]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="15" pageNumber="1030" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8-2.2 mm</quantity>
; length of forebody
<quantity id="4C959B4B2905B83ACD9DFA6FFB92727F" box="[928,1081,537,559]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.05" metricValueMin="0.95" pageId="15" pageNumber="1030" unit="mm" value="1.0" valueMax="1.05" valueMin="0.95">0.95-1.05 mm</quantity>
. Coloration: body blackish-brown to black; legs dark-brown to blackish-brown with paler tarsi; antennae blackish-brown.
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFA0FFC6E72B7" blockId="15.[178,1085,633,743]" pageId="15" pageNumber="1030">
Head (
<figureCitation id="13562A2B2905B83ACEC8FA0FFE9472C0" box="[245,319,633,656]" captionStart="Figs 33-38" captionStartId="10.[178,218,1506,1528]" captionTargetBox="[179,1080,211,1483]" captionTargetId="figure@10.[178,1085,208,1493]" captionTargetPageId="10" captionText="Figs 33-38: Myllaena ambulans: (33) forebody; (34-37) median lobe of aedeagus in lateral and in ventral view; (8) paramere. Scale bars: 33: 0.5 mm; 34-38: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776234" httpUri="https://zenodo.org/record/3776234/files/figure.png" pageId="15" pageNumber="1030">Fig. 33</figureCitation>
) wedge-shaped. Eyes relatively small, not protruding from lateral contours of head, shorter than the distance from posterior margin of eye to posterior constriction of head in lateral view. Antenna approximately
<quantity id="4C959B4B2905B83ACC84FAC2FCA07299" box="[697,779,692,714]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="1030" unit="mm" value="0.6">0.6 mm</quantity>
long; antennomeres IV-VIII approximately as long as broad (sometimes weakly oblong), IX-X transverse.
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFA81FD04737A" blockId="15.[178,1085,759,810]" pageId="15" pageNumber="1030">
Pronotum (
<figureCitation id="13562A2B2905B83ACF1BFA81FED9735D" box="[294,370,759,781]" captionStart="Figs 33-38" captionStartId="10.[178,218,1506,1528]" captionTargetBox="[179,1080,211,1483]" captionTargetId="figure@10.[178,1085,208,1493]" captionTargetPageId="10" captionText="Figs 33-38: Myllaena ambulans: (33) forebody; (34-37) median lobe of aedeagus in lateral and in ventral view; (8) paramere. Scale bars: 33: 0.5 mm; 34-38: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776234" httpUri="https://zenodo.org/record/3776234/files/figure.png" pageId="15" pageNumber="1030">Fig. 33</figureCitation>
) 1.3-1.4 times as broad as long and 1.3-1.4 times as broad as head; posterior margin not sinuate near posterior angles.
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFB4FFC9D73D9" blockId="15.[178,1086,825,906]" pageId="15" pageNumber="1030">
Elytra (
<figureCitation id="13562A2B2905B83ACF39FB4FFEFE7300" box="[260,341,825,848]" captionStart="Figs 33-38" captionStartId="10.[178,218,1506,1528]" captionTargetBox="[179,1080,211,1483]" captionTargetId="figure@10.[178,1085,208,1493]" captionTargetPageId="10" captionText="Figs 33-38: Myllaena ambulans: (33) forebody; (34-37) median lobe of aedeagus in lateral and in ventral view; (8) paramere. Scale bars: 33: 0.5 mm; 34-38: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776234" httpUri="https://zenodo.org/record/3776234/files/figure.png" pageId="15" pageNumber="1030">Fig. 33</figureCitation>
) short, approximately 0.65 times as long as pronotum. Hind wings reduced, rudiments only slightly protruding from under the elytra when fully unfolded. Posterior margin of abdominal tergite VIII with palisade fringe.
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFBECFC787457" blockId="15.[178,1085,921,1032]" pageId="15" pageNumber="1030">
<specimenCount id="9D6BFD272905B83ACE8FFBECFF6373FF" box="[178,200,922,943]" pageId="15" pageNumber="1030" type="male">♂</specimenCount>
: posterior margin of tergite VIII obtusely pointed in the middle (
<figureCitation id="13562A2B2905B83ACD62FBEFFC0773E0" box="[863,940,921,944]" captionStart="Figs 39-47" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[185,1079,211,1485]" captionTargetId="figure@11.[177,1084,208,1493]" captionTargetPageId="11" captionText="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776236" httpUri="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Fig. 39</figureCitation>
); sternite VIII (
<figureCitation id="13562A2B2905B83ACE84FBC1FEAD739D" box="[185,262,951,973]" captionStart="Figs 39-47" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[185,1079,211,1485]" captionTargetId="figure@11.[177,1084,208,1493]" captionTargetPageId="11" captionText="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776236" httpUri="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Fig. 40</figureCitation>
) of similar length as sternite VIII, posterior margin convex and with long thin setae; median lobe of aedeagus
<quantity id="4C959B4B2905B83ACC25FBA2FD0573B9" box="[536,686,980,1002]" metricMagnitude="-4" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.5" metricValueMin="3.2" pageId="15" pageNumber="1030" unit="mm" value="0.33499999999999996" valueMax="0.35" valueMin="0.32">0.32-0.35 mm</quantity>
long and shaped as in
<figureCitation id="13562A2B2905B83ACDFCFBA2FB9373B9" box="[961,1080,980,1002]" captionStart="Figs 33-38" captionStartId="10.[178,218,1506,1528]" captionTargetBox="[179,1080,211,1483]" captionTargetId="figure@10.[178,1085,208,1493]" captionTargetPageId="10" captionText="Figs 33-38: Myllaena ambulans: (33) forebody; (34-37) median lobe of aedeagus in lateral and in ventral view; (8) paramere. Scale bars: 33: 0.5 mm; 34-38: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776234" httpUri="https://zenodo.org/record/3776234/files/figure.png" pageId="15" pageNumber="1030">Figs 34-37</figureCitation>
; paramere (
<figureCitation id="13562A2B2905B83ACF23FB87FECC7458" box="[286,359,1009,1032]" captionStart="Figs 33-38" captionStartId="10.[178,218,1506,1528]" captionTargetBox="[179,1080,211,1483]" captionTargetId="figure@10.[178,1085,208,1493]" captionTargetPageId="10" captionText="Figs 33-38: Myllaena ambulans: (33) forebody; (34-37) median lobe of aedeagus in lateral and in ventral view; (8) paramere. Scale bars: 33: 0.5 mm; 34-38: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776234" httpUri="https://zenodo.org/record/3776234/files/figure.png" pageId="15" pageNumber="1030">Fig. 38</figureCitation>
) nearly
<quantity id="4C959B4B2905B83ACF86FB87FDA37457" box="[443,520,1009,1031]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="1030" unit="mm" value="0.5">0.5 mm</quantity>
long, with very long and slender apical lobe.
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFC61FD547437" blockId="15.[178,1085,1047,1127]" pageId="15" pageNumber="1030">
<specimenCount id="9D6BFD272905B83ACE8FFC61FF6E747C" box="[178,197,1047,1068]" pageId="15" pageNumber="1030" type="female">♀</specimenCount>
: tergite VIII (
<figureCitation id="13562A2B2905B83ACF6EFC61FE36747D" box="[339,413,1047,1069]" captionStart="Figs 39-47" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[185,1079,211,1485]" captionTargetId="figure@11.[177,1084,208,1493]" captionTargetPageId="11" captionText="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776236" httpUri="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Fig. 41</figureCitation>
) of similar shape as in male; sternite VIII (
<figureCitation id="13562A2B2905B83ACD6EFC61FC36747D" box="[851,925,1047,1069]" captionStart="Figs 39-47" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[185,1079,211,1485]" captionTargetId="figure@11.[177,1084,208,1493]" captionTargetPageId="11" captionText="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776236" httpUri="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Fig. 42</figureCitation>
) approximately as long as tergite VIII, posterior margin very weakly convex, nearly truncate, with long and rather stout marginal setae; spermatheca as in
<figureCitation id="13562A2B2905B83ACC8CFC27FD547437" box="[689,767,1105,1127]" captionStart="Figs 39-47" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[185,1079,211,1485]" captionTargetId="figure@11.[177,1084,208,1493]" captionTargetPageId="11" captionText="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776236" httpUri="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Fig. 43.</figureCitation>
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFC01FF7975E1" blockId="15.[178,1085,1143,1457]" pageId="15" pageNumber="1030">
C o m p a r a t i v e n o t e s:
<taxonomicName id="4C6D4D2D2905B83ACC2DFC01FD7274DC" authorityName="Assing" authorityYear="2018" box="[528,729,1143,1164]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="ambulans">
<emphasis id="B919EABC2905B83ACC2DFC01FD7274DC" box="[528,729,1143,1164]" italics="true" pageId="15" pageNumber="1030">Myllaena ambulans</emphasis>
</taxonomicName>
belongs to a group of small-sized species including
<emphasis id="B919EABC2905B83ACF55FCE3FE7B74FA" box="[360,464,1173,1194]" italics="true" pageId="15" pageNumber="1030">M. minuta</emphasis>
(GRAVENHORST, 1806),
<emphasis id="B919EABC2905B83ACCF3FCE3FC9674F9" box="[718,829,1172,1194]" italics="true" pageId="15" pageNumber="1030">M. gracilis</emphasis>
(MATTHEWS, 1838), and
<taxonomicName id="4C6D4D2D2905B83ACE8FFCC4FE9F7497" authorityName="Kraatz" authorityYear="1853" box="[178,308,1202,1223]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="B919EABC2905B83ACE8FFCC4FE9F7497" box="[178,308,1202,1223]" italics="true" pageId="15" pageNumber="1030">M. infuscata</emphasis>
</taxonomicName>
KRAATZ, 1853. It is distinguished from
<taxonomicName id="4C6D4D2D2905B83ACCDEFCC4FCCE7497" authorityName="Kraatz" authorityYear="1853" box="[739,869,1202,1223]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="B919EABC2905B83ACCDEFCC4FCCE7497" box="[739,869,1202,1223]" italics="true" pageId="15" pageNumber="1030">M. infuscata</emphasis>
</taxonomicName>
, the only species of this group recorded from
<collectingCountry id="F37A763E2905B83ACFFEFCB9FDB774B5" box="[451,540,1231,1253]" name="Armenia" pageId="15" pageNumber="1030">Armenia</collectingCountry>
and with elytra of reduced length, by even shorter elytra (
<taxonomicName id="4C6D4D2D2905B83ACEC6FC9BFED17551" authorityName="Kraatz" authorityYear="1853" box="[251,378,1260,1282]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="B919EABC2905B83ACEC6FC9BFED17551" box="[251,378,1260,1282]" italics="true" pageId="15" pageNumber="1030">M. infuscata</emphasis>
</taxonomicName>
: elytra usually 0.75-0.80 times as long as pronotum), very short hind wings, darker average coloration (
<taxonomicName id="4C6D4D2D2905B83ACC23FD7CFD09754F" authorityName="Kraatz" authorityYear="1853" box="[542,674,1290,1311]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="B919EABC2905B83ACC23FD7CFD09754F" box="[542,674,1290,1311]" italics="true" pageId="15" pageNumber="1030">M. infuscata</emphasis>
</taxonomicName>
: body mostly dark-brown to blackishbrown), the shapes of the apex of the ventral process (lateral view) and internal structures of the aedeagus, a less oblong and posteriorly less acute female tergite VIII, and by a larger distal portion of the spermatheca capsule with a more pronounced cuticular invagination. For illustrations of the primary sexual characters of
<taxonomicName id="4C6D4D2D2905B83ACD09FDF6FC1E75C4" authorityName="Kraatz" authorityYear="1853" box="[820,949,1407,1429]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1030" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="B919EABC2905B83ACD09FDF6FC1E75C4" box="[820,949,1407,1429]" italics="true" pageId="15" pageNumber="1030">M. infuscata</emphasis>
</taxonomicName>
see
<figureCitation id="13562A2B2905B83ACDDBFD09FF7975E1" captionStart="Figs 39-47" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[185,1079,211,1485]" captionTargetId="figure@11.[177,1084,208,1493]" captionTargetPageId="11" captionText="Figs 39-47: Myllaena ambulans (39-43) and M. infuscata from Armenia (44-47): (39) male tergite VIII; (40) male sternite VIII; (41) female tergite VIII; (42) female sternite VIII; (43, 46-47) spermatheca; (44-45) median lobe of aedeagus in lateral and in ventral view. Scale bars: 39-42: 0.2 mm; 43-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776236" httpUri="https://zenodo.org/record/3776236/files/figure.png" pageId="15" pageNumber="1030">Figs 44- 47.</figureCitation>
</paragraph>
<paragraph id="8BD236AE2905B83ACE8FFDB4FD23767F" blockId="15.[178,1085,1473,1583]" pageId="15" pageNumber="1030">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: This species is currently known from three localities in North, Central, and South
<collectingCountry id="F37A763E2905B83ACD36FDA9FCCF75A5" box="[779,868,1503,1525]" name="Armenia" pageId="15" pageNumber="1030">Armenia</collectingCountry>
(
<figureCitation id="13562A2B2905B83ACD4EFDA9FC1175A5" box="[883,954,1503,1525]" captionStart="Map 3" captionStartId="16.[178,224,1007,1028]" captionTargetBox="[177,1085,206,993]" captionTargetId="figure@16.[177,1085,206,993]" captionTargetPageId="16" captionText="Map 3: Distribution of Myllaena ambulans in Armenia." figureDoi="http://doi.org/10.5281/zenodo.3776242" httpUri="https://zenodo.org/record/3776242/files/figure.png" pageId="15" pageNumber="1030">Map 3</figureCitation>
). The specimens were collected from moist litter near streams, on stream and river banks, and in a swampy meadow at altitudes of
<quantity id="4C959B4B2905B83ACFC4FE6FFD2F767F" box="[505,644,1561,1583]" metricMagnitude="3" metricUnit="m" metricValue="2.01" metricValueMax="2.34" metricValueMin="1.68" pageId="15" pageNumber="1030" unit="m" value="2010.0" valueMax="2340.0" valueMin="1680.0">1680-2340 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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