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<document id="584BD3833083EC51EF432961B0CD8D1C" ID-DOI="10.11646/zootaxa.3852.5.3" ID-GBIF-Dataset="629360af-f7bc-42f5-9d4a-3ae23923922d" ID-ISSN="1175-5326" ID-Zenodo-Dep="251817" ID-ZooBank="F3201303-1585-43B4-A7F8-04AD38DD7565" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460886698860" checkinUser="plazi" docAuthor="Gumovsky, Alex" docDate="2014" docId="03BFAB19FF82FE2BFF4C1DBAFDF0F839" docLanguage="en" docName="zt03852p561.pdf" docOrigin="Zootaxa 3852 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trisecodes africanum Gumovsky, sp. n." docType="treatment" docVersion="8" lastPageNumber="555" masterDocId="FF86D361FF83FE29FFDB1856FF8EFF80" masterDocTitle="A new species of Trisecodes from the Afrotropics (Hymenoptera, Eulophidae)" masterLastPageNumber="561" masterPageNumber="553" pageNumber="554" updateTime="1698359123770" updateUser="plazi">
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<mods:title id="0BDC7BADAA13E2D4EA5D8B3C14B488DF">A new species of Trisecodes from the Afrotropics (Hymenoptera, Eulophidae)</mods:title>
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<mods:namePart id="D262636D01F04F4A4BDCB6D048E2E3CA">Gumovsky, Alex</mods:namePart>
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<mods:date id="57D7366DBAA671BC78AFC9630F6858FB">2014</mods:date>
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<treatment id="03BFAB19FF82FE2BFF4C1DBAFDF0F839" ID-DOI="http://doi.org/10.5281/zenodo.6144910" ID-GBIF-Taxon="119586572" ID-Zenodo-Dep="6144910" LSID="urn:lsid:plazi:treatment:03BFAB19FF82FE2BFF4C1DBAFDF0F839" httpUri="http://treatment.plazi.org/id/03BFAB19FF82FE2BFF4C1DBAFDF0F839" lastPageId="2" lastPageNumber="555" pageId="1" pageNumber="554">
<subSubSection id="C30C4984FF82FE28FF4C1DBAFCDAF8E5" pageId="1" pageNumber="554" type="nomenclature">
<paragraph id="8BA91A0FFF82FE28FF4C1DBAFDF4F986" blockId="1.[151,634,1516,1575]" box="[151,634,1516,1542]" pageId="1" pageNumber="554">
<heading id="D0E1AD63FF82FE28FF4C1DBAFDF4F986" bold="true" box="[151,634,1516,1542]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
<emphasis id="B962C61DFF82FE28FF4C1DBAFDF4F986" bold="true" box="[151,634,1516,1542]" pageId="1" pageNumber="554">
<taxonomicName id="4C16618CFF82FE28FF4C1DBAFDA5F986" authority="Gumovsky" authorityName="Gumovsky" box="[151,555,1516,1542]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="africanum" status="sp. nov.">
<emphasis id="B962C61DFF82FE28FF4C1DBAFE13F986" bold="true" box="[151,413,1516,1542]" italics="true" pageId="1" pageNumber="554">Trisecodes africanum</emphasis>
Gumovsky
</taxonomicName>
,
<taxonomicNameLabel id="A2517B66FF82FE28FDE31DBAFDF4F986" box="[568,634,1516,1542]" pageId="1" pageNumber="554" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA91A0FFF82FE28FF4C1E58FE84F9A7" blockId="1.[151,634,1516,1575]" box="[151,266,1550,1575]" pageId="1" pageNumber="554">
(
<figureCitation id="132D068AFF82FE28FF441E58FE8FF9A7" box="[159,257,1550,1575]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1886,1908]" captionStartId-1="4.[151,250,1951,1973]" captionStartId-2="5.[151,250,1953,1975]" captionTargetBox-0="[161,1431,195,1853]" captionTargetBox-1="[170,1416,193,1928]" captionTargetBox-2="[239,1348,193,1930]" captionTargetId-0="figure@3.[151,1435,195,1862]" captionTargetId-1="figure@4.[170,1416,193,1930]" captionTargetId-2="figure@5.[239,1348,193,1932]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." captionText-1="FIGURE 2. Trisecodes africanum Gumovsky, female holotype, SEM. A and B. Female holotype (Uganda, Semuliki forest). A. Habitus in lateral view (white arrows point to apical margin of fore wing infumation). B. Head and mesosoma, close-up. C F. Sculpture of dorsum viewed at increased magnifications." captionText-2="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri-0="https://zenodo.org/record/251818/files/figure.png" httpUri-1="https://zenodo.org/record/251819/files/figure.png" httpUri-2="https://zenodo.org/record/251820/files/figure.png" pageId="1" pageNumber="554">Figs 13</figureCitation>
)
</paragraph>
<paragraph id="8BA91A0FFF82FE28FF4C1E06FCDAF8E5" blockId="1.[151,1437,1616,2037]" pageId="1" pageNumber="554">
<emphasis id="B962C61DFF82FE28FF4C1E06FEF6F9E9" bold="true" box="[151,376,1616,1641]" pageId="1" pageNumber="554">Comparative notes</emphasis>
. This species is similar to the type-species of the genus,
<taxonomicName id="4C16618CFF82FE28FC341E07FB0AF9E8" box="[1007,1156,1617,1640]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="agromyzae">
<emphasis id="B962C61DFF82FE28FC341E07FB0AF9E8" box="[1007,1156,1617,1640]" italics="true" pageId="1" pageNumber="554">T. agromyzae</emphasis>
</taxonomicName>
, but differs in having the mesosoma distinctly strigate dorsally (
<figureCitation id="132D068AFF82FE28FD9C1E22FD05F90D" box="[583,651,1652,1677]" captionStart="FIGURE 2" captionStartId="4.[151,250,1951,1973]" captionTargetBox="[170,1416,193,1928]" captionTargetId="figure@4.[170,1416,193,1930]" captionTargetPageId="4" captionText="FIGURE 2. Trisecodes africanum Gumovsky, female holotype, SEM. A and B. Female holotype (Uganda, Semuliki forest). A. Habitus in lateral view (white arrows point to apical margin of fore wing infumation). B. Head and mesosoma, close-up. C F. Sculpture of dorsum viewed at increased magnifications." httpUri="https://zenodo.org/record/251819/files/figure.png" pageId="1" pageNumber="554">Fig. 2</figureCitation>
) with tiny elongate ridges forming virtually linear rather than circular meshes (in contrast, the mesosoma is alveolate dorsally in
<taxonomicName id="4C16618CFF82FE28FC8E1ECFFC7EF930" box="[853,1008,1689,1712]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="agromyzae">
<emphasis id="B962C61DFF82FE28FC8E1ECFFC7EF930" box="[853,1008,1689,1712]" italics="true" pageId="1" pageNumber="554">T. agromyzae</emphasis>
</taxonomicName>
,
<figureCitation id="132D068AFF82FE28FBD81ECEFBC4F930" box="[1027,1098,1688,1713]" captionStart="FIGURE 4" captionStartId="6.[151,250,1366,1388]" captionTargetBox="[167,1426,185,1349]" captionTargetId="figure@6.[167,1427,185,1350]" captionTargetPageId="6" captionText="FIGURE 4. Trisecodes agromyzae Delvare &amp; LaSalle, paratype, SEM. A. Habitus in lateral view. B. Head and mesosoma in dorsal view (black arrows point to median groove on mesoscutum and scutellum). C, sculpture of dorsum of mesosoma." httpUri="https://zenodo.org/record/251821/files/figure.png" pageId="1" pageNumber="554">Fig. 4</figureCitation>
C), and the mesoscutum and scutellum lack a longitudinal median groove (
<figureCitation id="132D068AFF82FE28FD421EEAFD52F955" box="[665,732,1724,1749]" captionStart="FIGURE 2" captionStartId="4.[151,250,1951,1973]" captionTargetBox="[170,1416,193,1928]" captionTargetId="figure@4.[170,1416,193,1930]" captionTargetPageId="4" captionText="FIGURE 2. Trisecodes africanum Gumovsky, female holotype, SEM. A and B. Female holotype (Uganda, Semuliki forest). A. Habitus in lateral view (white arrows point to apical margin of fore wing infumation). B. Head and mesosoma, close-up. C F. Sculpture of dorsum viewed at increased magnifications." httpUri="https://zenodo.org/record/251819/files/figure.png" pageId="1" pageNumber="554">Fig. 2</figureCitation>
) (groove present in
<taxonomicName id="4C16618CFF82FE28FC651EEBFBDAF954" box="[958,1108,1725,1748]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="agromyzae">
<emphasis id="B962C61DFF82FE28FC651EEBFBDAF954" box="[958,1108,1725,1748]" italics="true" pageId="1" pageNumber="554">T. agromyzae</emphasis>
</taxonomicName>
,
<figureCitation id="132D068AFF82FE28FBB91EEAFB27F955" box="[1122,1193,1724,1749]" captionStart="FIGURE 4" captionStartId="6.[151,250,1366,1388]" captionTargetBox="[167,1426,185,1349]" captionTargetId="figure@6.[167,1427,185,1350]" captionTargetPageId="6" captionText="FIGURE 4. Trisecodes agromyzae Delvare &amp; LaSalle, paratype, SEM. A. Habitus in lateral view. B. Head and mesosoma in dorsal view (black arrows point to median groove on mesoscutum and scutellum). C, sculpture of dorsum of mesosoma." httpUri="https://zenodo.org/record/251821/files/figure.png" pageId="1" pageNumber="554">Figs 4</figureCitation>
B, C). In addition, the scutellum has only 1 pair of setae posteriorly (3 pairs of setae in
<taxonomicName id="4C16618CFF82FE28FC791EB7FBB2F978" box="[930,1084,1761,1784]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="agromyzae">
<emphasis id="B962C61DFF82FE28FC791EB7FBB2F978" box="[930,1084,1761,1784]" italics="true" pageId="1" pageNumber="554">T. agromyzae</emphasis>
</taxonomicName>
), the disc of the fore wing is darkened in its basal three-fifths (
<figureCitation id="132D068AFF82FE28FDCD1F52FDD2F89D" box="[534,604,1796,1821]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="1" pageNumber="554">Figs 1</figureCitation>
E, F and 2A) as opposed to transparent in
<taxonomicName id="4C16618CFF82FE28FB9F1F53FB55F89C" box="[1092,1243,1797,1820]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="agromyzae">
<emphasis id="B962C61DFF82FE28FB9F1F53FB55F89C" box="[1092,1243,1797,1820]" italics="true" pageId="1" pageNumber="554">T. agromyzae</emphasis>
</taxonomicName>
(
<figureCitation id="132D068AFF82FE28FB371F52FABCF89D" box="[1260,1330,1796,1821]" captionStart="FIGURE 4" captionStartId="6.[151,250,1366,1388]" captionTargetBox="[167,1426,185,1349]" captionTargetId="figure@6.[167,1427,185,1350]" captionTargetPageId="6" captionText="FIGURE 4. Trisecodes agromyzae Delvare &amp; LaSalle, paratype, SEM. A. Habitus in lateral view. B. Head and mesosoma in dorsal view (black arrows point to median groove on mesoscutum and scutellum). C, sculpture of dorsum of mesosoma." httpUri="https://zenodo.org/record/251821/files/figure.png" pageId="1" pageNumber="554">Fig. 4</figureCitation>
), and the metapleuron is evenly reticulate and not clearly delimited from the propodeum (metapleuron smooth and delimited from propodeum by a broad, shallow groove in
<taxonomicName id="4C16618CFF82FE28FD751F1BFCCAF8E4" box="[686,836,1869,1892]" class="Insecta" family="Eulophidae" genus="Trisecodes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="agromyzae">
<emphasis id="B962C61DFF82FE28FD751F1BFCCAF8E4" box="[686,836,1869,1892]" italics="true" pageId="1" pageNumber="554">T. agromyzae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C4984FF82FE2BFF1C1F26FB97FA65" lastPageId="2" lastPageNumber="555" pageId="1" pageNumber="554" type="description">
<paragraph id="8BA91A0FFF82FE28FF1C1F26FEDCF875" blockId="1.[151,1437,1616,2037]" pageId="1" pageNumber="554">
<emphasis id="B962C61DFF82FE28FF1C1F26FEDFF809" bold="true" box="[199,337,1904,1929]" pageId="1" pageNumber="554">Description</emphasis>
.
<emphasis id="B962C61DFF82FE28FE851F26FE3DF809" box="[350,435,1904,1929]" italics="true" pageId="1" pageNumber="554">Female</emphasis>
(
<figureCitation id="132D068AFF82FE28FE1A1F26FD8DF808" box="[449,515,1904,1929]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="1" pageNumber="554">Figs 1</figureCitation>
A, B, F and 2, 3). Length
<quantity id="4CEEB7EAFF82FE28FCF81F26FC46F808" box="[803,968,1904,1929]" metricMagnitude="-4" metricUnit="m" metricValue="7.749999999999999" metricValueMax="8.5" metricValueMin="7.0" pageId="1" pageNumber="554" unit="mm" value="0.7749999999999999" valueMax="0.85" valueMin="0.7">0.700.85 mm</quantity>
. Body dark, antennae brownish, tips of all tibiae and tarsi (except for darker pretarsi) pale (
<figureCitation id="132D068AFF82FE28FD601FC2FD71F82D" box="[699,767,1940,1965]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="1" pageNumber="554">Figs 1</figureCitation>
A, B). Wings transparent except for somewhat darker, pale brown area covering basal 3/5 (more discernible in reflected light,
<figureCitation id="132D068AFF82FE28FC4A1FEEFC5EF850" box="[913,976,1976,2001]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="1" pageNumber="554">Fig. 1</figureCitation>
F, and under SEM,
<figureCitation id="132D068AFF82FE28FB6F1FEEFB79F850" box="[1204,1271,1976,2001]" captionStart="FIGURE 2" captionStartId="4.[151,250,1951,1973]" captionTargetBox="[170,1416,193,1928]" captionTargetId="figure@4.[170,1416,193,1930]" captionTargetPageId="4" captionText="FIGURE 2. Trisecodes africanum Gumovsky, female holotype, SEM. A and B. Female holotype (Uganda, Semuliki forest). A. Habitus in lateral view (white arrows point to apical margin of fore wing infumation). B. Head and mesosoma, close-up. C F. Sculpture of dorsum viewed at increased magnifications." httpUri="https://zenodo.org/record/251819/files/figure.png" pageId="1" pageNumber="554">Fig. 2</figureCitation>
A). Veins dark brown (
<figureCitation id="132D068AFF82FE28FF351F8AFEA3F875" box="[238,301,2012,2037]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="1" pageNumber="554">Fig. 1</figureCitation>
B).
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C18C1FF46FE50" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
Head in dorsal view 2.5× as long as broad, in frontal view (
<figureCitation id="132D068AFF81FE2BFC8618C1FC10FF30" box="[861,926,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
A) 1.2× as broad as high. POL approximately 3× OOL, MDO as long as OOL and twice as long as OCL, compound eye approximately 1.66× as high as broad. Mouth opening wider than length of malar space. Inner margin of eyes subparallel. Genae nearly straight. Frons with V-shaped frontal sulcus and suture-shaped scrobal grooves. Subtorular grooves long, extending from torulus to lateral edge of mouth opening (
<figureCitation id="132D068AFF81FE2BFDCE1971FDD2FEC0" box="[533,604,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Figs 3</figureCitation>
B, C). Mandibles bidentate (
<figureCitation id="132D068AFF81FE2BFC401971FC6CFEC0" box="[923,994,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Figs 3</figureCitation>
B, F). Malar suture sulcus-like, slightly curved basally and terminally (
<figureCitation id="132D068AFF81FE2BFE20191AFDCEFEE4" box="[507,576,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
F). Lower edge of antennal toruli situated slightly below lower eye margin (
<figureCitation id="132D068AFF81FE2BFF441939FF6DFE08" box="[159,227,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
A). Upper frons and areas near frontal sulcus with strigate sculpture (similar to dorsum of mesosoma,
<figureCitation id="132D068AFF81FE2BFAAB1939FF2AFE2C" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
A), areas near scrobal grooves finely alveolate and lower face alutaceous with sculpture of broader meshes (
<figureCitation id="132D068AFF81FE2BFAB419C2FF2BFE50" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C198AFCE8FD04" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
Antenna (
<figureCitation id="132D068AFF81FE2BFEEF198AFEF7FE74" box="[308,377,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Figs 3</figureCitation>
CE). Scape 4.55.0× as long as wide. Pedicel about as long as broad. F1 very transverse, ringlike, F2 and F3 somewhat longer (
<figureCitation id="132D068AFF81FE2BFDC019A9FDECFD98" box="[539,610,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Figs 3</figureCitation>
D, E). Clava 2-segmented, its first segment subquadrate, second segment longer than wide, with terminal spine comprising approximately 1/3 length of segment (
<figureCitation id="132D068AFF81FE2BFBA51A72FB4BFDBC" box="[1150,1221,548,573]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Figs 3</figureCitation>
D, E). All flagellar segments with curved, multiporous plate sensilla and mushroom-shaped sensilla. Multiporous plate sensilla fused to surface of respective segments in their basal 1/3 (
<figureCitation id="132D068AFF81FE2BFD071A3AFCACFD04" box="[732,802,620,645]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[239,1348,193,1930]" captionTargetId="figure@5.[239,1348,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. Trisecodes africanum Gumovsky, female holotype, SEM. A. Head in frontal view. B. Detail of lower face. C E. Detail of antennae. F. Lower face in lateral view. (White arrows point to subtorular grooves in B, C, F, and black arrow to malar suture in F)." httpUri="https://zenodo.org/record/251820/files/figure.png" pageId="2" pageNumber="555">Figs 3</figureCitation>
D, E).
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C1AC6FEFEFCB8" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
Mesosoma about 1.3× as long as broad, strigate dorsally (
<figureCitation id="132D068AFF81FE2BFCBB1AD9FC29FD28" box="[864,935,655,680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1951,1973]" captionTargetBox="[170,1416,193,1928]" captionTargetId="figure@4.[170,1416,193,1930]" captionTargetPageId="4" captionText="FIGURE 2. Trisecodes africanum Gumovsky, female holotype, SEM. A and B. Female holotype (Uganda, Semuliki forest). A. Habitus in lateral view (white arrows point to apical margin of fore wing infumation). B. Head and mesosoma, close-up. C F. Sculpture of dorsum viewed at increased magnifications." httpUri="https://zenodo.org/record/251819/files/figure.png" pageId="2" pageNumber="555">Fig. 2</figureCitation>
). Mesoscutum about 1.8× as broad as long, scutellum slightly longer than mesoscutum and as long as broad, without a furrow in mid part. Mesoscutum with 3 pairs of setae on midlobe; posterior end of scutellum with 1 pair of setae. Axillula present as a short, blunt projection. Spur of fore tibia (calcar) short and straight, fore basitarsus with a row of 5 large and 1 small paddlelike setae ventrally.
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C1B13FD88FC25" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">Metanotum with dorsellum appearing as a narrow reticulate bar with reticulate lateral panels. Propodeum and metapleuron not clearly separated and evenly reticulate. Lower and upper mesepimeron alutaceous, transepimeral sulcus traceable as a curved line.</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C1BE6FD8FFB44" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
Fore wing approximately 1.88× as long as broad. Submarginal vein broken where subcosta meets parastigma; subcosta with 1 dorsal seta. Marginal vein approximately 1.75× as long as costal cell, slightly more than half wing breadth. Stigmal vein very short, stigma sessile, postmarginal vein mostly reduced. Disc with a line of about 13 setae originating near uncus of stigmal vein and extending apically; another line of about 25 setae originating from stigma and extending to anterior margin of fore wing; and another row of about 14 setae connecting base of parastigma with stigmal vein (
<figureCitation id="132D068AFF81FE2BFE341C32FDBDFBFC" box="[495,563,1124,1149]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Figs 1</figureCitation>
A, B). Basal and cubital folds complete setose (speculum closed), subcubital fold along retinaculum as a row of 6 or 7 short setae. Marginal fringe slightly longer than breadth of marginal vein in its broadest part (
<figureCitation id="132D068AFF81FE2BFEAE1CFAFE36FB44" box="[373,440,1196,1221]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Figs 1</figureCitation>
A, B).
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C1C87FE8AFAD5" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
Metasoma (
<figureCitation id="132D068AFF81FE2BFE911C99FE03FB68" box="[330,397,1231,1256]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Figs 1</figureCitation>
A, B, F, 2A) as long as or slightly longer than mesosoma; petiole reduced, transverse. Gaster about 1.4×1.7× as long as wide. Tergite 2 with a row of 4 or 5 hairs laterally. Each of following tergites with one dorsal line of hairs. Basal end of ovipositor nearly reaching base of gaster. Cercal setae short, situated on small tubercles.
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C1D09FB97FA65" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
<emphasis id="B962C61DFF81FE2BFF1C1D09FE8FFAF8" box="[199,257,1375,1400]" italics="true" pageId="2" pageNumber="555">Male</emphasis>
(
<figureCitation id="132D068AFF81FE2BFECA1D09FEDBFAF8" box="[273,341,1375,1400]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Figs 1</figureCitation>
CE). Length
<quantity id="4CEEB7EAFF81FE2BFE201D09FD09FAF8" box="[507,647,1375,1400]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="555" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 mm</quantity>
. All characters identical to female except antennal flagellar segments longer (
<figureCitation id="132D068AFF81FE2BFF2A1DD2FEBDFA1C" box="[241,307,1412,1437]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Fig. 1</figureCitation>
D), mostly with one ring-like multiporous plate sensilla each. Fore wing infumation paler than in female, traceable in reflected light (
<figureCitation id="132D068AFF81FE2BFDE31DF1FDF5FA40" box="[568,635,1447,1472]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Fig. 1</figureCitation>
E); marginal fringe 23× longer than breadth of marginal vein in its broadest part (
<figureCitation id="132D068AFF81FE2BFEEC1D9AFEF9FA64" box="[311,375,1484,1509]" captionStart="FIGURE 1" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[161,1431,195,1853]" captionTargetId="figure@3.[151,1435,195,1862]" captionTargetPageId="3" captionText="FIGURE 1. Trisecodes africanum Gumovsky, light microscopy. A and B. Female holotype. A. Habitus in lateral view. B. Head and mesosoma, close-up. C E. Male paratype. C. Habitus in dorsal view. D. Antennae, close-up. E. Fore wing close-up. F. Female paratype, habitus in lateral view. (Black arrow points to apical margin of proximal dark infumation in D and F)." httpUri="https://zenodo.org/record/251818/files/figure.png" pageId="2" pageNumber="555">Fig. 1</figureCitation>
E). Gaster rectangular in dorsal view, 1.4× as long as broad.
</paragraph>
</subSubSection>
<subSubSection id="C30C4984FF81FE2BFF1C1DB9FA1CF960" pageId="2" pageNumber="555" type="materials_examined">
<paragraph id="8BA91A0FFF81FE2BFF1C1DB9FB5AF9AC" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
<emphasis id="B962C61DFF81FE2BFF1C1DB9FEFFF988" bold="true" box="[199,369,1519,1544]" pageId="2" pageNumber="555">
<typeStatus id="54ADA4ADFF81FE2BFF1C1DB9FE8CF988" box="[199,258,1519,1544]" pageId="2" pageNumber="555">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54ADA4ADFF81FE2BFE591DA6FE64F988" box="[386,490,1520,1544]" pageId="2" pageNumber="555" type="holotype">Holotype</typeStatus>
♀,
<collectingCountry id="F3015A9FFF81FE2BFDC01DA6FDFBF988" box="[539,629,1520,1544]" name="Uganda" pageId="2" pageNumber="555">Uganda</collectingCountry>
, Semuliki National Park,
<geoCoordinate id="EE227CC8FF81FE2BFC791DA6FBC6F988" box="[930,1096,1520,1544]" direction="north" orientation="latitude" pageId="2" pageNumber="555" precision="1" value="0.84188056">N 0°50'30.77&quot;</geoCoordinate>
,
<geoCoordinate id="EE227CC8FF81FE2BFB8C1DA6FA89F988" box="[1111,1287,1520,1544]" direction="east" orientation="longitude" pageId="2" pageNumber="555" precision="1" value="30.103937">E 30°06'14.17&quot;</geoCoordinate>
,
<date id="FFA83CCFFF81FE2BFACD1DA6FA16F988" box="[1302,1432,1520,1544]" pageId="2" pageNumber="555" value="2013-03-19">19.III.2013</date>
; semi-deciduous forest with moister areas near cut logs; altitude
<quantity id="4CEEB7EAFF81FE2BFCBA1E42FC26F9AC" box="[865,936,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="555" unit="m" value="670.0">670 m</quantity>
(A. Gumovsky) (
<collectionCode id="ED0782CAFF81FE2BFBB51E42FB46F9AC" box="[1134,1224,1556,1580]" pageId="2" pageNumber="555">BMNH</collectionCode>
).
</paragraph>
<paragraph id="8BA91A0FFF81FE2BFF1C1E6FFA1CF960" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
<typeStatus id="54ADA4ADFF81FE2BFF1C1E6FFEB6F9D1" box="[199,312,1593,1617]" pageId="2" pageNumber="555" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D10D186FF81FE2BFE9C1E6EFEFBF9D0" box="[327,373,1592,1616]" pageId="2" pageNumber="555" type="generic">2 ♂</specimenCount>
, same data as
<typeStatus id="54ADA4ADFF81FE2BFDF71E6EFD00F9D0" box="[556,654,1592,1616]" pageId="2" pageNumber="555" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED0782CAFF81FE2BFD7A1E6EFD77F9D0" box="[673,761,1592,1616]" pageId="2" pageNumber="555">BMNH</collectionCode>
,
<collectionCode id="ED0782CAFF81FE2BFCD11E6EFCECF9D0" box="[778,866,1592,1616]" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="555">RMCA</collectionCode>
); 2 ♀,
<collectingCountry id="F3015A9FFF81FE2BFC601E6EFB9BF9D0" box="[955,1045,1592,1616]" name="Uganda" pageId="2" pageNumber="555">Uganda</collectingCountry>
, near Najembe, Mabira forest,
<geoCoordinate id="EE227CC8FF81FE2BFA531E6EFE90F9F4" direction="north" orientation="latitude" pageId="2" pageNumber="555" precision="1" value="0.3901361">N 0°23'24.49&quot;</geoCoordinate>
,
<geoCoordinate id="EE227CC8FF81FE2BFEF41E0BFE4BF9F5" box="[303,453,1628,1653]" direction="east" orientation="longitude" pageId="2" pageNumber="555" precision="1" value="33.01817">E 33°1'5.41&quot;</geoCoordinate>
, logged patch in the forest,
<date id="FFA83CCFFF81FE2BFCC71E0AFC38F9F4" box="[796,950,1628,1652]" pageId="2" pageNumber="555" value="2013-02-22" valueMax="2013-02-23" valueMin="2013-02-22">22-23.II.2013</date>
(A. Gumovsky) (
<collectionCode id="ED0782CAFF81FE2BFB571E0AFB6BF9F4" box="[1164,1253,1628,1652]" pageId="2" pageNumber="555">BMNH</collectionCode>
,
<collectionCode id="ED0782CAFF81FE2BFB2E1E0AFAC3F9F4" box="[1269,1357,1628,1652]" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="555">RMCA</collectionCode>
);
<specimenCount id="9D10D186FF81FE2BFABE1E0AFA19F9F4" box="[1381,1431,1628,1652]" pageId="2" pageNumber="555" type="generic">3 ♂</specimenCount>
;
<collectingCountry id="F3015A9FFF81FE2BFF4C1E29FF66F918" box="[151,232,1663,1688]" name="Guinea-Bissau" pageId="2" pageNumber="555">Guinea</collectingCountry>
(Conakry), Nimba Mountain, Gouan Camp, Gallery forest of Zie, nr. “Station de Pompage Zie”, canopy of trees, under-story shrub layer, Fogging 01, Mt Nimba, Zié forest,
<geoCoordinate id="EE227CC8FF81FE2BFCA01EF3FB8BF93C" box="[891,1029,1700,1724]" direction="north" orientation="latitude" pageId="2" pageNumber="555" precision="15" value="7.673055">N 07°40'23''</geoCoordinate>
<geoCoordinate id="EE227CC8FF81FE2BFBD61EF3FB13F93C" box="[1037,1181,1700,1724]" direction="west" orientation="longitude" pageId="2" pageNumber="555" precision="15" value="-8.373334">W 08°22'24''</geoCoordinate>
<quantity id="4CEEB7EAFF81FE2BFB7E1EF2FB72F93C" box="[1189,1276,1700,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="555" unit="m" value="1250.0">1250 m</quantity>
,
<date id="FFA83CCFFF81FE2BFAD21EF2FAFEF93C" box="[1289,1392,1700,1724]" pageId="2" pageNumber="555" value="2011-10-03">3.X.2011</date>
(A. Hernard &amp; D. Van den Spiegel) (
<collectionCode id="ED0782CAFF81FE2BFDDC1E9EFDEEF960" box="[519,608,1736,1760]" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="555">RMCA</collectionCode>
); 1 ♀,
<collectingCountry id="F3015A9FFF81FE2BFD701E9EFCADF960" box="[683,803,1736,1760]" name="Cameroon" pageId="2" pageNumber="555">Cameroon</collectingCountry>
, Nkoemvone,
<date id="FFA83CCFFF81FE2BFC1F1E9EFB07F960" box="[964,1161,1736,1760]" pageId="2" pageNumber="555" value="1980-03-30" valueMax="1980-04-19" valueMin="1980-03-30">30.III-19.IV.1980</date>
(D. Jackson) (
<collectionCode id="ED0782CAFF81FE2BFAF71E9EFA08F960" box="[1324,1414,1736,1760]" pageId="2" pageNumber="555">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C4984FF81FE2BFF1C1EBAFD7EF8F0" pageId="2" pageNumber="555" type="biology_ecology">
<paragraph id="8BA91A0FFF81FE2BFF1C1EBAFD7EF8F0" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="555">
<emphasis id="B962C61DFF81FE2BFF1C1EBAFE91F885" bold="true" box="[199,287,1772,1797]" pageId="2" pageNumber="555">Biology</emphasis>
. Unknown. In
<collectingCountry id="F3015A9FFF81FE2BFE121EBAFD93F885" box="[457,541,1772,1797]" name="Guinea-Bissau" pageId="2" pageNumber="555">Guinea</collectingCountry>
, specimens were collected from the forest canopy by fogging. In both localities in
<collectingCountry id="F3015A9FFF81FE2BFF6F1F59FE80F8A8" box="[180,270,1807,1832]" name="Uganda" pageId="2" pageNumber="555">Uganda</collectingCountry>
, numerous sweepings were conducted under forest canopy, but only those in recent clearings produced specimens. It is possible that the species is normally a canopy inhabitant and the collection of specimens in lower forest levels was associated with recently felled trees.
</paragraph>
</subSubSection>
<subSubSection id="C30C4984FF81FE2BFF1C1F2AFA01F815" box="[199,1423,1916,1941]" pageId="2" pageNumber="555" type="distribution">
<paragraph id="8BA91A0FFF81FE2BFF1C1F2AFA01F815" blockId="2.[151,1437,151,1977]" box="[199,1423,1916,1941]" pageId="2" pageNumber="555">
<emphasis id="B962C61DFF81FE2BFF1C1F2AFED7F815" bold="true" box="[199,345,1916,1941]" pageId="2" pageNumber="555">Distribution</emphasis>
.
<collectingCountry id="F3015A9FFF81FE2BFEBC1F2AFE30F815" box="[359,446,1916,1941]" name="Uganda" pageId="2" pageNumber="555">Uganda</collectingCountry>
(Semuliki forest, Mabira forest),
<collectingCountry id="F3015A9FFF81FE2BFCEC1F2AFC22F814" box="[823,940,1916,1940]" name="Cameroon" pageId="2" pageNumber="555">Cameroon</collectingCountry>
(Nkoemvone),
<collectingCountry id="F3015A9FFF81FE2BFB871F2AFB23F815" box="[1116,1197,1916,1941]" name="Guinea-Bissau" pageId="2" pageNumber="555">Guinea</collectingCountry>
(Nimba Mountain).
</paragraph>
</subSubSection>
<subSubSection id="C30C4984FF81FE2BFF1C1FF6FDF0F839" box="[199,638,1952,1977]" pageId="2" pageNumber="555" type="etymology">
<paragraph id="8BA91A0FFF81FE2BFF1C1FF6FDF0F839" blockId="2.[151,1437,151,1977]" box="[199,638,1952,1977]" pageId="2" pageNumber="555">
<emphasis id="B962C61DFF81FE2BFF1C1FF6FEC8F839" bold="true" box="[199,326,1952,1977]" pageId="2" pageNumber="555">Etymology</emphasis>
. Named for its distribution.
</paragraph>
</subSubSection>
</treatment>
</document>