255 lines
33 KiB
XML
255 lines
33 KiB
XML
<document id="D7D4A3A5A14E247172C76D8A7F8A8651" ID-ISSN="0024-4082" ID-ZooBank="0810A91E-7D8D-4F56-B468-9D5B3A6115E9" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630094174565" checkinUser="felipe" docAuthor="Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D." docDate="2021" docId="03C00E7BFFFD6649FDF429C23D8BA195" docLanguage="en" docName="zlaa052.pdf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Toktokkus Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith 2021" docType="key" docVersion="2" lastPageNumber="894" masterDocId="FFF97603FFF66642FFE4291C3802A521" masterDocTitle="First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group" masterLastPageNumber="901" masterPageNumber="883" pageNumber="894" updateTime="1699143134663" updateUser="plazi">
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<mods:title id="957A0E2D078770CD11036060B8FCC003">First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group</mods:title>
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<mods:namePart id="9AEC90CF9014C0F345AC206A02A35E9D">Kamiński, Marcin J.</mods:namePart>
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<mods:namePart id="15B652E71C3832CD65EC0F93A36A2E13">Gearner, Olivia M.</mods:namePart>
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<mods:namePart id="B1B96FA639FD8DD7002F753E6A056942">Kanda, Kojun</mods:namePart>
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<mods:namePart id="5F36444AC254F038B8CC2EAED2A9E625">Swichtenberg, Kali</mods:namePart>
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<mods:namePart id="E5DA076B8C286CCEC3AF386491C34301">Purchart, Luboš</mods:namePart>
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<mods:namePart id="C4C32936B47B1A437B04908103B5949E">Smith, Aaron D.</mods:namePart>
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<mods:title id="DF8FFCB0FB81132092DEF22C01901574">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="7DEA8DB6AFEEF8E0040C14A211158E0F">2021</mods:date>
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KEY TO THE SPECIES OF THE GENUS
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<taxonomicName id="4C69C4EEFFFD6649FC4829C33C34A5D7" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" box="[940,1078,223,246]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="901" phylum="Arthropoda" rank="genus">
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<emphasis id="B91D637FFFFD6649FC4829C33C34A5D7" box="[940,1078,223,246]" italics="true" pageId="11" pageNumber="894">TOKTOKKUS</emphasis>
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<subSubSection id="C373ECE6FFFD6649FF28281B3D8BA195" pageId="11" pageNumber="894" type="key">
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<paragraph id="8BD6BF6DFFFD6649FF28281B3D8AA43D" blockId="11.[188,1417,263,1205]" box="[204,1416,263,285]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF28281B3D8AA43D" box="[204,1416,263,285]" pageId="11" pageNumber="894">
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1. Apex of elytra not sloped (dorsal view) and depressed with clear margin (
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<figureCitation id="1352A3E8FFFD6649FBCE281B3C79A43D" box="[1066,1147,263,285]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3B</figureCitation>
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)...............
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<taxonomicName id="4C69C4EEFFFD6649FB0828143D8AA43D" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1260,1416,263,285]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="congolensis">
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<emphasis id="B91D637FFFFD6649FB0828143D8AA43D" box="[1260,1416,263,285]" italics="true" pageId="11" pageNumber="894">T. congolensis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BD6BF6DFFFD6649FF33283A3D8AA41A" blockId="11.[188,1417,263,1205]" box="[215,1416,294,316]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF33283A3D8AA41A" box="[215,1416,294,316]" pageId="11" pageNumber="894">
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- Apex of elytra sloped (dorsal view) and flat to slightly convex (e.g.
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<figureCitation id="1352A3E8FFFD6649FC3F283A3C29A41A" box="[987,1067,294,316]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3A</figureCitation>
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) ...............................................2
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</keyLead>
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</paragraph>
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<keyStep id="309DA7C8FFFD6649FF2828583D8AA494" pageId="11" pageNumber="894">
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<paragraph id="8BD6BF6DFFFD6649FF2828583D8AA459" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF2828583D8AA459" pageId="11" pageNumber="894">
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2. Margin of prosternal collar expanded and folded out into a large lip (
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<figureCitation id="1352A3E8FFFD6649FC1528583C43A478" box="[1009,1089,324,346]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3C</figureCitation>
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); elytra round, almost heart- shaped (
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<figureCitation id="1352A3E8FFFD6649FEA8287F399FA459" box="[332,413,355,377]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3C</figureCitation>
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); declivous portion of elytra with little to no tubercles.....................................
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<taxonomicName id="4C69C4EEFFFD6649FB1B28783D8AA459" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1279,1416,355,377]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="tschinkeli">
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<emphasis id="B91D637FFFFD6649FB1B28783D8AA459" box="[1279,1416,355,377]" italics="true" pageId="11" pageNumber="894">T. tschinkeli</emphasis>
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</taxonomicName>
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<paragraph id="8BD6BF6DFFFD6649FF33289E3D8AA494" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF33289E3D8AA494" pageId="11" pageNumber="894">- Margin of prosternal collar not expanded, only occasionally folded out; elytra round or elongate; declivous portion of elytra with tuberculate rows .........................................................................................................3</keyLead>
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<keyStep id="309DA7C8FFFD6649FF2828A33D8AA4D2" pageId="11" pageNumber="894">
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<paragraph id="8BD6BF6DFFFD6649FF2828A33D8AA4F5" blockId="11.[188,1417,263,1205]" box="[204,1416,447,469]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF2828A33D8AA4F5" box="[204,1416,447,469]" pageId="11" pageNumber="894">
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3. Disc of pronotum with prominent punctures (
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<figureCitation id="1352A3E8FFFD6649FD1428A33B41A4F5" box="[752,835,447,469]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3D</figureCitation>
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); gold setae present on elytra (
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<figureCitation id="1352A3E8FFFD6649FB4628A33CF6A4F5" box="[1186,1268,447,469]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3D</figureCitation>
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).....
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<taxonomicName id="4C69C4EEFFFD6649FAC228DC3D8AA4F5" authorityName="Gearner" authorityYear="2021" box="[1318,1416,447,469]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="herero">
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<emphasis id="B91D637FFFFD6649FAC228DC3D8AA4F5" box="[1318,1416,447,469]" italics="true" pageId="11" pageNumber="894">T. herero</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BD6BF6DFFFD6649FF3328C23D8AA4D2" blockId="11.[188,1417,263,1205]" box="[215,1416,478,500]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF3328C23D8AA4D2" box="[215,1416,478,500]" pageId="11" pageNumber="894">- Disc of pronotum only with micropunctures; elytra not covered with setae................................................4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309DA7C8FFFD6649FF2828E03D8AA710" pageId="11" pageNumber="894">
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<paragraph id="8BD6BF6DFFFD6649FF2828E03D8AA733" blockId="11.[188,1417,263,1205]" box="[204,1416,508,530]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF2828E03D8AA733" box="[204,1416,508,530]" pageId="11" pageNumber="894">
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4. Tubercles on elytral sides round/globular, vertically direct (
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<figureCitation id="1352A3E8FFFD6649FC6828E03BF8A730" box="[908,1018,508,530]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 2B, C</figureCitation>
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) ......................................................5
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</keyLead>
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</paragraph>
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<paragraph id="8BD6BF6DFFFD6649FF332B073D8AA710" blockId="11.[188,1417,263,1205]" box="[215,1416,539,561]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF332B073D8AA710" box="[215,1416,539,561]" pageId="11" pageNumber="894">
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- Tubercles on elytral sides pointed, directed posteriorly (e.g.
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<figureCitation id="1352A3E8FFFD6649FC742B073BDDA711" box="[912,991,539,561]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3F</figureCitation>
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) ..........................................................6
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309DA7C8FFFD6649FF282B253D8AA78A" pageId="11" pageNumber="894">
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<paragraph id="8BD6BF6DFFFD6649FF282B253D8BA74C" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
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<keyLead id="30981C58FFFD6649FF282B253D8BA74C" pageId="11" pageNumber="894">
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5. Elytral tubercles dense (1.0–1.5 diameters apart), laterally reaching humerus; elytra round (
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<figureCitation id="1352A3E8FFFD6649FAC82B263D82A76E" box="[1324,1408,570,592]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 2B</figureCitation>
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) ...........................................................................................................................................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FAFD2B453D8BA74C" ID-CoL="4NCBY" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1899" box="[1305,1417,600,622]" class="Insecta" family="Tenebrionidae" genus="Psammodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="mulleri">
|
||
<emphasis id="B91D637FFFFD6649FAFD2B453D8BA74C" box="[1305,1417,600,622]" italics="true" pageId="11" pageNumber="894">T. mulleri</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6BF6DFFFD6649FF332B6B3D8AA78A" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF332B6B3D8AA78A" pageId="11" pageNumber="894">
|
||
- Elytral tubercles sparse (2–3 diameters apart), laterally terminating prior to humerus; elytra elongate to round (
|
||
<figureCitation id="1352A3E8FFFD6649FEDA2B8A398DA78A" box="[318,399,662,684]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 2C</figureCitation>
|
||
)................................................................................................................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FAEE2B8A3D8AA78A" ID-CoL="6WBRW" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1908" box="[1290,1416,662,683]" class="Insecta" family="Tenebrionidae" genus="Psammodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="schultzei">
|
||
<emphasis id="B91D637FFFFD6649FAEE2B8A3D8AA78A" box="[1290,1416,662,683]" italics="true" pageId="11" pageNumber="894">T. schultzei</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309DA7C8FFFD6649FF282BA83D8AA7C9" pageId="11" pageNumber="894">
|
||
<paragraph id="8BD6BF6DFFFD6649FF282BA83D8AA7EB" blockId="11.[188,1417,263,1205]" box="[204,1416,692,714]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF282BA83D8AA7EB" box="[204,1416,692,714]" pageId="11" pageNumber="894">6. Elytral tubercles sparse (4–6 lateral rows), deeply angled...........................................................................7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6BF6DFFFD6649FF332BCF3D8AA7C9" blockId="11.[188,1417,263,1205]" box="[215,1416,723,745]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF332BCF3D8AA7C9" box="[215,1416,723,745]" pageId="11" pageNumber="894">
|
||
- Elytral tubercles dense (more than 6 lateral rows), slightly angled (
|
||
<figureCitation id="1352A3E8FFFD6649FC0D2BCF3C5AA7C9" box="[1001,1112,723,745]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 2B–F</figureCitation>
|
||
).........................................8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309DA7C8FFFD6649FF282BEE3D8AA665" pageId="11" pageNumber="894">
|
||
<paragraph id="8BD6BF6DFFFD6649FF282BEE3D8AA626" blockId="11.[188,1417,263,1205]" box="[204,1416,754,776]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF282BEE3D8AA626" box="[204,1416,754,776]" pageId="11" pageNumber="894">
|
||
7. Elytral tubercles distinct, not confluent, nearly reaching humeri (
|
||
<figureCitation id="1352A3E8FFFD6649FC312BEE3C24A626" box="[981,1062,754,776]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3A</figureCitation>
|
||
) ................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FAF42BEE3D8AA626" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1296,1416,754,775]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="barclayi">
|
||
<emphasis id="B91D637FFFFD6649FAF42BEE3D8AA626" box="[1296,1416,754,775]" italics="true" pageId="11" pageNumber="894">T. barclayi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6BF6DFFFD6649FF332A0D3D8AA665" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF332A0D3D8AA665" pageId="11" pageNumber="894">
|
||
- Elytral tubercles small and short, confluent into rows, terminating well before humeri (
|
||
<figureCitation id="1352A3E8FFFD6649FACC2A0C3D7DA604" box="[1320,1407,784,806]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 2A</figureCitation>
|
||
) ......................................................................................................................................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FB132A2C3D8AA665" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1896" box="[1271,1416,815,837]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="mashunus">
|
||
<emphasis id="B91D637FFFFD6649FB132A2C3D8AA665" box="[1271,1416,815,837]" italics="true" pageId="11" pageNumber="894">T. mashunus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309DA7C8FFFD6649FF282A523D8AA69E" pageId="11" pageNumber="894">
|
||
<paragraph id="8BD6BF6DFFFD6649FF282A523D8AA6A0" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF282A523D8AA6A0" pageId="11" pageNumber="894">
|
||
8. Microtubercles present between tuberculate rows (e.g.
|
||
<figureCitation id="1352A3E8FFFD6649FC922A523BC8A642" box="[886,970,846,868]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3E</figureCitation>
|
||
), tuberculate rows rarely elevated on ridges ...............................................................................................................................................................9
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6BF6DFFFD6649FF332A973D8AA69E" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF332A973D8AA69E" pageId="11" pageNumber="894">- Elytral tubercles all relatively the same size, no microtubercles present between rows, tuberculate rows often elevated on ridges ................................................................................................................................10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309DA7C8FFFD6649FF282AD43D8AA118" pageId="11" pageNumber="894">
|
||
<paragraph id="8BD6BF6DFFFD6649FF282AD43D8AA6DD" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF282AD43D8AA6DD" pageId="11" pageNumber="894">
|
||
9. Body size fairly small (18.0–23.0 mm); elytral tubercles relatively dense, almost confluent; tubercles relatively short ....................................................................................................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FB592AF43D8AA6DD" box="[1213,1416,999,1021]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="tuberculipennis">
|
||
<emphasis id="B91D637FFFFD6649FB592AF43D8AA6DD" box="[1213,1416,999,1021]" italics="true" pageId="11" pageNumber="894">T. tuberculipennis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6BF6DFFFD6649FF332D1A3D8AA118" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF332D1A3D8AA118" pageId="11" pageNumber="894">
|
||
- Body size medium to large (28.0–32.0 mm); elytral tubercles less dense, only occasionally confluent; tubercles taller ................................................................................................................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FAF52D393D8AA118" authorityName="Gearner" authorityYear="2021" box="[1297,1416,1060,1082]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="makuya">
|
||
<emphasis id="B91D637FFFFD6649FAF52D393D8AA118" box="[1297,1416,1060,1082]" italics="true" pageId="11" pageNumber="894">T. makuya</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309DA7C8FFFD6649FF582D5F3D8BA195" pageId="11" pageNumber="894">
|
||
<paragraph id="8BD6BF6DFFFD6649FF582D5F3D8AA157" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF582D5F3D8AA157" pageId="11" pageNumber="894">
|
||
10. Elytral tubercles large, often confluent (
|
||
<figureCitation id="1352A3E8FFFD6649FD232D5F3B18A179" box="[711,794,1091,1113]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3F</figureCitation>
|
||
); tuberculate rows extend over humeri to scutellum; humeri prominent; disk of elytra where smooth often flat (
|
||
<figureCitation id="1352A3E8FFFD6649FC672D7D3BD0A156" box="[899,978,1121,1143]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 3F</figureCitation>
|
||
) .........................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FB1A2D7E3D8AA157" authorityName="Kaminski" authorityYear="2021" box="[1278,1416,1121,1143]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="waclawae">
|
||
<emphasis id="B91D637FFFFD6649FB1A2D7E3D8AA157" box="[1278,1416,1121,1143]" italics="true" pageId="11" pageNumber="894">T. waclawae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6BF6DFFFD6649FF332D9D3D8BA195" blockId="11.[188,1417,263,1205]" pageId="11" pageNumber="894">
|
||
<keyLead id="30981C58FFFD6649FF332D9D3D8BA195" pageId="11" pageNumber="894">
|
||
- Tubercles medium sized (
|
||
<figureCitation id="1352A3E8FFFD6649FDE22D9C3A6DA1B7" box="[518,623,1152,1174]" captionStart="Figure 2" captionStartId="4.[145,223,1514,1536]" captionTargetBox="[155,1419,208,1459]" captionTargetId="figure-156@4.[145,1425,195,1474]" captionTargetPageId="4" captionText="Figure 2. Habitus, morphological details, and labels of species representing Toktokkus: A, syntype of Toktokkus mashunus; B, syntype of T. mulleri; C, holotype of T. schultzei; D, specimen of T. tuberculipennis; E, specimen (locus typicus) of T. vialis sellatus; F, specimen of T. vialis vialis. Scale bar = 10.0 mm." pageId="11" pageNumber="894">Fig. 2E, F</figureCitation>
|
||
), not confluent; tuberculate rows end at humeri; humeri not prominent; disc of elytra where smooth generally convex ....................................................................................
|
||
<taxonomicName id="4C69C4EEFFFD6649FACA2D833D8BA195" ID-CoL="4NCFP" baseAuthorityName="Burchell" baseAuthorityYear="1822" box="[1326,1417,1183,1204]" class="Insecta" family="Tenebrionidae" genus="Psammodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="894" phylum="Arthropoda" rank="species" species="vialis">T. vialis</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</table>
|
||
</treatment>
|
||
</document> |