treatments-xml/data/03/C0/30/03C030765321FFB7FE83B3DAFAFFF920.xml
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<document id="3FA081B5010938D3E84B397A25E1EAFB" ID-DOI="10.5281/zenodo.158124" ID-GBIF-Dataset="45e277ac-5921-4c12-b8ac-1ca07ec3857c" ID-ISSN="1175­5326" ID-Zenodo-Dep="158124" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461028957239" checkinUser="plazi" docAuthor="Tomikawa, Ko, Hendrycks, Ed A. &amp; Mawatari, Shunsuke F." docDate="2004" docId="03C030765321FFB7FE83B3DAFAFFF920" docLanguage="en" docName="zt00674.pdf" docOrigin="Zootaxa 674" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Dactylopleustes yoshimurai Tomikawa, Hendrycks &amp; Mawatari, 2004, sp. nov." docType="treatment" docUuid="43443907-473F-495A-A0F2-802C49A13100" docUuidSource="ZooBank" docVersion="10" lastPageNumber="10" masterDocId="FFF9480E5323FFBEFF8BB123FFD2FFF6" masterDocTitle="A new species of the genus Dactylopleustes (Crustacea: Amphipoda: Pleustidae) from Japan, with a partial redescription of D. echinoides Bousfield and Hendrycks, 1995" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1698573089058" updateUser="plazi">
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<mods:title id="6EADD3FE8F28C7C857365FA9748D1856">A new species of the genus Dactylopleustes (Crustacea: Amphipoda: Pleustidae) from Japan, with a partial redescription of D. echinoides Bousfield and Hendrycks, 1995</mods:title>
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<mods:name id="F82EA07D628AA8336187ABB30380C654" type="personal">
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<mods:namePart id="977ED55099B5B440746C63159AF0106B">Tomikawa, Ko</mods:namePart>
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<mods:namePart id="2E858F37B456C8C8F64D188078C1D463">Hendrycks, Ed A.</mods:namePart>
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<mods:namePart id="5F056DC514E52B5C6AED5C64D8FEE72F">Mawatari, Shunsuke F.</mods:namePart>
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<mods:title id="0E3AA0CF5A756738368E7603AB05F110">Zootaxa</mods:title>
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<mods:part id="7BF9B0242D376BC241445CE2BD80DA81">
<mods:date id="0368A51E5C0167E64D30D368F8D426AA">2004</mods:date>
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<mods:number id="C0BE060CC9CCC8C725A941AB4CB2A8A5">674</mods:number>
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<mods:identifier id="5FC8C0A9C23503A4C5B2B4FB20DE2DC1" type="DOI">10.5281/zenodo.158124</mods:identifier>
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<mods:identifier id="2A854F4AC336C33F308BC6106E69D0B0" type="Zenodo-Dep">158124</mods:identifier>
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<treatment id="03C030765321FFB7FE83B3DAFAFFF920" ID-DOI="http://doi.org/10.5281/zenodo.6273430" ID-GBIF-Taxon="119607143" ID-Zenodo-Dep="6273430" LSID="urn:lsid:zoobank.org:act:43443907-473F-495A-A0F2-802C49A13100" httpUri="http://treatment.plazi.org/id/03C030765321FFB7FE83B3DAFAFFF920" lastPageId="9" lastPageNumber="10" pageId="2" pageNumber="3">
<subSubSection id="C373D2EB5321FFBCFE83B3DAFE75FCA8" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BD681605321FFBCFE83B3DAFD68FCE5" blockId="2.[264,884,657,862]" box="[264,698,761,787]" pageId="2" pageNumber="3">
<heading id="D09E360C5321FFBCFE83B3DAFD68FCE5" bold="true" box="[264,698,761,787]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B91D5D725321FFBCFE83B3DAFD68FCE5" bold="true" box="[264,698,761,787]" pageId="2" pageNumber="3">
<taxonomicName id="4C69FAE35321FFBCFE83B3DAFD87FCE5" ID-CoL="33WCC" box="[264,597,761,787]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="yoshimurai" status="sp. nov.">
<emphasis id="B91D5D725321FFBCFE83B3DAFD87FCE5" bold="true" box="[264,597,761,787]" italics="true" pageId="2" pageNumber="3">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EE0095321FFBCFDD7B3DAFD68FCE5" box="[604,698,761,787]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD681605321FFBCFE83B23FFE75FCA8" blockId="2.[264,884,657,862]" pageId="2" pageNumber="3">
[Japanese name: Ezo­tengunouniyadori, new] (
<figureCitation id="13529DE55321FFBCFE9BB267FEAEFCA8" box="[272,380,836,862]" captionStart="FIGURE 1" captionStartId="3.[264,367,1797,1821]" captionTargetBox="[270,1311,326,1782]" captionTargetId="figure@3.[190,1325,308,1788]" captionTargetPageId="3" captionText="FIGURE 1. Dactylopleustes yoshimurai sp. nov., holotype female, 4.3 mm (NSMT­Cr 15894); paratype juvenile, 3.4 mm (CMNC 2004 ­ 1306). Lacinia mobilis of left mandible is indicated by an arrowhead. Scale bars = 0.1 mm unless indicated otherwise." httpUri="https://zenodo.org/record/158125/files/figure.png" pageId="2" pageNumber="3">Figures 1</figureCitation>
5)
</paragraph>
</subSubSection>
<subSubSection id="C373D2EB5321FFBCFE83B2AAFE2FFA88" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BD681605321FFBCFE83B2AAFE7BFC54" blockId="2.[264,1323,905,1171]" box="[264,425,905,930]" pageId="2" pageNumber="3">
<emphasis id="B91D5D725321FFBCFE83B2AAFE7BFC54" box="[264,425,905,930]" italics="true" pageId="2" pageNumber="3">
<typeStatus id="54D23FC25321FFBCFE83B2AAFEEDFC54" box="[264,319,905,930]" pageId="2" pageNumber="3">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BD681605321FFBCFEB3B292FEB0FB65" blockId="2.[264,1323,905,1171]" pageId="2" pageNumber="3">
<typeStatus id="54D23FC25321FFBCFEB3B292FE78FC3D" box="[312,426,945,971]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
: ovigerous female,
<quantity id="4C912C855321FFBCFD13B292FD24FC3D" box="[664,758,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="3" unit="mm" value="4.3">4.3 mm</quantity>
(12 slides), NSMT­Cr 15894, Itanki, Muroran City, Hokkaido,
<geoCoordinate id="EE5DE7A75321FFBCFE45B2FAFDE3FC05" box="[462,561,985,1011]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="42.333332">42°20'N</geoCoordinate>
,
<geoCoordinate id="EE5DE7A75321FFBCFDCBB2FAFD72FC05" box="[576,672,985,1011]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="141.01666">141°1'E</geoCoordinate>
,
<date id="FFD7A7A05321FFBCFD24B2FAFCA9FC05" box="[687,891,985,1011]" pageId="2" pageNumber="3" value="2003-10-17">17 October 2003</date>
, collector K. Yoshimura.
<typeStatus id="54D23FC25321FFBCFB3BB2FAFAF5FC05" box="[1200,1319,985,1011]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
: one ovigerous female,
<quantity id="4C912C855321FFBCFD99B522FDBEFBED" box="[530,620,1025,1051]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="3" unit="mm" value="5.4">5.4 mm</quantity>
(undissected), NSMT­Cr 15895; two juveniles,
<quantity id="4C912C855321FFBCFB14B522FB2BFBED" box="[1183,1273,1025,1051]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="3" unit="mm" value="3.3">3.3 mm</quantity>
and
<quantity id="4C912C855321FFBCFE83B50AFEB7FBB5" box="[264,357,1065,1091]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="3" unit="mm" value="3.4">3.4 mm</quantity>
(24 slides); three juveniles, 2.9, 2.9,
<quantity id="4C912C855321FFBCFCA5B50AFC59FBB5" box="[814,907,1065,1091]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="3" unit="mm" value="3.2">3.2 mm</quantity>
(undissected),
<collectionCode id="ED7819A55321FFBCFBC8B50AFB4BFBB5" box="[1091,1177,1065,1091]" pageId="2" pageNumber="3">CMNC</collectionCode>
2004­1306, same data as
<typeStatus id="54D23FC25321FFBCFE2CB572FDC3FB9D" box="[423,529,1105,1131]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
. All specimens were found in association with the urchin,
<taxonomicName id="4C69FAE35321FFBCFB40B572FE89FB64" class="Echinoidea" family="Strongylocentrotidae" genus="Strongylocentrotus" kingdom="Animalia" order="Camarodonta" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="intermedius">
<emphasis id="B91D5D725321FFBCFB40B572FE89FB64" italics="true" pageId="2" pageNumber="3">S. intermedius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD681605321FFBCFE83B5E7FE65FB2B" blockId="2.[264,1323,1220,1406]" box="[264,439,1220,1245]" pageId="2" pageNumber="3">
<emphasis id="B91D5D725321FFBCFE83B5E7FE65FB2B" box="[264,439,1220,1245]" italics="true" pageId="2" pageNumber="3">Other material</emphasis>
</paragraph>
<paragraph id="8BD681605321FFBCFEB3B5CFFE2FFA88" blockId="2.[264,1323,1220,1406]" pageId="2" pageNumber="3">
Two juveniles (
<quantity id="4C912C855321FFBCFE67B5CFFD98FAF0" box="[492,586,1260,1286]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="3" unit="mm" value="3.3">3.3 mm</quantity>
,
<quantity id="4C912C855321FFBCFDDEB5CFFD61FAF0" box="[597,691,1260,1286]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="3" unit="mm" value="3.4">3.4 mm</quantity>
, slide mounts), taken from
<emphasis id="B91D5D725321FFBCFC79B5CFFB72FAF3" box="[1010,1184,1260,1285]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C69FAE35321FFBCFC79B5CFFB4EFAF3" box="[1010,1180,1260,1285]" class="Echinoidea" family="Strongylocentrotidae" genus="Strongylocentrotus" kingdom="Animalia" order="Camarodonta" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="intermedius">S. intermedius</taxonomicName>
,
</emphasis>
Fukushima Town, Hokkaido,
<date id="FFD7A7A05321FFBCFE57B437FD6CFAD8" box="[476,702,1300,1326]" pageId="2" pageNumber="3" value="2003-12-17">17 December 2003</date>
, collector K. Yoshimura:
<specimenCount id="9D6F4AE95321FFBCFC67B437FBA9FAD8" box="[1004,1147,1300,1326]" pageId="2" pageNumber="3" type="juvenile">17 juveniles</specimenCount>
(
<quantity id="4C912C855321FFBCFB01B437FACFFAD8" box="[1162,1309,1300,1326]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.9" metricValueMin="2.7" pageId="2" pageNumber="3" unit="mm" value="3.3" valueMax="3.9" valueMin="2.7">2.73.9 mm</quantity>
), taken from
<taxonomicName id="4C69FAE35321FFBCFE05B41FFC52FAA0" authority="Tomiura, Noboribetsu City" authorityName="Tomiura, Noboribetsu City" box="[398,896,1340,1366]" class="Echinoidea" family="Strongylocentrotidae" genus="Strongylocentrotus" kingdom="Animalia" order="Camarodonta" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="intermedius">
<emphasis id="B91D5D725321FFBCFE05B41FFDE7FAA3" box="[398,565,1340,1365]" italics="true" pageId="2" pageNumber="3">S. intermedius</emphasis>
, Tomiura, Noboribetsu City
</taxonomicName>
, Hokkaido,
<date id="FFD7A7A05321FFBCFB86B41FFB23FAA0" box="[1037,1265,1340,1366]" pageId="2" pageNumber="3" value="2003-11-19">19 November 2003</date>
, collector K. Yoshimura.
</paragraph>
</subSubSection>
<subSubSection id="C373D2EB5321FFBCFE83B497FD44F910" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BD681605321FFBCFE83B497FEADFA3B" blockId="2.[264,1321,1460,1766]" box="[264,383,1460,1485]" pageId="2" pageNumber="3">
<emphasis id="B91D5D725321FFBCFE83B497FEADFA3B" box="[264,383,1460,1485]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD681605321FFBCFEB3B4FFFD44F910" blockId="2.[264,1321,1460,1766]" pageId="2" pageNumber="3">Antenna 1 subequal to antenna 2, accessory flagellum minute, one­articulate; pereopods 57, posterodistal lobes of basis angular, anterior margins of propodus with 1521, 1319, and 1316 short spines respectively; upper lip, medial notch shallow; lower lip, outer lobes horizontal; mandible, palp article 2 with setae, accessory spine row with 1314 slender blades; maxilla 1, outer plate with 1718 spines; maxilla 2, outer plate with 912 stout, spine­like setae; maxilliped, inner plate with 6 small apical spines; uropods slender; telson length 1.41.5 times width.</paragraph>
</subSubSection>
<subSubSection id="C373D2EB5321FFB7FE83B63FFB35FD30" lastPageId="9" lastPageNumber="10" pageId="2" pageNumber="3" type="description">
<paragraph id="8BD681605321FFBCFE83B63FFE40F8C3" blockId="2.[264,859,1820,1886]" box="[264,402,1820,1845]" pageId="2" pageNumber="3">
<emphasis id="B91D5D725321FFBCFE83B63FFE40F8C3" box="[264,402,1820,1845]" italics="true" pageId="2" pageNumber="3">Description</emphasis>
</paragraph>
<paragraph id="8BD681605321FFBCFEB3B667FC89F8A8" blockId="2.[264,859,1820,1886]" box="[312,859,1860,1886]" pageId="2" pageNumber="3">
<typeStatus id="54D23FC25321FFBCFEB3B667FE7AF8A8" box="[312,424,1860,1886]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
, ovigerous female, NSMT­Cr 15894.
</paragraph>
<caption id="DF16D1E85320FFBDFE83B626FC5AF894" httpUri="https://zenodo.org/record/158125/files/figure.png" pageId="3" pageNumber="4" targetBox="[270,1311,326,1782]" targetPageId="3">
<paragraph id="8BD681605320FFBDFE83B626FC5AF894" blockId="3.[264,1323,1797,1890]" pageId="3" pageNumber="4">
<emphasis id="B91D5D725320FFBDFE83B626FE5CF8EB" bold="true" box="[264,398,1797,1821]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
<taxonomicName id="4C69FAE35320FFBDFE12B625FD14F8EB" box="[409,710,1798,1821]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="yoshimurai" status="sp. nov.">
<emphasis id="B91D5D725320FFBDFE12B625FD14F8EB" box="[409,710,1798,1821]" italics="true" pageId="3" pageNumber="4">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EE0095320FFBDFD5BB625FCFBF8EB" box="[720,809,1798,1821]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, 4.3 mm (NSMT­Cr 15894); paratype juvenile, 3.4 mm (CMNC 2004­1306). Lacinia mobilis of left mandible is indicated by an arrowhead. Scale bars = 0.1 mm unless indicated otherwise.
</paragraph>
</caption>
<paragraph id="8BD681605327FFBAFEB3B03FFC15FDB8" blockId="4.[264,1324,284,779]" pageId="4" pageNumber="5">Head: length (excluding rostrum) exceeding pereonite 1; eye large, reniform; rostrum, blunt, deep, reaching about 0.5 times antenna 1 peduncular article 1; lateral head lobe narrowly rounded. Antenna 1: short, length about 0.3 times body length, length ratio of peduncular articles 13 = 1: 0.54: 0.28; accessory flagellum small, scale­like, one­articulate; primary flagellum eight­articulate, articles with distal setae. Antenna 2: length subequal to antenna 1, peduncular article 4, length subequal to article 5, both with clusters of setae; flagellum short, about 0.7 times as long as peduncle length, nine­articulate, article 1 long, 1.7 times as long as article 2, articles with distal setae.</paragraph>
<paragraph id="8BD681605327FFBAFEB3B37FFC46FCFD" blockId="4.[264,1324,284,779]" pageId="4" pageNumber="5">Pereonites, Pleonites, and Urosomites: smooth. Coxae: coxa 1 shorter than 2, length 0.78 times coxa 2, ventrally rounded, with four cusps on hind corner; coxa 23 with four, and three cusps on hind corner; coxa 4, posteroventral lobe subacute, posterior margin strongly excavated; coxa 5, aequilobate, anterior lobe bare, posterior lobe with fine setae; coxa 6 with fine setae on posterior margin; coxa 7 bare.</paragraph>
<caption id="DF16D1E85327FFBAFE83B617FCD8F898" httpUri="https://zenodo.org/record/158126/files/figure.png" pageId="4" pageNumber="5" targetBox="[283,1312,867,1844]" targetPageId="4">
<paragraph id="8BD681605327FFBAFE83B617FCD8F898" blockId="4.[264,1324,1844,1902]" pageId="4" pageNumber="5">
<emphasis id="B91D5D725327FFBAFE83B617FE5CF8BA" bold="true" box="[264,398,1844,1868]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="4C69FAE35327FFBAFE12B617FD14F8BD" box="[409,710,1844,1867]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="yoshimurai" status="sp. nov.">
<emphasis id="B91D5D725327FFBAFE12B617FD14F8BD" box="[409,710,1844,1867]" italics="true" pageId="4" pageNumber="5">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EE0095327FFBAFD5AB617FCF8F8BD" box="[721,810,1844,1867]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, 4.3 mm (NSMT­Cr 15894). Scale bars = 0.1 mm unless indicated otherwise.
</paragraph>
</caption>
<paragraph id="8BD681605326FFBBFEB3B03FFB5EFC90" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
Upper lip (
<typeStatus id="54D23FC25326FFBBFE4FB03FFDFCFEC0" box="[452,558,284,310]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
, CMNC 2004­1306): ventral margin shallowly incised, lobes slightly asymmetric apically. Lower lip: outer lobes broad, horizontal; inner lobes deep. Mandible: left and right incisors eight and seven dentate respectively; left lacinia mobilis broad, eight­dentate, right lacinia mobilis absent; left and right accessory spine rows with 13 and 14 slender blades respectively; palp three­articulate, ratio of articles 13 = 1: 2.5: 4.5, article 1 unarmed, article 2 with four inner marginal setae, article 3 with five marginal, four apical setae, and outer facial fine setae. Maxilla 1: inner plate (
<typeStatus id="54D23FC25326FFBBFBD1B32FFB16FDD0" box="[1114,1220,524,550]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
, CMNC 2004­1306) small, with apical seta; outer plate broad, rounded apically, with 17 slender spine­teeth; palp two­articulate, longer than outer plate, article 2 with six apical spines. Maxilla 2: inner plate with 15 setae apically; outer plate distinctly bent distomedially, rounded apically, with 12 spine­like setae and three setae apically. Maxilliped: inner plate short, not reaching base of palp, apical margin sloping medially, with six evenly spaced apical button­like spines, two setae apically, and three setae medially; outer plate narrow, bent medially, with a strong apical spine and setae; palp four­articulate, article 2 1.1 times as long as article 1, articles 3 and 4 with micropectination on medial margin.
</paragraph>
<paragraph id="8BD681605326FFBBFEA2B257FBB7FB00" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">Gnathopod 1: basis, anterior margin setose, posterior margin weakly expanded, with medial setae; ischium short, with posterodistal seta; merus, ventral margin setose; carpus, length 2.1 times width, ventral margin with two clusters of setae and distal setae, with medial setae; propodus narrowly subrectangular, slightly curved, shorter than carpus, length 2.4 times width, ventral margin with 2 spines and setae, with medial setae, palm transverse; dactylus stout, overhanging palm, length 0.4 times article 6, with setae. Gnathopod 2: as for gnathopod 1 except basis narrower and longer; carpus, length 2.8 times width, ventral margin with three clusters of setae and distal setae, with medial setae; propodus, slightly narrow distally, length 2.8 times width, ventral margin with one spine and setae, palm slightly shorter than that of gnathopod 1, slightly parachelate.</paragraph>
<paragraph id="8BD681605326FFBBFEB3B427FAD3F920" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">Pereopod 3: basis, anterior margin setose, with medial setae; ischium short, with posterodistal setae; ratio of meruspropodus = 1: 1: 1.3, posterior margin of propodus with 16 short spines; dactylus, posterior margin finely pectinate. Pereopod 4: as for pereopod 3 except basis setose on posterior margin; ratio of meruspropodus = 1: 0.7: 1, propodus relatively shorter and thicker than that of pereopod 3, posterior margin of propodus with 21 short spines. Pereopod 5: basis, posterior submargin with fine setae, posterodistal lobe angular; ischium short, anterior margin with spines; ratio of meruspropodus = 1: 0.66: 1, anterior margins spinose; propodus, anterior margin with 21 short spines, length 3.6 times width; dactylus, anterior margin weakly serrated. Pereopod 6: as for pereopod 5 except basis, inner surface with setae; ratio of meruspropodus = 1: 0.85: 1.3, propodus, anterior margin with 19 short spines, length 3.7 times width. Pereopod 7: as for pereopod 6 except; ratio of meruspropodus = 1: 0.77: 1, propodus, anterior margin with 16 short spines.</paragraph>
<paragraph id="8BD681605326FFB8FEB3B7C7FCCAFE20" blockId="5.[264,1324,284,1870]" lastBlockId="6.[264,1323,284,470]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">Coxal gills: present on pereopods 26, pear shaped, laminar, largest gill on pereopod 4. Brood plates: broad, with simple marginal setae of medium length. Pleopods: peduncles, outer margin setose, retinaculae paired; pleopod 1, inner and outer ramus 11 and 12­ articulate; pleopod 2, same as pleopod 1; pleopod 3, inner and outer ramus 10 and 11­articulate. Epimeron 1: narrow, posterior margin straight, with two anteroventral setae. Epimeron 2: posteroventral corner acuminate, ventral margin with 3 setae. Epimeron 3: wider than 1 and 2, posterior margin weakly convex, posteroventral corner produced into a broad tooth, ventral margin with seven setae.</paragraph>
<caption id="DF16D1E85325FFB8FE83B610FCD8F89B" httpUri="https://zenodo.org/record/158127/files/figure.png" pageId="6" pageNumber="7" targetBox="[322,1322,554,1819]" targetPageId="6">
<paragraph id="8BD681605325FFB8FE83B610FCD8F89B" blockId="6.[264,1324,1843,1901]" pageId="6" pageNumber="7">
<emphasis id="B91D5D725325FFB8FE83B610FE5CF8BD" bold="true" box="[264,398,1843,1867]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
<taxonomicName id="4C69FAE35325FFB8FE12B617FD14F8BD" box="[409,710,1844,1867]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="yoshimurai" status="sp. nov.">
<emphasis id="B91D5D725325FFB8FE12B617FD14F8BD" box="[409,710,1844,1867]" italics="true" pageId="6" pageNumber="7">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EE0095325FFB8FD5AB617FCF8F8BD" box="[721,810,1844,1867]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, 4.3 mm (NSMT­Cr 15894). Scale bars = 0.1 mm unless indicated otherwise.
</paragraph>
</caption>
<caption id="DF16D1E85324FFB9FE83B71DFCD8F98E" httpUri="https://zenodo.org/record/158128/files/figure.png" pageId="7" pageNumber="8" targetBox="[268,1308,337,1563]" targetPageId="7">
<paragraph id="8BD681605324FFB9FE83B71DFCD8F98E" blockId="7.[264,1324,1598,1656]" pageId="7" pageNumber="8">
<emphasis id="B91D5D725324FFB9FE83B71DFE5CF9A0" bold="true" box="[264,398,1598,1622]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
<taxonomicName id="4C69FAE35324FFB9FE12B71CFD14F9A0" box="[409,710,1599,1622]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yoshimurai" status="sp. nov.">
<emphasis id="B91D5D725324FFB9FE12B71CFD14F9A0" box="[409,710,1599,1622]" italics="true" pageId="7" pageNumber="8">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EE0095324FFB9FD5AB71DFCF8F9A3" box="[721,810,1598,1621]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, 4.3 mm (NSMT­Cr 15894). Scale bars = 0.1 mm unless indicated otherwise.
</paragraph>
</caption>
<paragraph id="8BD681605324FFB6FEB3B78FFB4FFDB8" blockId="7.[264,1323,1708,1894]" lastBlockId="8.[264,1324,284,590]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">Uropods 13 slender, rami narrowly lanceolate, lacking terminal spine, marginally serially spinose. Uropod 1: peduncle spinose, length subequal to inner ramus, with nine dorsolateral and 7 dorsomedial spines; outer ramus slightly shorter than inner, with eight dorsolateral and four dorsomedial marginal spines; inner ramus with five dorsolateral and six dorsomedial marginal spines. Uropod 2: peduncle with one apicodorsal and one apicomedial spine, length 0.7 times inner ramus; outer ramus length 0.6 times inner ramus, with six dorsolateral and two dorsomedial marginal spines; inner ramus with four dorsolateral and five dorsomedial marginal spines. Uropod 3: peduncle length length 0.6 times inner ramus, with one apicodorsal spine, with two inner marginal setae; outer ramus length 0.5 times inner ramus, with three dorsolateral and one dorsomedial marginal spine; inner ramus with five dorsolateral and five dorsomedial marginal spines. Telson: with ventral keel, length 1.5 times width, dorsal surface with two setae, four penicillate setae, and 15 microscopic setules, lateral margins parallel. Egg number: 10. Male unknown.</paragraph>
<paragraph id="8BD68160532BFFB6FE12B62BFD18F892" blockId="8.[264,1323,1800,1892]" pageId="8" pageNumber="9">
<taxonomicName id="4C69FAE3532BFFB6FE12B62BFD14F8E9" box="[409,710,1800,1823]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yoshimurai" status="sp. nov.">
<emphasis id="B91D5D72532BFFB6FE12B62BFD14F8E9" box="[409,710,1800,1823]" italics="true" pageId="8" pageNumber="9">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EE009532BFFB6FD5BB62BFCFBF8E9" box="[720,809,1800,1823]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="54D23FC2532BFFB6FCBCB62BFC46F8E9" box="[823,916,1800,1823]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
female,
<quantity id="4C912C85532BFFB6FC71B62BFB9DF8E9" box="[1018,1103,1800,1823]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="9" unit="mm" value="4.3">4.3 mm</quantity>
(NSMT­Cr 15894). Setae on pleopod rami are omitted. Pleopod retinacula (coupling spines) are shown enlarged. Scale bars =
<quantity id="4C912C85532BFFB6FEDAB66EFE70F892" box="[337,418,1869,1892]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="0.1">0.1 mm</quantity>
unless indicated otherwise.
</paragraph>
<paragraph id="8BD68160532AFFB7FE83B03FFDBBFE70" pageId="9" pageNumber="10">
<table id="F96973C0532A0041FE83B03FFAF9FE70" box="[264,1323,284,390]" gridcols="9" gridrows="3" pageId="9" pageNumber="10">
<tr id="35598322532A0041FE83B03FFAF9FEC3" box="[264,1323,284,309]" gridrow="0" pageId="9" pageNumber="10" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="7688EA5E532A0041FE83B03FFD7BFEC3" box="[264,681,284,309]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="B91D5D72532AFFB7FE83B03FFE57FEC3" box="[264,389,284,309]" italics="true" pageId="9" pageNumber="10">Etymology</emphasis>
</th>
</tr>
<tr id="35598322532A0041FE83B067FAF9FEA8" box="[264,1323,324,350]" gridrow="1" pageId="9" pageNumber="10">
<th id="7688EA5E532A0041FE83B067FD7BFEA8" box="[264,681,324,350]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">The species is named in honour</th>
<td id="7688EA5E532A0041FD3BB067FD1AFEA8" box="[688,712,324,350]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">of</td>
<td id="7688EA5E532A0041FD44B067FD21FEA8" box="[719,755,324,350]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">the</td>
<td id="7688EA5E532A0041FD71B067FCB3FEA8" box="[762,865,324,350]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">collector</td>
<td id="7688EA5E532A0041FCE3B067FC7CFEA8" box="[872,942,324,350]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">Keizo</td>
<td id="7688EA5E532A0041FC3EB067FBE9FEA8" box="[949,1083,324,350]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">Yoshimura,</td>
<td id="7688EA5E532A0041FBC9B067FBA7FEA8" box="[1090,1141,324,350]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">who</td>
<td id="7688EA5E532A0041FBF7B067FB15FEA8" box="[1148,1223,324,350]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">kindly</td>
<td id="7688EA5E532A0041FB45B067FAF9FEA8" box="[1230,1323,324,350]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">donated</td>
</tr>
<tr id="35598322532A0041FE83B04FFAF9FE70" box="[264,1323,364,390]" gridrow="2" pageId="9" pageNumber="10" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="7688EA5E532A0041FE83B04FFD7BFE70" box="[264,681,364,390]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">the amphipods for us to study.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BD68160532AFFB7FE83B09FFEAFFE23" blockId="9.[264,1321,444,550]" box="[264,381,444,469]" pageId="9" pageNumber="10">
<emphasis id="B91D5D72532AFFB7FE83B09FFEAFFE23" box="[264,381,444,469]" italics="true" pageId="9" pageNumber="10">Condition</emphasis>
</paragraph>
<paragraph id="8BD68160532AFFB7FEB3B0C7FCF0FDD0" blockId="9.[264,1321,444,550]" pageId="9" pageNumber="10">Without the upper lip and the inner lobe of the maxilla 1. The right uropod 3 is small, possibly due to an early stage of regeneration.</paragraph>
<paragraph id="8BD68160532AFFB7FE83B37FFEA3FD83" blockId="9.[264,1323,604,710]" box="[264,369,604,629]" pageId="9" pageNumber="10">
<emphasis id="B91D5D72532AFFB7FE83B37FFEA3FD83" box="[264,369,604,629]" italics="true" pageId="9" pageNumber="10">Va r i a t i o n</emphasis>
</paragraph>
<paragraph id="8BD68160532AFFB7FEB3B3A7FB35FD30" blockId="9.[264,1323,604,710]" pageId="9" pageNumber="10">No males were collected, so sexual dimorphism could not be documented. The spine numbers of the propodus of pereopods 57 decrease in the smaller sized individuals.</paragraph>
</subSubSection>
<subSubSection id="C373D2EB532AFFB7FE83B3DFFB93F9A8" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BD68160532AFFB7FE83B3DFFEBCFCE3" blockId="9.[264,1324,764,1630]" box="[264,366,764,789]" pageId="9" pageNumber="10">
<emphasis id="B91D5D72532AFFB7FE83B3DFFEBCFCE3" box="[264,366,764,789]" italics="true" pageId="9" pageNumber="10">Remarks</emphasis>
</paragraph>
<paragraph id="8BD68160532AFFB7FEB3B207FE41FBD8" blockId="9.[264,1324,764,1630]" pageId="9" pageNumber="10">
The new species is most similar to
<emphasis id="B91D5D72532AFFB7FD5BB207FBCAFCCB" box="[720,1048,804,829]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C69FAE3532AFFB7FD5BB207FBC6FCCB" box="[720,1044,804,829]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="echinoides">Dactylopleustes echinoides</taxonomicName>
.
</emphasis>
The two differ from all other species of
<taxonomicName id="4C69FAE3532AFFB7FE43B26FFD51FC93" box="[456,643,844,869]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B91D5D72532AFFB7FE43B26FFD51FC93" box="[456,643,844,869]" italics="true" pageId="9" pageNumber="10">Dactylopleustes</emphasis>
</taxonomicName>
in having the following combination of characters: 1) the eye is large, reniform; 2) coxa 1 is distinctly shorter than coxa 2, with four cusps on the hind corner; 3) basis of gnathopods 12 are setose; 4) outer lobe of the lower lip is horizontal; 5) the medial notch of the upper lip is shallow; 6) palp article 2 of the mandible is much longer than article 1, with setae and 7) palp article 4 of the maxilliped is slender (see key below).
</paragraph>
<paragraph id="8BD68160532AFFB7FEB3B51FFB93F9A8" blockId="9.[264,1324,764,1630]" pageId="9" pageNumber="10">
<taxonomicName id="4C69FAE3532AFFB7FEB3B51FFDADFBA3" box="[312,639,1084,1109]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yoshimurai">
<emphasis id="B91D5D72532AFFB7FEB3B51FFDADFBA3" box="[312,639,1084,1109]" italics="true" pageId="9" pageNumber="10">Dactylopleustes yoshimurai</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="4C69FAE3532AFFB7FC5CB51FFBA8FBA3" box="[983,1146,1084,1109]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="echinoides">
<emphasis id="B91D5D72532AFFB7FC5CB51FFBA8FBA3" box="[983,1146,1084,1109]" italics="true" pageId="9" pageNumber="10">D. echinoides</emphasis>
</taxonomicName>
by the following character states (states of
<taxonomicName id="4C69FAE3532AFFB7FDEAB547FCD3FB8B" box="[609,769,1124,1149]" class="Malacostraca" family="Pleustidae" genus="Dactylopleustes" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="echinoides">
<emphasis id="B91D5D72532AFFB7FDEAB547FCD3FB8B" box="[609,769,1124,1149]" italics="true" pageId="9" pageNumber="10">D. echinoides</emphasis>
</taxonomicName>
are in parentheses): 1) antenna 1 with 8­articulate flagellum (vs 46), antenna 2 with 9­articulate flagellum (vs 45); 2) coxa 2 with four cusps on the hind corner (vs 3); 3) length of the propodus of gnathopod 1 is 2.42.8 times width (vs 2); 4) gnathopod 1 palm of the propodus is transverse (vs oblique); 5) pereopods 57, propodus with 1521, 1319, and 1316 spines respectively (vs 11, 7, and 8); 6) pereopods 57, posterodistal lobes of basis are angular (vs round); 7) pereopods 57, length of propodus is 3.63.7 times width (vs 2.72.9); 8) pereopods 57, anterior margins of dactyls with very weak teeth (vs strong teeth); 9) uropods 13 are slender and long (vs short, stout); 10) telson, length 1.5 times width (vs 2 times); 11) outer plate of maxilla 1 with 17 18 slender, spine­teeth (vs 15); 12) outer plate of the maxilla 2 with 912 stout, spine­like setae apically (vs 6); 13) accessory spine row of the mandible with 1314 slender blades (vs 7); 14) palp article 3 of the mandible is 1.71.8 times as long as article 2 (vs 1.4) and 15) palp article 2 of the maxilliped is longer than article 1 (vs shorter).
</paragraph>
</subSubSection>
<subSubSection id="C373D2EB532AFFB7FE83B7B7FAFFF920" box="[264,1325,1684,1750]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BD68160532AFFB7FE83B7B7FE45F95B" blockId="9.[264,1325,1684,1750]" box="[264,407,1684,1709]" pageId="9" pageNumber="10">
<emphasis id="B91D5D72532AFFB7FE83B7B7FE45F95B" box="[264,407,1684,1709]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
</paragraph>
<paragraph id="8BD68160532AFFB7FEB3B79FFAFFF920" blockId="9.[264,1325,1684,1750]" box="[312,1325,1724,1750]" pageId="9" pageNumber="10">Known from shallow water around Muroran, Noboribetsu, and Fukushima, Hokkaido.</paragraph>
</subSubSection>
</treatment>
</document>